Gudeodiscus (Gudeodiscus) soosi Pall-Gergely , 2013

Pall-Gergely, Barna & Asami, Takahiro, 2016, A new species of Gudeodiscus Pall-Gergely, 2013 from China, with extraordinary conchological and anatomical features (Gastropoda, Pulmonata, Plectopylidae), ZooKeys 564, pp. 1-19 : 7-10

Gudeodiscus (Gudeodiscus) soosi Pall-Gergely, 2013 Figures 2 F–J, 3 D–F, 4 D–E, 6, 7C, 8B, 9 D–F

Gudeodiscus soosi Páll-Gergely, In: Páll-Gergely & Hunyadi, Archiv für Molluskenkunde 142(1): 31-32, figs 42 a–b, 66. 2013

Characters of the genital structure

(Figs 6, 7c, 8B). Two specimens were anatomically examined. Shells, ethanol-preserved bodies and radulae on double-faced adhesive tape are deposited in coll. JUO.

One of the specimens was aphallic, i.e. the male part of the genitalia was entirely missing. The right ommatophoral retractor passes between the penis and the vagina of the second specimen. Atrium extremely short, slender, long; penis moderately long, spindle shaped; inner wall of the penis with approx. 14 low longitudinal folds which join each other in the direction of the atrium resulting in fewer number of folds posteriorly; on the penial wall of the apical part of the penis there are slit-like “pockets” arranged in a transversal row; no calcareous granules have been found inside these pockets; the penial caecum is situated on the apical portion of the penis, it is approx. one third of the length of the penis; its inner wall is ornamented with several rhomboid papillae with holes in the middle of each papillae; no calcareous granules were found in them; retractor muscle inserts on the apical part of the penial caecum; retractor muscle very long, branched off from the columellar muscle; epiphallus enters penis laterally, at the joint of the penis and the larger penial caecum; its inner wall with three strong longitudinal folds; vagina slightly longer and thicker than the penis; the inner wall of the vagina is with irregular, low, longitudinal folds; the bursa copulatrix starts on the proximal part of the vagina; its base is not thickened; the stalk is slender and very long, the bursa is oval, more thickened than in the other species; diverticulum starts at the end of the vagina, therefore the base of the diverticulum and the base of the bursa copulatrix are very far from each other; diverticulum very slender, without thickening at it end; it is approximately as long as the bursa copulatrix; spermoviduct contained several developing eggs.

Characters of the radula

(Fig. 9 D–E). Radula elongated, but not very slender, central tooth present, laterals 7 or 8 (it is difficult to decide whether the 8th row belong to the laterals or the marginals), standing in straight lines (perpendicular to the central column); marginals approximately 12-13; marginals are placed in slightly oblique rows; central tooth wide-based triangular, smaller than the endocone of the first lateral, but approximately as large as the ectocone; laterals bicuspid, ectocones triangular, endocones have rather parallel margins with triangular tip; marginals usually tricuspid (= the endocone has two cusps); occasionally the innermost cusp is also divided into two cusps resulting in three cusps for the structure equivalent to the endocone of the laterals; some of the external marginals have both the endocone and the ectocone di vided into two cusps; all cusps pointed, the incision between the innermost two cusps (= two cusps of the endocone) is deep.

Differential diagnosis.

See under Gudeodiscus longiplica sp. n.

Remarks.

We cannot rule out the possibility that the aphallic individual was a hybrid (see Schilthuizen et al. 2011).