Scaphisoma densepunctatum, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1369

5.2.7.7. Scaphisoma densepunctatum nov.sp. ( Figs 83, 84 )

T y p e m a t e r i a l: Holotype 6: Mindanao, 30km NW of Maramag, 13.-17.May Bagongsilang, 1700m Bolm lgt., 1996 (SMNS).

Description: Length 2.05 mm, width 1.35 mm. Head and thorax blackishbrown, elytra dark reddish-brown, becoming lighter posterior mid-length. Abdomen dark reddish-brown, with apical segments lighter. Femora and tibiae light, ochreous, tarsi yellowish. Pronotum and elytra not microsculptured and not iridescent. Antennomeres with length/width ratio as: III 15/10: IV 30/8: V 40/9: VI 60/11: VII 60/15: VIII 46/11: IX 54/13: X 52/12: XI 70/13. Pronotum finely and densely punctate, punctures sharply delimited, distinct at 20 times magnification; lateral contours rounded, lateral margin carinae visible in dorsal view except near basis, lateral striae impunctate. Tip of scutellum exposed. Elytra with lateral margin carinae exposed in dorsal view, apical margins rounded, inner apical angles not prominent, situated about in level with outer angles; sutural margin not raised, sutural striae fairly deep, somewhat bent at base, not extending laterally, parallel from level of tip of scutellum toward apical third, punctate as elytral disc; adsutural areas narrow, finely and densely punctate. Lateral striae appearing impunctate. Elytra with discal punctation fine and dense, coarser than pronotal punctation, punctures sharply delimited, with diameters mostly about as large as halves of puncture intervals. Hind wings fully developed. Hypomera and mesanepisterna smooth, not microsculptured. Mesepimera about as long as two thirds of intervals to mesocoxae and about twice as long as wide. Metaventrite lacking microsculpture; median area convex, impressed between metacoxae, very finely and densely punctate; punctation becoming sparser near lateral margins; antecoxal puncture row absent. Submesocoxal areas about 0.04 mm, as fifth of intervals to metacoxae; submesocoxal lines convex, distinctly punctate. Metanepisterna flat, weakly narrowed anteriad, not impressed along straight suture. Tibiae straight. Abdomen with strigulate microsculpture; sternite 1 very finely punctate; submetacoxal areas 0.05 mm long, about as fourth of intervals to apical margin; submesocoxal lines convex, distinctly punctate.

M a l e: [Protarsi broken off and missing]. Mesotarsomeres 1 to 3 conspicuously enlarged, with segments I about twice as wide as, segments II as wide as, and segments III somewhat narrower than apices of mesotibiae. Aedeagus ( Figs 83, 84 ) 1.10 mm long, symmetrical, strongly sclerotized. Apical process of median lobe almost much shorter than basal bulb, inflexed, with oblique ventral side, tapering, with acute tip; articular processes large, prominent. Parameres narrowed toward apical third and curved in dorsal view and almost straight, reaching beyond tip of median lobe; almost straight, evenly wide in middle section in lateral view. Internal sac tubular, without rods or flagellum, with minute denticular structures.

E t y m o l o g y: The species epithet is a Latin adjective meaning densely punctate. Comparative notes: This species may be readily distinguished by the strongly enlarged mesotarsomeres 1 and 2. Its aedeagal characters suggest relationships with S. binaluanum PIC, though the aedeagus is notably larger and stronger sclerotized.

Figures

Figs 83 - 88: Aedeagi of Scaphisoma in dorsal and lateral views; (83, 84), S. densepunctatum nov. sp., scale = 0.2 mm; (85, 86), S. hexamerum nov. sp., scale 0.1 mm; (87, 88), S. kodadai nov. sp., scale = 0.1 mm.