Goniopsara mystica ( Melichar, 1899 )

Gnezdilov, Vladimir M., 2024, Revision of the genus Goniopsara Metcalf, 1952 (Hemiptera: Fulgoroidea: Nogodinidae), with the description of a new species from Laos, Zootaxa 5523 (5), pp. 564-572 : 565-569

Goniopsara mystica ( Melichar, 1899)

( Figs 1–13 )

Goniopsis mystica Melichar 1899: 292 .

Goniopsara mystica Metcalf 1952: 226 .

Goniopsara mystica : Meng et al. 2014: 87 , figs 28–30.

Type material examined. Holotype, ♂, “ Singapore / Biró 1898” [printed] // “ mystica Mel. [hand-written in ink] / det. Melichar [printed]” // “typus” [red, printed] ( HNHM).

Other material examined. Singapore: 1♂, “Coll. I.R.Sc.N.B. / Singapore Nee / Soon Swamp Forest / Mal Trap 1 / Station 25028 / Leg. P. Grootaert ” ( RBINS). [According to the station number the specimen was collected 24 March 2005 (P. Grootaert, pers. com.)] . 1♀, “ Singapore / Coll. Baker ” [printed] // “1922 / 2” [printed and hand-written] // “ mystica / Mel.” [hand-written] ( SNSD) .

Emended description. Structure. Coryphe concave, transverse, without carina, six times as wide as long at midline; anterior margin obtusely angular; posterior margin deeply concave. Metope visible from above, elongate, with strongly concave lateral margins below eyes and slightly concave upper margin ( Figs 2 , 4 ); lateral margins (keels) foliate, overhanging pedicels. Metope smooth, with very week median carina running from its upper margin to its middle and with strong V-shaped carina above straight metopoclypeal suture ( Fig. 4 ). Postclypeus with median and lateral carinae. Anteclypeus with median carina. Ocelli present above scapi. Rostrum reaching hind coxae, with 2 nd segment longer than 3 rd one; 3 rd segment narrowing apically ( Fig. 5 ). Pedicel large, bowl-shaped. Pronotum twice as long as coryphe, with median carina. Paradiscal fields of pronotum not narrow, with high keel-shaped margins. Paranotal lobes of pronotum large, elongate, with convex lower margin. Mesonotum large, four times as long as pronotum, with median and lateral carinae. Tegulae large. Forewings elongate, without hypocostal plate, almost not narrowing apically, with narrow precostal area bearing few transverse veins in its proximal half; clavus long, 4/5 of whole wing length, opened, without apical claw ( Figs 1, 3 ). Basal cell large, nearly triangular. Radius and median running from the basal cell separately. Radius firstly furcating closely to basal cell. Median firstly furcating soon after radius, at the level of knee bulge (bulla), second (posterior) branch (M 2) in female furcating near wing middle and both branches (M 1 and M 2) in male and female multifurcating apically. Cubitus anterior tri-furcate (CuA 3), firstly furcating near to wing middle and becoming multifurcate after claval apex. Hind wings tri-lobed, with weak cubital and vannal clefts ( Fig. 3 ). Basal cell large. Hind wing vein branching sequence: R 5, firstly furcating after the coupling lobe; r-m 1; M 4, furcating apically; m-cua 1; CuA 8, firstly furcating after wing middle; icua 3–4 (transverse veins between the branches of CuA); CuP 1; Pcu 1, with apex turned to cubital cleft; A 1 2; A 2 2. Hind tibia with two lateral spines subapically and 11 apical spines. First metatarsomere large and wide, with two large latero-apical spines and eight intermediate spines arranged in strongly curved horse-shoe shape. Second metatarsomere smaller than first one, with only two large latero-apical spines. First and second metatarsomeres with dense long setae ventrally. Arolium of pretarsus with straight hind margin not protruding beyond the claw apices in dorsal view.

Coloration. Metope, coryphe, pro- and mesonotum yellowish brown to dark brown ( Figs 1, 2 ). Head laterally (lora and preocular fields), paranotal lobes of pronotum, and abdomen yellowish brown, except black lateral margins of sternite VII and gonocoxae VIII in female. Postclypeus yellowish brown, with brown to dark brown median carina and lateral stripes. Rostrum with black apex. Scapus and pedicel light yellow to brown. Forewings brown to dark brown, with whitish band (two large spots partly fused) after its middle and with a large whitish spot at furcation of median vein near to claval suture ( Fig. 3 ). Hind wings translucid, with apical and posterior areas infuscate and with dark brown veins. Legs yellowish brown, with dark brown bands on femora and tibiae. Claws brown to dark brown. Dorso-lateral plates of arolium of pretarsus and apices of leg spines black. Anterior connective laminae of gonapophyses VIII light yellow. Anal tube dark brown. Gonoplacs with dark brown areas of denticles.

Male genitalia structure ( Figs 8–13 ). Pygofer massive, wide, with a large semicircular lobe below the anal tube and with a large lobe-shaped process on hind margin medially surrounded by dense long setae on each side ( Fig. 8 ). Anal tube large, nearly as long as pygofer vertically, narrow in dorsal view, with narrowly rounded apex ( Figs 8, 11 ). Anal column short. Phallobase massive, enlarged apically, strongly curved in lateral view, with pair of nearly quadrate lobes basally and with pair of rounded lobes above ventral aedeagal hooks ( Fig. 10 ). Ventral phallobase lobe massive, with pair of hemisphaerical processes below ventral aedeagal hooks ( Fig. 9 ). Aedeagus with two pairs of long acuminate apically hooks—one pair directed apically and other pair (ventral hooks) directed basally (shaded on the Figs 9, 10 ). Connective with large cup. Style large, with vertically elongate plate and long and narrow capitulum bearing smooth apical tooth (in lateral view) and narrowing apically in dorsal view ( Figs 12, 13 ).

Female terminalia ( Figs 6, 7 ). Hind margin of sternite VII with short semicircular processes (lobes) medially ( Fig. 7 ). Anal tube large, flattened and enlarged apically ( Fig. 6 ). Anal column small.

Total length. Female— 13 mm. Male— 12 mm. ( Melichar (1899) mentioned 14 mm. for the holotype).

Figures

FIGURE 1–3. Goniopsara mystica (Melichar, 1899), Singapore. (1) Holotype, male, lateral view; (2) same, frontal view; (3) female, fore and hind wings (Scale bar—2 mm.).

FIGURE 4–7. Goniopsara mystica (Melichar, 1899), female (Singapore). (4) head, fronto-lateral view; (5) rostrum, lateral view; (6) anal tube, caudal view; (7) sternite VII, ventral view (Scale bar—0.6 mm.).

FIGURE 8–13. Goniopsara mystica (Melichar, 1899), male genitalia (Singapore). (8) pygofer and anal tube, lateral view; (9) penis, ventral view; (10) penis and connective, lateral view; (11) anal tube, dorsal view; (12) style, lateral view; (13) capitulum of style, dorsal view. Not to scale.