Pycnomerus sulcatissimus (Reitter, 1880)

Lord, Nathan P. & Leschen, Richard A. B., 2014, Illustrated Catalogue and Type Designations of the New Zealand Zopheridae (Coleoptera: Tenebrionoidea), Zootaxa 3809 (1), pp. 1-127 : 74-75

Pycnomerus sulcatissimus (Reitter, 1880)

( Figs. 413–417 )

Penthelispa sulcatissima Reitter, 1880b: 5 . Reitter 1880c: 175. Hetschko 1930: 66.

Pycnomerus sulcatissimus: Maddison 2010: 426 (attributed to Reitter, although author and year were not in parentheses).

Pycnomerus sulcatissimus Sharp, 1886: 389 , pl. 12, fig. 22. Broun 1893b: 1095 (reprinted excerpt of Sharp 1886: 389). Hutton 1904: 171. Broun 1912: 421. Hudson 1923: 370. Hetschko 1930: 63. New Synonymy.

Type locality: Penthelispa sulcatissima Reitter : assumed to be Greymouth, as Reitter received specimens from Helms, who collected extensively in Greymouth. Pycnomerus sulcatissimus Sharp : Greymouth.

Broun number: Penthelispa sulcatissima Reitter : none given. Pycnomerus sulcatissimus Sharp : 1949.

Remarks: The nomenclatural history of this species is complex. Reitter first described this species (1880b: 5) under the genus Penthelispa . Sharp (1886: 389) later describes the species Pycnomerus sulcatissimus , at the end of the description stating: “I have retained for this species the trivial name under whith it has been distributed by Herr Reitter.” It is apparent Sharp was unaware of Reitter’s earlier description, and, using material sent to him from Reitter, described the species under Pycnomerus . Sharp elected to not recognize the genus Penthelispa , as he stated (1876: 25): “Pascoe and Leconte have proposed to distinguish the Pycnomeri with distinctly 11-jointed antennae by the name of Penthelispa . Erichson, who pointed out this character [in his 1845 description of Pycnomerini ], considered it unnecessary to make distinct generic names for the two forms; and the present species indicates the correctness of his judgement; for the antennae are just intermediate in structure between the two forms.” The genera Pycnomerus and Penthelispa were later synonymized by Sharp 1894: 474 (missed by Hetschko 1930); thus, Pycnomerus sulcatissimus Sharp is rendered a subjective synonym, as well as a secondary homonym, of Pycnomerus sulcatissimus (Reitter) . Hetschko (1930) listed both species under their respective genera. Reitter did not mention the number of specimens examined of Penthelispa sulcatissima . Sharp did not mention the number of specimens examined of Pycnomerus sulcatissimus . We considered all specimens not given as variants as syntypes. In order to stabilize these names, a lectotype and eight paralectotypes are here designated from the material of Penthelispa sulcatissima Reitter and a lectotype and four paralectotypes are here designated from the material of Pycnomerus sulcatissimus Sharp.

Type material examined: Penthelispa sulcatissima Reitter : Lectotype ( MNHN): card-mounted, “TYP. REITTER // New Zeeland Helms sulcatissima m. [in Reitter’s hand] // Penthelispa sulcatissima Rt TYPE Slip. 85 [green label, in S.A. Ślipiński’s hand]” . Paralectotypes ( MNHN): 5, individually mounted on separate cards and pins [card with three black lines near base, middle line thicker], with identical labels, “Now. Zeeland Helms Reitter. [label bordered with thin black line]” . Paralectotype ( MNHN): card-mounted [card with three black lines near base, middle line thicker], lacking labels, but on with same card and pin type as lectotypes and other paralectotypes. Paralectotype ( MNHN): card-mounted [card with three black lines near base, middle line thicker], “ EX. COLL. REITTER” . Paralectotype ( MNHN): card-mounted [card with black border and one black line near base], “ EX. COLL. REITTER”. Pycnomerus sulcatissimus Sharp : Lectotype ( BMNH): card-mounted, “ Pycnomerus sulcatissimus Type D.S. Greymouth. Helms. [written at base of card in Sharp’s hand] // Type [round label with red border] // Greymouth, [red underline] New Zealand. Helms. // Sharp Coll. 1905-313.” Paralectotype ( BMNH): “ Pycnomerus sulcatissimus D.S. Greymouth. [written at base of card in Sharp’s hand] / / Greymouth, New Zealand. [red underline] Helms. // Sharp Coll. 1905-313.” Paralectotype ( BMNH): “ Pycnomerus sulcatissimus Greymouth. [written at base of card in Sharp’s hand] // I 22 [handwritten] Greymouth, New Zealand. [red underline] Helms. // Sharp Coll. 1905-313.” Paralectotype ( BMNH): “ ♂ Pycnomerus sulcatissimus D.S. Greymouth Helms [written at base of card in Sharp’s hand] // Greymouth, New Zealand. [red underline] Helms. // Sharp Coll. 1905-313.” Paralectotype ( BMNH): “ Pycnomerus sulcatissimus Greymouth, 1885 [written at base of card in Sharp’s hand] // Greymouth, New Zealand. [red underline] Helms. / / Sharp Coll. 1905-313.”

Figures

FIGURES 406–414. Primary type specimens and labels of New Zealand Zopheridae: Pycnomerini. 406–407) Holotype, Pycnomerus simplex Broun, 1880; labels; 408–410) Lectotype, Pycnomerus simulans Sharp, 1876; card-mount; labels; 411–412) Lectotype, Pycnomerus sinuatus Broun, 1893; labels; 413–414) Lectotype, Pycnomerus sulcatissimus (Reitter, 1880; as Penthelispa); labels.

FIGURES 415–421. Primary type specimens and labels of New Zealand Zopheridae: Pycnomerini. 415–417) Lectotype, Pycnomerus sulcatissimus Sharp, 1886, jr. syn. and secondary hom. of Pycnomerus sulcatissimus (Reitter, 1880); card-mount; labels; 418–419) Holotype, Pycnomerus suteri Broun, 1909; labels; 420) Holotype, Pycnomerus tenuiculus Broun, 1914; 421) Holotype labels, Pycnomerus tenuiculus Broun, 1914.