Mischocyttarus anchicaya Silveira, 2015

Silveira, Orlando Tobias, Silva, Suzanna de Sousa & Felizardo, Sherlem Patricia de Seixas, 2015, Notes on social wasps of the group of Mischocyttarus (Omega) punctatus (Ducke), with description of six new species (Hymenoptera, Vespidae, Polistinae), Revista Brasileira de Entomologia 59 (3), pp. 154-168 : 162

publication ID	https://doi.org/ 10.1016/j.rbe.2015.07.006
persistent identifier	https://treatment.plazi.org/id/0D340C17-FFB3-6E20-FFAE-FE85FA28FA34
treatment provided by	Felipe
scientific name	Mischocyttarus anchicaya Silveira
status	sp. nov.

Treatment

Mischocyttarus anchicaya Silveira sp. nov. ( Figs. 3 View Figs , 11F, I View Fig and 12B View Fig )

Holotype: female, Colombia, Valle , Anchicaya, nr. Buenaventura, 16/i/1977, M. Cooper ( NHM).

Diagnosis. Female: wing length 6.5–7.0 mm; clypeus higher than wide; first metasomal segment distinctly short, LSI/HMP 1.37–1.45; pronotal carina exceptionally low at sides, with very narrow lamella; lamella of anteromedial margin of pronotum less reflexed at middle than in other species; mesoscutum with punctures of two very distinct size-classes, the smaller ones often hard to visualize; frons with medium-sized punctures and nearly indistinguishable micropunctures; punctures on head just anterior to occipital carina weaker; yellow colored parts often more restricted, propodeal spots very small; female clypeus yellow with black or blackish marks, pronotum and mesopleura with extensive black areas.

Description. Female: length of fore wing 6.5–7.0 mm. Head nearly as high as wide in frontal view, FHH/ INTOW about 1.02–1.05; clypeus distinctly higher than wide, H/WCL 1.04–1.13, apex narrowly truncate; malar space nearly obsolete; tentorial pit much closer to eye margin than to antennal socket; ocelli as in an equilateral triangle, narrowly separated from eyes, POL/OOL about 1.6; occipital carina well developed, distinct in lateral view and ending fairly below the level of ocular sinus and outwards in relation to margin of pre-foraminal concavity; gena considerably narrower than the upper lobe of the eye. Pronotum without a lateral fovea, central part of the anterior margin of pronotum with the lamella raised and reflexed, with contour reaching just a little more than 180 degrees, much less reflexed than in other species of this group; region immediately behind the lamella without secondary margin; humeral angle well developed, produced into a very weak lobe, pronotal carina exceptionally low at sides for this group, lamella very narrow measuring at sides only ca. 0.09 mm, total width of carina ( WCAR) only about 1.07–1.17 times larger than that of mesoscutum; mesoscutum only a fraction longer than wide, L/WMS 1.04–1.09. Fore wing well-elongated, LDIS /HMP 2.0–2.10. Inner claw of hind tarsus with the apex uniformly rounded, but not enlarged or spoon-shaped. Propodeum with median furrow narrow and rather deep for this group, developed on 4/5 of the length of propodeal dorsum, propodeal valve with upper lamellar bor- der narrow and strongly oblique, with subtriangular outline. First segment of metasoma relatively short ( Figs. 3 View Figs and 12B View Fig ), LSI/HMP 1.37–1.45, with the apex 1.83–1.94 times wider than the basal petiole; spiracles weakly prominent.

Sculpture: very strong and conspicuous. Disk of clypeus with medium-sized punctures and nearly indistinguishable micropunctures, area close to the ventral margin finely reticulate without punctures, shining; upper interantennal area and frons with well visible medium-sized punctures and nearly indistinguishable micropunctures. Mesoscutum sculpture with ordered aspect, punctures of two very distinct size-classes, the larger ones very distinct measuring ca. 0.06 mm or more, the smaller very small and shallow, with less than 0.02 mm, often hard to visualize ( Figs.11F and I View Fig ); mesopleuron and propodeum sculpture similar to that of mesoscutum, but with small-sized punctures still finer; integument in general very shining.

Vestiture:eyes bare; head and mesosoma with very conspicuous whitish appressed or decumbent pilosity; longer and stouter hairs on clypeus, frons and mesoscutum, those on pleura and propodeum still longer but finer and rather decumbent; metasomal terga with appressed pubescence, and more erect hairs on distal segments; first metasomal sternum with very smooth and shining integument, nearly absolutely glabrous except for rather sparse fine and curved hairs; remaining sterna with appressed pubescence, with longer and more erect hairs on distal segments, besides those of glandular brushes.

Color: black (see Fig. 12B View Fig ). Antenna above, most of legs, disk of metasomal terga and sterna, dark brown. Ventral side of antenna very light brown. Large spot on clypeus with grading tones of brown, connected to an interantennal dark mark. Apical and distal margins and teeth of mandible; posterior region of head and mouth parts; posterior area on pronotum at sides; anterior half of mesepisternal plate; posterior area on mesopleuron; large median area on mesosternum; antero-lateral aspect of tegula; whole metapleuron and latero-anterior region of propodeum; extensive areas on fore coxa, femur, trochanter, tibia and tarsus; anterior aspect of mid and hind coxae; femora, trochanters, tibiae, and tarsi beneath, light reddish brown. Most of mandible, lateral region and small spot on apical region of clypeus; inner and outer orbits; narrow supra-antennal concave “V-like” mark, interrupted at center; narrow band encircling anterior region of pronotum (over carina and descending on sides) and narrow strip along posterior margin; posterior half of mesepisternal plate; articular region of mesopleuron; small posterior spot on tegula; anterior half of scutellum (divided by a median line); mesoaxillae; two spots at sides on metanotum; metaxillae; two small spots on propodeum; valvular region; dorso-lateral stripe on fore tibia; dorsum of first and fifth segment of fore tarsus; one stripe on mid coxa; two stripes on hind coxa; distal mark on mid femora; elongated stripe on dorsal aspect of mid tibia; apical spot on mid tibia; distal band on metasomal tergum I; narrow distal and lateral margins of terga II to VI (broadened sublaterally); distal expanded region of sternum I; distal margin (broadened medially) of sterna II to VI, yellow. Wings hyaline with brown veins.

Male: unknown.

Nest: unknown.

Distribution: Colombia (Nariño, Valle).

Etymology: the specific epithet is in reference to the Anchicayá River in Colombia.

Remarks. This new species is very similar to M. chalucas in many important aspects, such as the scarce development of the pronotal carina and anterior pronotal lamella medially, as well as the relatively short first metasomal segment. However, it can be readily differentiated by the narrower clypeus in the female, the weaker punctures of the head just anterior to the occipital carina, and by its color pattern: female clypeus yellow with black or blackish marks and pronotum and mesopleura with extensive black areas ( Fig. 12B View Fig ).

Examined material (Paratypes): Colombia, Nariño, 1 female, Barbacoas , 2–6/v/1976; Valle, 1 female (N. 171), 1 female (N. 172), 3 females, Anchicaya, nr. Buenaventura, 16/i/1977, M. Cooper ( NHM) .