Arachnospila (Arachnospila) gussakovskii Lelej & Loktionov

Lelej, Arkady S. & Loktionov, Valery M., 2011, Review of the nominotypical subgenus of Arachnospila Kincaid (Hymenoptera: Pompilidae) of Russia and neighbouring countries with the lectotypification of enigmatic Pompilus sogdianus Morawitz and description of new species, Zootaxa 2882, pp. 1-18 : 9-11

Arachnospila (Arachnospila) gussakovskii Lelej & Loktionov , sp. nov.

( Figs 2, 5 , 22, 23, 35, 44, 45)

Diagnosis of male. This new species is distinguished from other species of nominotypical subgenus; by having deeply emarginated S6 with strong setae in the bottom of emargination ( Fig. 5 ); by having hypopygium basally without sharp differences between sclerotized and unsclerotized parts (Figs 22, 23 vs. 6–9), and by having black colored body ( Fig. 2 ). FEMALE. Unknown.

Description. MALE. Body length 13.0–13.2 mm. Fore wing length 10.1–10.6 mm. Head width 1.1–1.2 × its height. Frons evenly convex. Ocelli small, POD: OOD 0.74–0.80 ×. Ratio of genal median length to eye median length (lateral view) 0.60–0.65 ×. Malar space (minimum value) 0.25–0.27 × pedicel width (lateral view). Clypeus longitudinally convex; anterior border weakly arcuately emarginated with thin rim. Labrum weakly convex, anterior border with weak median emargination. Mandible bidentate. Flagellomere 1 length 2.3 × its width. Relation of scape, pedicel and two first flagellomeres 44–45: 18–20: 53: 54–55. Mesosoma length dorsally 1.5–1.6 × its maximum width. Pronotum median length 0.3 × its median width, posterior pronotal border obtuse-angulated. Metanotum median length 2.25–2.27 × metapostnotum median length. Metapostnotum shiny with transversal striae. Propodeum evenly convex, its median length 0.80–0.85 × maximum width. Longer spur of mid and hind tibia 0.75 × mid and hind tarsomere 1 respectively. Fore claws asymmetrical with subbasal tooth, mid and hind claws symmetrical with subbasal tooth. Wings infuscate with darker apical part, venation as in Figs 44, 45. Pterostigma small, 1.5–1.7 × longer than first abscissa of R. Second abscissa of R 0.9–1.2 × third abscissa of R. Nervulus weakly postfurcal. Cu almost touch wing border. S6 deeply emarginated with short strong setae in the bottom of emargination ( Fig. 5 ). S8 (hypopygium) with median carina, slightly convergent apically, basally without sharp differences between sclerotized and unsclerotized parts (Figs 22, 23). Genitalia as in Fig. 35 .

Head (except gena), scape ventrally with dense long brownish erect setae; gena and pleurae with dense long erect setae; fore coxa, propodeum laterally and posteriorly with scattered long erect setae; pronotum, mid coxa, T1 laterally with scattered long erect setae; mesonotal scutum, scutellum, femora and S2–S5 with sparse erect setae; T7 apically with dense short and sparse longer erect setae. Body and legs with iridescent brown micropubescence. Body regularly micropunctate. Body and legs black; outer eye orbit near vertex with small pale patch; T2 basally indistinctly reddish in holotype.

Type material. Holotype, 3, Tajikistan: Gissarskii Mountain Ridge, Anzob Pass, 3600 m, 31.VII.1937 (Gussakovskij) [ ZIN]. Paratype, 1 3 with the same label as holotype [ IBSS].

Distribution. Tajikistan.

Etymology. The specific name is dedicated to famous Russian entomologist V.V. Gussakovskij (1904–1948), who was the expert in wasps and excellent collector.

FIGURES 36–51. 36, 38, 40, 42, 44, 46, 48, 50, fore wing; 37, 39, 41, 43, 45, 47, 49, 51, hind wing. 36–39. Arachnospila sogdianoides , France. 40, 41. A. sogdiana , lectotype. 42, 43. A. binaeva (paralectotype of A. sogdiana ). 44, 45. A. gussakovskii sp. nov., paratype. 46–49. A. eisukei, Kunashir. 50, 51. A. fumipennis , Khabarovsk Terr. 36, 37, 40, 41, 44, 45, 48, 49, 3; 38, 39, 42, 43, 46, 47, 50, 51, Ƥ.

FIGURES 52–69. 52, 54, 56, 58, 60, 62, 64, 66–69, fore wing; 53, 55, 57, 59, 61, 63, 65, hind wing.52, 53. Arachnospila fumipennis, Primorkiy Terr. 54–57. A. amurensis, Primorkiy Terr. 58–61, 67. A. clericalis (58, 59, holotype; 60, 61, Irkutsk Prov.). 62–65. A. rufa , Leningrad Prov. 66. A. binaeva ; 68, 69. A. turcorum . 52, 53, 56, 57, 60, 61, 64, 65, 69, 3; 54, 55, 58, 59, 62, 63, 66–68, Ƥ. (66 from Haupt 1930; 67 from Wolf & Móczár 1972; 68, 69 from Zonstein 2002a).

Figures

FIGURES 1 – 5. 1, 4. Arachnospila sogdiana 3. 1. Lectotype, specimen and labels; 4. S 6, ventral view. 2, 5. A. gussakovskii sp. nov., holotype. 2. Specimen and labels; 5. S 6, ventral view. 3. A. binaeva, Ƥ specimens and labels (paralectotype of P. s o g d i - anus).

FIGURES 28 – 35. Male genitalia, ventral view. 28. Arachnospila sogdianoides, Chita Prov. 29. A. sogdiana, lectotype. 30. A. fumipennis, Irkutsk Prov. 31. A. eisukei, Iturup. 32. A. amurensis, Primorkiy Terr. 33. A. clericalis Irkutsk Prov. 34. A. rufa, Omsk. 35. A. gussakovskii sp. nov., holotype.