Anomalophylla moxiensis, Ahrens, 2005

Ahrens, Dirk, 2005, Taxonomic revision of the genus Anomalophylla Reitter, 1887 (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), Zootaxa 1076 (1), pp. 1-62 : 57-59

Anomalophylla moxiensis sp. n.

( Fig. 4N–P , 8 )

Type material. Holotype: ♂ ”1998 China Expedition J. Farka č, D. Král, J. Schneider & A.

Smetana / China: Sichuan, Moxi 29°13’N 102°10’E, 1600 m 2.VII.1998, D. Král ( DEI) .

Paratypes: 21 ♂♂, 9 ♀♀ same data as holotype ( TICB, CA, CK) , 8 ♂♂, 4 ♀♀ CHINA :

SICHUAN; Moxi ; 29°13’N 102°10’ E; 1600 m; 2.vii.1998 ; / 1998 China Expedition J. Farka č, D. Král, J. Schneider & A. Smetana ( TICB), 3 ♂♂ China Sichuan, Moxi VI.1993

N. Hackel lgt. ( CN, CA), 1 ♂ “ China Sichuan Gongga Shan, Moxi 1300 m, 10–11.VII.96 29°13’N 102°10’E / collected by J. Farka č, P. Kabátek & A. Smetana ( NHMB), 11 ♂♂ “W SICHUAN 1–7.VII.1994 29°37’N 102°07’E 1200–1900 m Moxi­Hailougou lgt. J. Farka č

& D. Král ( NHMB, CK), 3 ♂♂, 2 ♀♀ “ China Cebt. Sichuan Luding Co. Moxi env. 22.5 .–

10.6. Benes ( CA), 15 ♂♂, 21 ♀♀ “C. CHINA, W. Sichuan, Luding Xian, Moxi , 9 –

14.vii.1999, V. Benes leg.” ( TICB), 2 ♂♂, 6 ♀♀ “ China West Sichuan Moximian Liding

Co. 13.–18.7.94 Benes” ( CA), 3 ♂♂ “ CHINA Sichuan, Gongga Shan, Hailuogou , btw .

Camp 1 & 2, 1900–2500 m 4.VII.98 29°35’N 102°00’E C50 / collected by A. Smetana, J.

Farkac and P. Kabátek ” ( HAHC, CA), 3 ♂♂, 1 ♀ “W Sichuan 2.–6.VI.1994 29°36’N

102°06’E 1900–2900 m Gonggashan­Hailougou lgt. D. Kral & J. Farkac ” ( CK), 3 ♂♂, 1 ♀

” Sichuan, Moxi Gongashan Mts. 28.VI.–2.VII.1994 Bolm lgt., 1650 m ” ( TICB), 3 ♂♂ , 1

♀ ” CHINA Sichuan pr. Liziping 28.6.– 3.7.1991 R. Dunda lgt.” ( CN, CA) , 1 ♂ “ China (Sichuan) Pass zw. Yaan u. Fulin ( Wa­shan olim) ~ 1900 m 6./ 7.VII.1995 Heinz leg.” ( CDKC) , 1 ♂, 1 ♀ “ Siao­Lou Chasseurs du P. Dejean 1904 / Muséum Paris 1952 Coll. R. Oberthür ” ( MNHN) , 1 ♂, 2 ♀♀ “ Su­Tchuen Siao­Lou 1897 / Muséum Paris 1952 Coll. R. Oberthür ” ( MNHN) , 1 ♂ “ Thibet Tatsienlou Mgt. Biet / Muséum Paris 1952 Coll. R. Oberthür ” ( MNHN) , 1 ♂ “ Thibet Tatsienlou Chasseurs Thibétains 1896 / Muséum Paris 1952 Coll. R. Oberthür ” ( MNHN) , 2 ♂♂, 7 ♀♀ “ Museum Paris Se­Tchouen Env. de Tatsien­Lou Mo­Sy­Mien Pèvre Aubert 1902” ( MNHN) .

Holotype description. Length: 6.1 mm, length of elytra: 3.6 mm, width: 3.2 mm. Body oblong, black; dorsal surface dull; head and pronotum with long, dense, erect setae; brown pilosity on elytra sparse; pilosity on head, pronotum, elytra. Head: Labroclypeus transverse, widest medially with lateral margins moderately convex, convergent anteriorly and toward base; anterior angles strongly rounded; lateral border and ocular canthus producing distinct blunt angle; anterior and lateral margin strongly reflexed; anterior margin shallowly sinuate medially; surface almost flat medially, moderately shiny with double punctation; fine, glabrous punctures mixed with coarse, dense punctures each bearing a long, erect seta. Frontoclypeal suture distinct; weakly curved, slightly elevated; smooth area anterior to eye as wide as long. Ocular canthus moderately long, slender; finely, densely punctate; densely setose. Frons with double punctation; fine glabrous punctures mixed with coarse, moderately dense punctures each bearing a long, erect seta; punctation less dense, predominantly fine, almost glabrous on posterior part of frons. Eyes small, ratio of diameter / interocular width: 0.46. Antenna with ten antennomeres, brown; club with five equal in length antennomeres; club 2.5 times as long as the remaining antennomeres combined, reflexed. Prementum weakly elevated, transversely carinate; distinctly concave apical to elevation. Pronotum: widest adjacent to base; lateral margins in posterior half moderately convex and convergent anteriorly, in anterior half strongly convex and convergent; anterior angles not produced and strongly rounded; posterior angles moderately rounded; anterior margin medially weakly convex with distinct, fine marginal line; basal margin medially without fine marginal line. Pronotal surface with dense, double punctation; fine, glabrous punctures mixed with large punctures bearing a moderately long erect seta; large punctures lacking on centre of disc. Anterior and lateral borders setose; basal margin of hypomeron not produced ventrally, not transversely sulcate anterior to base. Scutellum : moderately long; apex moderately sharp; with fine, sparse punctures; minute setae present in punctures. Elytra: oblong, widest medially; striae indistinctly impressed, finely densely punctate; intervals distinctly convex with fine punctures along striae; punctures on odd intervals with long, erect setae which are anteriorly longer, denser; sutural interval with robust, long, single seta. Epipleural edge fine, ending at strongly convex external apical angle of elytra; epipleura densely setose, strongly curved in anterior third of elytra; apical border chitinous, without short microtrichomes. Venter: Ventral surface dull with fine, dense punctures. Metasternum with dense, long setae, setae partially appressed, partially erect. Metacoxa glabrous near articulation of the leg; elsewhere with fine, long appressed setae. Abdominal sternites with indistinct transverse row of coarse punctures bearing thick setae between fine, dense punctation; all sternites with fine, long setae; tegument of abdominal sternites (60x magnification) with fine polygonal mesh pattern formed by microtrichomes; penultimate sternite at midline with longitudinally impressed line. Mesosternum between mesocoxae narrow, narrower than mesofemur, with irregularly scattered, strong setae. Ratio of length of metepisternum / metacoxa: 1 / 1.73. Pygidium strongly convex; posteriorly shiny; with fine, dense punctures bearing fine, moderately long setae; pilosity beside apical margin denser; without smooth midline. Legs: moderately slender, with shiny surface. Femora with two longitudinal rows of setae; finely, densely punctate and setose; anterior edge of metafemur acute, lacking adjacent serrated line; posterior margin weakly convex with a few fine setae medially; ventral posterior margin weakly widened in apical half; posterior margin smooth ventrally and dorsally. Metatibia moderately slender, long, widest at apex; ratio width / length: 1 / 3.3; dorsally sharply margined, dorsal edge serrate with two groups of spines, basal group at one third, apical one at two thirds of metatibial length, basally with a few single spines in punctures; lateral face longitudinally convex with dense, fine punctures; ventral edge serrated with five strong, long, equally spaced spines; medial face finely punctate; apex sharply truncate interiorly near tarsal articulation. Tarsomeres dorsally glabrous, coarsely punctate; ventrally with sparse, short setae; metatarsomeres dorsally without longitudinal impressions, ventrally with strongly serrated ridge adjacent to strong longitudinal carina, laterally with strong longitudinal carina; first metatarsomere slightly shorter than following two tarsomeres combined, one third of length longer than upper tibial spur. Protibia short, bidentate. Protarsal claws asymmetrical; basal tooth of inner protarsal claw widened, truncate apically. Aedeagus: Fig. 4N–P .

Intraspecific variation. Length: 6.1–7.6 mm, length of elytra: 3.6–4.6 mm, width: 3.2– 3.8 mm. Coloration varies strongly: sometimes legs and elytra reddish brown, elytra apically and laterally dark, entirely black (including legs), entirely reddish brown, reddish brown with dark apex of elytra, or reddish brown with dark pronotum and dark apex of elytra. ♀: antennal club with three antennomeres and distinctly shorter than the remaining antennomeres combined; eyes equal in size to that of male.

Diagnosis. This species differs from the similar A. huashanensis by the shape of parameres ( Fig. 4N–P ). The right paramere is curved ventrally and its basal lobe is almost as wide as long and not produced distally, its medial margin is abruptly reflexed ventrally.

Etymology. The new species is named after Moxi, the type locality.

Figures

FIGURE 4. (A–C) Anomalophylla wulingshanica sp. n. (holotype); (D–F) A. ganhaiziensis sp. n. (holotype); (G–J) A. dongchuanensis sp. n. (holotype); (K–M) A. qinlingensis sp. n. (holotype); (N–P) A. moxiensis sp. n. (holotype); (Q–S) A. plagipennis sp. n. (holotype). (A, D, G, K, N, Q) aedeagus, left side lateral view; (B, E, H, L, O, R) parameres, dorsal view; (C, F, J, M, P, S) aedeagus, right side lateral view. Scale bar = 0.5 mm.

FIGURE 8. Present known distribution of Anomalophylla bicolor, A. dongchuanensis, A. ganhaiziensis, A. huashanica, A. kangdingensis, A. kozlovi, A. mawi, A. moxiensis, A. plagipennis, A. qinlingensis, and A. wulingshanica.