Triasaria olympia, Shear, 2020

Shear, William A., 2020, The millipede family Striariidae Bollman, 1893: I. Introduction to the family, synonymy of Vaferaria Causey with Amplaria Chamberlin, the new subfamily Trisariinae, the new genus Trisaria, and three new species (Diplopoda, Chordeumatida, Striarioidea), Zootaxa 4758 (2), pp. 275-295 : 290-294

Triasaria olympia , new species

Figs. 48–54

Types: Male holotype, eight male and three female paratypes from WASHINGTON: Thurston Co.: Hospital Creek, above confluence with Skookumchuck River , N46°46.396’, W122°35.133’, collected 15 December 2003, by William Leonard & Kelly McAllister, deposited in Burke Memorial Museum, University of Washington , Seattle, Washington.

Etymology: The species epithet is a noun in apposition, after the city of Olympia, Washington.

Diagnosis: Distinct from congeners in the form of the gonopods. The median angiocoxite anteriormost branches (mac1, Figs. 48 , 52 ) bear a posterior array of long, thin spines not present in T. rex or T. washingtonensis , and the median branch of the lateral angiocoxite (lac2, Figs. 50 , 53 ) has an irregular array of spines in T. olympia such that the branch does not appear comb-like. The ninth legs ( Figs. 51 , 54 ) are much more substantially reduced and fused than in the other two species.

Description: Male paratype from Hospital Creek. Length ca. 10 mm, greatest width 0.6 mm. Body form and secondary sexual modifications as described for genus. Gonopods ( Figs. 48–50 , 52, 53 ) robust, mesal angiocoxite 2-branched, anteriormost branch (mac1, Figs. 52, 53 ) longest, set laterally with long, acute spines; posteriormost branch (mac2, Fig. 53 ) smaller, blunt. Lateral angiocoxite with anteriormost branch (lac1, Figs. 48, 50 , 53 ) curving first laterally, then sharply posteriorly, distal part with many mesally directed, long, acute spines. Lateral angiocoxite mesal branch (lac2, Figs. 50 , 53 ) shorter, curved posteriorly, densely set with complex array of mesally directed spines, some branched. Lateral angiocoxite posteriormost branch (lac3, Figs. 52, 53 ) large, blunt, simple, slightly sigmoid. Colpocoxite (cc, Fig. 53 ) relatively small, membranous, with small, triangular teeth. Ninth legpair with all elements completely fused. Telopodite portion (t9, Fig. 54 ) narrowed, cupped anteriorly, acute, triangular lateral tooth near apex. Coxa with no gland pore detected, with relatively small anterior lobe (cl, Fig. 54 ). Sternal spiracle (sp, Fig. 54 ) very large.

Distribution: WASHINGTON: Thurston Co.: Wetland beside Kennedydell Park, 50’ asl, N46°59.249’, W122°58.190’, 4 April 2004, W. Leonard, m. Ranier Road, 3.0 mi NW of Military Road, Fort Lewis Military Reservation, 29 November 2003, W. Leonard, m, ff. Woodard Bay Natural Resource Conservation, maple leaves, 2 March 2003, W. Leonard, m. Olympia, Watershed Park, 11 February 2003, W. Leonard, m. The Evergreen State College, Olympia, beside main parking lot, 14 February 2003, W. Leonard, C. Richart, m. McAllister Springs, 60–123’ asl, 47°07.837’N, 122°43.722’W, 7 February 2004, W. Leonard, C. Richart, m, ff. Same, but 47°02.946’N, 122°43.678’W, 22 February 2004, m. Capitol State Forest, Potosi Creek, 46°58.59’N, 123°06.17’W, 18 December 2002, J. Ziegstrum, R. Shoals, m. Capitol State Forest, along Rock Candy Mtn. Road, 70’ asl, N47°01.54’, W123°04.41’, 18 December 2004, W. Leonard, m, f.Capitol State Forest, 2 mi S of Rock Candy Mtn, 10 December 2002, W. Leonard, m. Capitol State Forest, Bob Baumer’s Grove, 287’ asl, N46°53.76’, W123°05.78’, 14 November 2004, W. Leonard, m. Black Lake and Belmore Road at 66 th Avenue, Tumwater, N46°59.34’, W122°57.99’, 19 November 2004, W. Leonard, m, f. Clallam Co.: Deer Park Road, 6.1 mi N of US 101, 1123’ asl, 14 June 2003, W. Leonard, m. Lewis Co.: Gifford Pinchot National Forest, Kraus Ridge Trail, 6 mi N of?Runate, 680’ asl, N46°27.87’, W121°53.86’, 21 September 2003, W. Leonard, m. Gifford Pinchot National Forest, FS25, 8.5 mi S of Rendell, 1050’ asl, N46°26.456’, W121°59.796’, 6 December 2003, W. Leonard, C. Richart, mm, ff. Gifford Pinchot National Forest, as above, but W46°76.489’, N121°59.890’, 21 December 2003, mm, ff. 604 Roswell Road, Centralia, N46°43.339’, W122°56.664’, 280’ asl, 25 January 2004, W. Leonard, C. Richart, m. SR508 at Bremer, 750’ asl, N46°35’30”, W122°25’55”, 6 December 2003, W. Leonard, C. Richart, mm, ff. SE end of Riffle Lake, 650’asl, N46°27.865’, W122°10.236’, 9 April 2004, W. Leonard, C. Richart, m, ff. Rainbow Falls State Park, SR6 16.1 mi W of I- 5,124 m asl, N46.6302°, W123.2316°, 2 April 2008, W. Leonard, C. Richart, S. Derkarabetian, mm. Grays Harbor Co.: Middle Fork Satsop River, 2.9 mi N of SR12, N47°01.98’, W123°31.55’, 31 December 2004, W. Leonard, mm, f. Capitol State Forest, Porter Creek Campground, N46°58.69’, W123°15.25’, 3 March 2005, W. Leonard, m. Brooklyn Road, 3.4 mi W of Garrard Creek Road, 300’ asl, N46°47.85’, W123°21.31’, 7 February 2005, W. Leonard, female presumably this species. Garrard Creek Road at Weyerhauser D-line, 150’ asl, N46°48.24’, W123°18.99’, 7 February 2005, W. Leonard, female presumably this species.

MAP. Washington State, USA, showing selected localities for species of Trisaria . Some symbols may represent more than one nearby locality.

Figures

FIGURES 48–51. Trisaria olympia. Fig. 48. Gonopods, anterior view. Fig. 49. Gonopods, posterior view. Fig. 50. Tip of left gonopod, posterior view. Fig. 41. Legpair 9 of male, anterior view. Labels as in Figs. 21–23.

FIGURES 52–54. Trisaria olympia. Fig. 52. Gonopods, anterior view. Fig. 53. Gonopods, posterior view. Fig. 54. Left leg 9 of male, anterior view; ap, apodeme, cl, coxal lobe; other labels for this and Figs. 52 & 53 as in Figs. 21–23.