Stenus, Latreille, 1797

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium IV. Staphylinine Group (Part 1) Euaesthetinae, Leptotyphlinae, Megalopsidiinae, Oxyporinae, Pseudopsinae, Solieriinae, Steninae, Bulletin of the American Museum of Natural History 2440 (265), pp. 1807-2440 : 1807-2440

— Puthz, 1966g: 132 ( Stenus ; lectotype designation: Ribeira de Janella-Porto Moniz; synonym cicindela Sharp, 1874: 85 ( Stenus ; Type locality: Southern Japan and northern China).

— Bernhauer and Schubert, 1911: 173 ( Stenus ; subgenus Hypostenus ; catalog).

— Nakane, 1963a: 86 ( Stenus ; characters; habitus photograph; Japan).

— Puthz, 1967d: 142 ( Stenus ; synonym of cicindeloides ).

— Puthz, 1968b: 45 ( Stenus ; synonym of dissimilis ).

— Puthz, 1981a: 11 ( Stenus ; synonym of cicindeloides ).

— Ryvkin, 1990: 174 ( Stenus ; synonym of cicindeloides ).

polypterus Bernhauer, 1938: 30 ( Stenus ; subgenus Hypostenus ; Type locality: Nordwestl. China: Chinkiang).

— Puthz, 1968b: 45 ( Stenus ; synonym of dissimilis ).

— Puthz, 1981a: 11 ( Stenus ; synonym of cicindeloides ).

— Ryvkin, 1990: 174 ( Stenus ; synonym of cicindeloides ).

coomani Cameron, 1940d: 250 ( Stenus ; subgenus Hypostenus ; Type locality: Tonkin: Hoa Binh).

— Puthz, 1968b: 45 ( Stenus ; synonym of dissimilis ).

— Puthz, 1981a: 11 ( Stenus ; synonym of cicindeloides ).

— Ryvkin, 1990: 174 ( Stenus ; synonym of cicindeloides ).

cincinnifer Puthz, 2000: 190 ( Stenus ; subgenus Hypostenus ; Type locality: Peru: Tampopata [ Note: Probably Tambopata]).

- DISTRIBUTION: Peru.

cinctiventris Cameron, 1938 , see: frater L. Benick, 1916 .

cinerascens Erichson, 1839 , see: melanarius Stephens, 1833 .

cinereus L. Benick, 1917b: 304 ( Stenus ; subgenus Hypostenus ; Type locality: Columbien: Muzo).

— Scheerpeltz, 1933: 1175 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1938c: 275 ( Stenus ; characters).

- DISTRIBUTION: Colombia.

circe Puthz, 1972 r: 501 ( Stenus ; subgenus Hypostenus ; Type locality: New Guinea:

Mt. Kaindi, 2400 m).

— Puthz, 1982a: 135 ( Stenus ; New Guinea).

- DISTRIBUTION: New Guinea.

circularis Gravenhorst, 1802: 157 ( Stenus ; Type locality: Not cited, but presumably Brunsvigae ).

— Latreille, 1804: 354 ( Stenus ; characters; Germany).

— Gyllenhal, 1810: 479 ( Stenus ; characters; Sweden).

— Ljungh, 1810: 160 ( Stenus ; characters; Sweden).

— Curtis, 1827: pl. 164 ( Stenus ; England).

— Mannerheim, 1830: 44 ( Stenus ; Finland; Sweden).

— Mannerheim, 1831: 458 ( Stenus ; Finland; Sweden).

— C. Sahlberg, 1832: 432 ( Stenus ; characters; Finland).

— Stephens, 1833: 289 ( Stenus ; characters; England).

— Runde, 1835: 17 ( Stenus ; characters; Germany).

— Lacordaire, 1835: 451 ( Stenus ; characters; France).

— Zetterstedt, 1838: 71 ( Stenus ; characters; habitat; Lapland).

— Erichson, 1839a: 556 ( Stenus ; characters; Germany).

— Heer, 1839: 222 ( Stenus ; characters; Switzerland).

— Stephens, 1839: 412 ( Stenus ; characters; England).

— Erichson, 1840: 717 ( Stenus ; characters; Europe).

— Laporte, 1840: 185 ( Stenus ; characters; France).

— Hochhuth, 1849: 176 ( Stenus ; characters; Tiflis).

— Redtenbacher, 1849: 727 ( Stenus ; characters; Austria).

— Leprieur, 1851: 197 ( Stenus ; characters).

— Fairmaire and Laboulbène, 1856: 585 ( Stenus ; characters; France).

— Thomson, 1857: 228 ( Stenus ; characters; Sweden).

— Redtenbacher, 1857: 222 ( Stenus ; characters; Austria).

— Kraatz, 1857d: 774 ( Stenus ; characters; Germany).

— Thomson, 1860: 219 ( Stenus ; characters; Scandinavia).

— Rye, 1864: 64 ( Stenus ; characters; England).

— Baudi, 1870: 386 ( Stenus ; characters).

— Hochhuth, 1872: 155 ( Stenus ; cited as circuralis; notes; Ukraine).

— Fauvel, 1873: 91 [= 1873a: 249] ( Stenus ; characters; France; Europe; Cyprus; Georgia;

Caspian Sea).

— Redtenbacher, 1874: 242 ( Stenus ; characters; Austria).

— Seidlitz, 1875: 253 ( Stenus ; characters; Baltic region).

— J. Sahlberg, 1876: 52 ( Stenus ; Finland).

— Rey, 1884: 304 [= 1884a: 152] ( Stenus ; subgenus Nestus ; characters; France).

— Fowler, 1888: 341 ( Stenus ; characters; Britain).

— Ganglbauer, 1895: 578 ( Stenus ; subgenus Nestus ; characters; Europe; Mediterranean region).

— Fauvel, 1902b: 75 ( Stenus ; Algeria [ Note: Doubtful locality according to Puthz, 1965c: 74];).

— Montandon, 1908: 78 ( Stenus ; Romania).

— Reitter, 1909: 157 ( Stenus ; subgenus Nestus ; characters; Germany).

— Bernhauer and Schubert, 1911: 156 ( Stenus ; catalog).

— Johansen, 1914: 509 ( Stenus ; subgenus Nestus ; characters; Denmark).

— L. Benick, 1929a: 36 ( Stenus ; subgenus Nestus ; characters in key).

— Portevin, 1929: 398 ( Stenus ; characters; France).

— Scheerpeltz, 1933: 1151 ( Stenus ; subgenus Stenus ; catalog).

— Wüsthoff, 1934a: Tafel II ( Stenus ; illustration of aedeagus).

— L. Benick, 1949: 117 ( Stenus ; characters; Finland; Norway; Germany; Russia; France; Britain;

Austria; Czechoslovakia; Hungary; Romania; Yugoslavia; Italy).

— Kevan and Allen, 1962: 215 ( Stenus ; characters; notes).

— Szujecki, 1961: 31 ( Stenus ; subgenus Nestus ; characters; Poland).

— Palm, 1961a: 97 ( Stenus ; subgenus Nestus ; characters; Sweden; Denmark; Norway; Finland).

— Horion, 1963: 353 ( Stenus ; subgenus Nestus ; Caucasus; west-Siberia; Russia; Fennoscandia;

Britain; Denmark; France to Poland-Russia; Italy; Bosnia).

— Lohse, 1964: 115 ( Stenus ; subgenus Nestus ; characters; central Europe).

— Puthz, 1965c: 74 ( Stenus ; characters; Norway; Sweden; Finland; Russia; England; France;

Italy; Yugoslavia; Siebenbürgen; Transylvania; Caucasus).

— Szujecki, 1968a: 720 ( Stenus ; Poland).

— Korge, 1971a: 6 ( Stenus ; Turkey).

— Puthz, 1972s: 254 ( Stenus ; Yugoslavia).

— Puthz, 1977d: 162 ( Stenus ; synonymic notes).

— Pope, 1977: 26 ( Stenus ; Britain).

— Lucht, 1987: 97 ( Stenus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland;

France; Benelux; Denmark).

— Ryvkin, 1990: 162 ( Stenus ; notes; Georgia).

— Puthz, 1991f: 169 ( Stenus ; Belorussia).

— Dauphin, 1993: 179 ( Stenus ; subgenus Nestus ; characters; France).

— Hromádka, 1993: 44 ( Stenus ; subgenus Nestus ; Czech Republic; Slovakia).

— Puthz and Zanetti, 1995: 17 ( Stenus ; Italy).

— Jelínek, 1999: 156 ( Stenus ; subgenus Nestus ; collecting note; Czech Republic).

- DISTRIBUTION: Europe, Russia, Georgia, Turkey. clavulus Hochhuth, 1849: 174 ( Stenus ; Type locality: Iberien).

— Bernhauer and Schubert, 1911: 179 ( Stenus ; subgenus Hemistenus ; catalog).

— L. Benick, 1929a: 61 ( Stenus ; subgenus Parastenus ; characters in key).

— Scheerpeltz, 1933: 1187 ( Stenus ; subgenus Hemistenus ; catalog).

— Wüsthoff, 1934a: Tafel III ( Stenus ; illustration of aedeagus).

— Puthz, 1977d: 162, 163 ( Stenus ; synonym of circularis ). brunneipalpis Roubal, 1928: 8 ( Stenus ; variety of circularis ; Type locality: Slovakia: Leopoldov).

— Scheerpeltz, 1933: 1152 ( Stenus ; synonym of circularis ).

— L. Benick, 1949: 130 ( Stenus ; subspecies of circularis ).

— Szujecki, 1961: 9 ( Stenus ; aberration of circularis ).

circumflexus Fauvel, 1895: 211 ( Stenus ; Type locality: Bírmanie Carin Asciuii Ghécu , 1400–1500 m; [ Note: See lectotype designation by Puthz, 1969]. Also cited from Bírmanie: Carin Ghécu, 1300–1400 m).

— Bernhauer and Schubert, 1911: 179 ( Stenus ; subgenus Hemistenus ; catalog).

— Cameron, 1930: 386 ( Stenus ; subgenus Mesostenus ; characters; Burma).

— Scheerpeltz, 1933: 1187 ( Stenus ; subgenus Hemistenus ; catalog).

— Puthz, 1969: 33 ( Stenus ; subgenus Parastenus ; lectotype designation: Carin Asciuii Ghécu, 14–1500 m; characters).

— Rougemont, 1981a: 343 ( Stenus ; characters; Burma).

— Rougemont, 1981b: 382 ( Stenus ; Thailand).

— Puthz, 1981a: 19 ( Stenus ; China).

— Puthz, 1981b: 146 ( Stenus ; characters).

— Rougemont, 1983c: 46 ( Stenus ; Thailand).

— Rougemont, 1987: 711 ( Stenus ; subgenus Parastenus ; Thailand).

- DISTRIBUTION: Myanmar, Thailand, China.

cirriformis Naomi, 1988e: 34 ( Stenus ; subgenus Parastenus ; Type locality: Japan: Hiroshima Pref.: Sandankyô).

— Naomi, 1988h: 67 ( Stenus ; characters; Japan).

— Naomi, 1997c: 6 ( Stenus ; characters; Japan).

- DISTRIBUTION: Japan.

cirrus L. Benick, 1940: 561 ( Stenus ; Type locality: N.W. China: Tienmushan ).

— Puthz, 1967s: 245 ( Stenus ; subgenus Parastenus ; China).

- DISTRIBUTION: China.

civicus L. Benick, 1941, see: japonicus Sharp, 1874 .

clainpanaini Bernhauer, 1912b: 136 ( Stenus ; Type locality: Beirut, Bikfaia im Libanon).

— L. Benick, 1929a: 66 ( Stenus ; subgenus Parastenus ; characters in key).

— Scheerpeltz, 1933: 1187 ( Stenus ; subgenus Hemistenus ; catalog).

— Fagel, 1968a: 193 ( Stenus ; Lebanon).

- DISTRIBUTION: Lebanon.

claritarsis Puthz, 1971 v: 100 ( Stenus ; Type locality: Lenkoran. Also cited from Romania,

and Hungary).

— Puthz, 1972s: 264 ( Stenus ; notes).

— Puthz, 1972d: 87 ( Stenus ; subgenus “ Hemistenus ”; characters; Austria; Hungary; Romania; Russia; Caucasus).

— Lohse and Lucht, 1989: 146 ( Stenus ; characters; Slovakia; Neusiedlersee).

— Ryvkin, 1990: 176 ( Stenus ; notes; Georgia; Armenia; Azerbaijan).

— Puthz, 1992c: 152 ( Stenus ; Austria).

— Hromádka, 1993: 45 ( Stenus ; subgenus Hemistenus ; Slovakia).

- DISTRIBUTION: Europe, Russia, Georgia, Armenia, Azerbaijan.

clarkei Rougemont, 1981: 106 ( Stenus ; Type locality: Ethiopia: Sidamo Prov. : 11 km SE of Aghere Selam, 2630 m).

- DISTRIBUTION: Ethiopia.

clarostigma Puthz, 1988b: 135 ( Stenus ; subgenus Stenus ; Type locality: Canada: Alberta: Waterton National Park, Rowe Creek, 5460 feet. Also cited from Idaho and Montana). — Campbell and Davies, 1991: 110 ( Stenus ; Canada).

- DISTRIBUTION: Canada, USA.

clavicornis Scopoli, 1763: 100 ( Staphylinus ; Type locality: Carniolia). — Villers, 1789: 419 ( Staphylinus ; characters; Germany).

— Gmelin, 1790: 2029 ( Staphylinus ; characters).

— Turton, 1802: 513 ( Staphylinus ; characters; Kiel).

— Marsham, 1802: 527 ( Staphylinus ; characters; Britain).

— Ljungh, 1804: 61 ( Stenus ; [ Note: Misidentification: Bernhauer and Schubert, 1911: 156 cited cicindeloides Ljungh as a synonym of clavicornis ; Ljungh did not describe the species as new, he attributed it to Schaller]; characters; Sweden).

— Gyllenhal, 1810: 469 ( Stenus ; [ Note: Misidentification: Bernhauer and Schubert, 1911: 156, cited boops Gyllenhal as a synonym of clavicornis Scopoli ; Gyllenhal did not describe the species as new, he attributed it to Gravenhorst]; characters; Sweden).

— Curtis, 1827: pl. 164 ( Stenus ; England).

— Erichson, 1839a: 545 ( Stenus ; synonym of speculator ).

— Erichson, 1839a: 545 ( Stenus ; [ Note: Cited as cicindeloides Ljungh , but Ljungh’s use was a misidentification]; synonym of speculator ).

— Erichson, 1840: 706 ( Stenus ; synonym of speculator ).

— Erichson, 1840: 706 ( Stenus ; [ Note: Cited as cicindeloides Ljungh , but Ljungh’s use was a misidentification]; synonym of speculator ).

— Dufour, 1843: 36 ( Stenus ; characters; France).

— Kraatz, 1857d: 762 ( Stenus ; [ Note: Cited as cicindeloides Ljungh , but Ljungh’s use was a misidentification]; synonym of speculator ).

— Fauvel, 1873: 84 [= 1873a: 242] ( Stenus ; characters; France; Europe; Caucasus; Siberia).

— Fauvel, 1874: 331 [= 1874b: 37] ( Stenus ; Turkey).

— J. Sahlberg, 1876: 48 ( Stenus ; synonym of speculator ).

— Rey, 1884: 234 [= 1884a: 82] ( Stenus ; [ Note: Cited as boops Gyllenhal ]; characters; France).

— Eppelsheim, 1893: 55 ( Stenus ; Baikal).

— Ganglbauer, 1895: 564 ( Stenus ; characters; Palaearctic region).

— Ganglbauer, 1895: 564 ( Stenus ; [ Note: Cited as cicindeloides Ljungh , but Ljungh’s use was a misidentification]; synonym of clavicornis ).

— Reitter, 1909: 156 ( Stenus ; characters; Germany).

— Bernhauer and Schubert, 1911: 156 ( Stenus ; catalog).

— Bernhauer and Schubert, 1911: 156 ( Stenus ; [ Note: Cited as cicindeloides Ljungh , but Ljungh’s use was a misidentification]; synonym of clavicornis ).

— Johansen, 1914: 492 ( Stenus ; subgenus Stenus ; characters; Denmark).

— L. Benick, 1922c: 161 ( Stenus ; Siberia).

— L. Benick, 1929a: 30 ( Stenus ; subgenus Stenus ; characters in key).

— Portevin, 1929: 396 ( Stenus ; characters; France).

— Cameron, 1930: 327 ( Stenus ; characters; India [ Note: Probably an erroneous locality according to Puthz (personal communication)]).

— Cameron, 1930: 328 ( Stenus ; [ Note: Cited as cicindeloides Ljungh , but Ljungh’s use was a misidentification]; synonym of clavicornis ).

— Scheerpeltz, 1933: 1152 ( Stenus ; subgenus Stenus ; catalog).

— Wüsthoff, 1934a: Tafel II ( Stenus ; illustration of aedeagus).

— West, 1937: 42 ( Stenus ; Faroe Islands).

— Szujecki, 1961: 25 ( Stenus ; subgenus Stenus ; characters; Poland).

— Palm, 1961a: 78 ( Stenus ; subgenus Stenus ; characters; Sweden; Denmark; Norway; Finland).

— Horion, 1963: 323 ( Stenus ; subgenus Stenus ; widespread in Europe).

— Lohse, 1964: 113 ( Stenus ; subgenus Stenus ; characters; central Europe).

— Smetana, 1964d: 59 ( Stenus ; subgenus Stenus ; Czechoslovakia).

— Raianu, 1964: 105 ( Stenus ; Romania).

— Puthz, 1967: 3 ( Stenus ; subgenus Stenus ; Portugal).

— Puthz, 1967m: 77 ( Stenus ; Mongolia).

— Puthz, 1967o: 230 ( Stenus ; Mongolia).

— Szujecki, 1968a: 719 ( Stenus ; Poland).

— Bordoni, 1970: 23 ( Stenus ; ecological notes; Italy).

— Puthz, 1971j: 138 ( Stenus ; Mongolia).

— Puthz, 1974k: 435 ( Stenus ; Korea).

— Bordoni, 1975h: 7 ( Stenus ; distribution; Italy).

— Puthz, 1975f: 124 ( Stenus ; Quebec).

— Pope, 1977: 26 ( Stenus ; Britain).

— Puthz, 1983e: 348 ( Stenus ; notes; Caucasus: Georgia; Daghestan).

— Anderson, 1984: 245 ( Stenus ; collecting notes; Ireland).

— Yuh, Paik, Kwon, and Lee, 1985: 230 ( Stenus ; Korea).

— Lucht, 1987: 96 ( Stenus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Naomi, 1988g: 58, 61 ( Stenus ; subgenus Stenus ; characters; Japan).

— Lohse and Lucht, 1989: 140 ( Stenus ; characters).

— Nowosad, 1990: 63 ( Stenus ; collected from mole nest; notes; Poland).

— Ryvkin, 1990: 154 ( Stenus ; notes; Russia; Georgia; Armenia; Azerbaijan).

— Whitehead, 1990d: 9 ( Stenus ; characters; Britain).

— Puthz, 1991f: 167 ( Stenus ; Belorussia).

— Dauphin, 1991: 286 ( Stenus ; subgenus Stenus ; characters; France).

— Campbell and Davies, 1991: 110 ( Stenus ; Canada).

— Welch, 1993: 228 ( Stenus ; ovariole number and ovary structure).

— Hromádka, 1993: 44 ( Stenus ; subgenus Stenus ; Slovakia; Czech Republic).

— J. Li, 1993: 24 ( Stenus ; China).

— Puthz, 1994j: 223 ( Stenus ; New Hampshire; Pennsylvania).

— Puthz and Zanetti, 1995: 17 ( Stenus ; Italy).

— Bordoni, 1995: 78 ( Stenus ; ecological notes; Italy).

— Betz, 1996: 25 ( Stenus ; labial adhesion capture apparatus).

— Jelínek, 1999: 154 ( Stenus ; collecting note; Czech Republic).

— Assing, 2001: 75 ( Stenus ; collected from xerothermous site; Germany).

- DISTRIBUTION: Europe, Russia, Georgia, Armenia, Azerbaijan, Turkey, Mongolia, China, Korea, Japan, Canada, USA.

buphthalmus Schrank, 1776: 72 ( Staphylinus ; cited as buphthalmos ; [ Note: The original spelling differs from that currently used but most authors have used buphthalmus rather than buphthalmos and have attributed the name to Schrank; the prevailing use of this incorrect subsequent spelling is maintained by application of Article 33.3.1]; Type locality: Not cited).

— Goeze, 1777: 723 ( Staphylinus ; characters).

— Schrank, 1781: 235 ( Staphylinus ; cited as buphthalmos ; characters; Austria).

— Villers, 1789: 424 ( Staphylinus ; characters).

— Gmelin, 1790: 2032 ( Staphylinus ; [ Note: Page misnumbered as 1032]; characters).

— Schrank, 1798: 643 ( Staphylinus ; characters; notes).

— Turton, 1802: 516 ( Staphylinus ; characters; Austria).

— Gravenhorst, 1806: 230 ( Stenus ; characters).

— Panzer, 1808: 16 ( Stenus ; characters; illustrations; Note: The clavicornis that Panzer cited in the first edition of Fauna Germanica is cited in 1808 as the same as buphthalmus ).

— Zetterstedt, 1828: 88 ( Stenus ; characters; Lapland).

— Mannerheim, 1830: 43 ( Stenus ; Finland; Sweden).

— Mannerheim, 1831: 457 ( Stenus ; Finland; Sweden).

— Zetterstedt, 1838: 70 ( Stenus ; characters; habitat; Lapland).

— Erichson, 1839a: 545 ( Stenus ; synonym of speculator ).

— Heer, 1839: 218 ( Stenus ; characters; Switzerland).

— Erichson, 1840: 706 ( Stenus ; synonym of speculator ).

— Kraatz, 1857d: 752 ( Stenus ; synonym of speculator ).

— Seidlitz, 1875: 255 ( Stenus ; characters; Baltic region).

— Rey, 1884: 234 [= 1884a: 82] ( Stenus ; synonym of boops ).

— Ganglbauer, 1895: 564 ( Stenus ; synonym of clavicornis ).

— Sainte-Claire Deville, 1906: 82 ( Stenus ; Corsica).

— Bernhauer and Schubert, 1911: 156 ( Stenus ; catalog).