Clusiodes, Coquillett

Owen Lonsdale, 2017, World Catalogue of the Druid Flies (Diptera: Schizophora: Clusiidae), Zootaxa 4333 (1), pp. 1-85 : 21-29

publication ID

https://doi.org/ 10.11646/zootaxa.4333.1.1

publication LSID

lsid:zoobank.org:pub:47AB9D4D-16FF-4A7E-B73F-2A65C2841FF4

DOI

https://doi.org/10.5281/zenodo.4752261

persistent identifier

https://treatment.plazi.org/id/103F3650-BF6F-FF8E-FF0C-FF11DB61FA92

treatment provided by

Plazi

scientific name

Clusiodes
status

 

Genus CLUSIODES Coquillett View in CoL

Heteroneura Fallén 1823: 2 [preoccupied by Heteroneura Fallén 1810 ) ( Platypezidae View in CoL )]. Type species, Heteroneura albimana Meigen, 1830 , subsequent designation by ICZN 2010: 344 (Opinion #2262). Gender feminine. Meigen 1830: 126 [redescription, listed in error as independent proposal in Neave (1939b: 640)]; Curtis 1837: column 274 [checklist, Britain], 1838: plate 721 [ albimana Meigen 1830 designated as type species, redescription, diagnosis, Britain]; Haliday 1838: 188 [Britain]; Zetterstedt 1838: column 784 [redescription, designation of Heteroneura muscaria Fallén 1823 (= Cnemacantha muscaria (Fallén) View in CoL , Lauxaniidae View in CoL ) as type species], 1848: 2787 [ Heteroneura geomyzinus Fallén 1823 designated as type species, redescription], 1852: 4816 [Scandinavia]; Westwood 1840: 152 [ Heteroneura albimana Meigen 1830 listed as type species, diagnosis, Britain]; Agassiz 1846: 519 [nomenclator, attributed to Meigen (1830)]; Rossi 1849: 78 [checklist, Austria]; Wilson 1855: 256 [description]; Loew 1862: 42 [catalogue, North America]; Osten Sacken 1878: 197 [catalogue, North America]; Schiner 1862: 37 [redescription, key to species, Austria], 1864: 72 [catalogue, Europe]; Siebke 1877: 181 [catalogue, Norway]; Osten Sacken 1878: 197 [catalogue, North America]; Williston 1896: 386 [key, St. Vincent], 1908: 319 [key, illustrations of head and wing]; Strobl 1897: 27 [catalogue, Transylvania]; Becker 1902: 313 [reexamination of Meigen types], 1905: 38 [catalogue, Palaearctic]; Czerny 1903: 73 [family and genus revision]; Aldrich 1905: 571 [catalogue, North America]; Kertész 1906: 320 [key]; Hendel 1910: 309 [nomenclature]; Schulze et al. 1930: 1540 [nomenclator]; Neave 1939b: 640 [nomenclator]; Lonsdale 2009a: 144 [proposal to ICZN]; ICZN 2010: 344 (Opinion #2262) [designation of H. albimana Meigen, 1830 as type species]).

Heteronevra . Macquart 1835: 587 [diagnosis, key]. Unjustified emendation of Heteroneura Fallén 1823 . Blanchard 1840: 628 [ Heteroneura Nubila Meigen (= albimana Meigen ) designated as type species]; Scudder 1882: 161 [nomenclator], 1884: 148 [nomenclator]; Schulze et al. 1930: 1540 [nomenclator]; Evenhuis et al. 2016: 70 [nomenclatural status, as nomen oblitum, junior synonym of Clusiodes View in CoL ].

Clusiodes Coquillett 1904: 93 View in CoL [replacement name for Heteroneura Fallén 1823 ]. Type species, Heteroneura albimanus Meigen 1830 , automatic designation. Gender masculine. Coquillett 1910: 525 [type species catalogue, North America]; Johnson 1913b: 97 [family and genus revision, eastern United States], 1925a: 245 (as genus and subgenus) [catalogue, New England]; Wahlgren 1917: 186 [review, Sweden]; Malloch 1918: 5 [key to species], 1922: 47 (as genus and subgenus) [key to subgenera and species]; Melander & Argo 1924: 13 [family and genus revision, key to subgenera and species]; Czerny 1928: 6 (as genus and subgenus) [family and genus revision, Palaearctic]; Schulze et al. 1927b: 730 [nomenclator]; Stackelberg 1933: 381 [key, European USSR], 1988: 505 [key, European USSR]; Tuomikoski 1933: 15 [Finland]; Séguy 1934: 351 [France]; Neave 1939a: 770 [nomenclator]; Kloet & Hincks 1945: 394 [checklist, Britain]; Hennig 1952: 227 [immature stages]; Frey 1960: 18 [family and genus revision, key, Indoaustralia, catalogue, world]; McAlpine 1960: 81 [family and genus revision, Australia—all species presently recognized as Hendelia View in CoL ]; Trojan 1962: 22 (as genus and subgenus) [key to subgenera and species, Poland]; Soós 1964a: 7 [key], 1984: 11 (as genus and subgenus) [catalogue, Palaearctic], 1987: 857 [key, Nearctic]; Steyskal 1965: 807 (as genus and subgenus) [catalogue, America north of Mexico]; Cole & Schlinger 1969: 426 [illustration of head, key to genera of Clusiidae View in CoL , western North America]; Sasakawa 1977: 236 (as genus and subgenus) [catalogue, Orient], 1987: 100 (as genus and subgenus) [revision, Orient/Pacific], 1998b: 223 [key to subgenera] 2011b: 20 [Oriental fauna]; Stubbs 1982: 89 (as genus and subgenus) [identification guide, biology, Britain]; Ferrar 1987: 121 [notes on biology]; Balthazar 1990: 416 [catalogue, Philippines, only included species Hendelia aberrans (Frey) ]; Woznica 1991: 208 [checklist, Poland]; DeBruyn 1991: 146 [catalogue, Belgium]; Caloren & Marshall 1998: 263 [redescription, biology, New world revision, phylogeny, synonymy of Labomyia Frey (now synonymized with Hendelia View in CoL ), Clusiaria Malloch and Columbiella Malloch , illustration of male and female genitalia, photos]; Marshall 2000: 4 [synonymy of Parahendelia McAlpine ]; Beuk 2002: 271 (as genus and subgenus) [checklist, Netherlands]; Greve 2005: 3 [Norway]; Sueyoshi 2006: 6 [Japan], 2014: 618 [catalogue, Japan]; Papp et al. 2006: 190 [identified and unidentified species, Thailand]; Lonsdale & Marshall 2007b: 123 [redefinition, maps, review, phylogeny, biology, taxonomic history, zoogeography, data on Nearctic species, discussion of C. melanostomus complex, illustration of immature stages, Old world, incorrect treatment of Gioenia Lioy as synonym], 2010: 1044 [Manual of Central American Diptera View in CoL ]; Lonsdale et al. 2010: 267 [family revision, key]; ICZN 2010: 344 [conservation of Clusiodes View in CoL in Clusiidae View in CoL by setting aside all previous type fixations and by designating Heteroneura albimana Meigen 1830 as type species for Heteroneura Fallén 1823 ]; Kahanpää 2014: 288 [checklist, Finland]; Evenhuis et al. 2016: 70 [nomenclatural status, nomen protectum].

Clusiaria Malloch 1922: 49 (as subgenus) [key to subgenera and species, notes]. Type species, Clusiodes duplicata Malloch 1922 View in CoL , by original designation (= melanostomus Loew ). Gender feminine. Johnson 1925a: 245 [catalogue, New England]; Czerny 1928: 6 [key, Palaearctic]; Neave 1939a: 770 [nomenclator]; Kloet & Hincks 1945: 394 [checklist, Britain]; Frey 1960: 16 [key to subgenera and species, Indoaustralia, catalogue, world]; Trojan 1962: 22 [key, Palaearctic]; Stubbs 1982: 89 [key, Britain]; Soós 1984: 12 [catalogue, Palaearctic]; Sasakawa 1987: 101 [key, Orient/Pacific]; Schumann et al. 1999: 144 [checklist, Germany]; Caloren & Marshall 1998: 263 [as synonym of Clusiodes View in CoL ]; Beuk 2002: 271 [checklist, Netherlands].

Columbiella Malloch 1922: 49 [as subgenus]. Type species, Clusiodes apiculata Malloch 1922 , by original designation. Gender feminine. Johnson 1925a: 246 [catalogue, New England]; Czerny 1928: 9 [Palaearctic]; Neave 1939a: 806 [nomenclator]; Kloet & Hincks 1945: 394 [checklist, Britain]; Frey 1960: 26 [catalogue, world]; Trojan 1962: 26 [key, Poland]; Steyskal 1965: 807 [catalogue, America north of Mexico]; Stubbs 1982: 89 [key, Britain]; Soós 1984: 12 [catalogue, Palaearctic]; Sasakawa 1987: 101 [key, Orient/Pacific]; Caloren & Marshall 1998: 263 [as synonym of Clusiodes View in CoL ]; Beuk 2002: 271 [checklist, Netherlands].

Clusioides . Misspelling of Clusiodes View in CoL . Curran 1934b: 363 [key]; Soós 1981: 97 [ Hungary, key]. Clusinodes. Misspelling of Clusiodes View in CoL . Craighead 1950: 541 [as subgenus within Sobarocephala View in CoL ].

albimanus (Meigen) . PA: Europe, Russia.

Heteroneura geomyzina var. b Fallén 1823: 3. Sweden. ST ♂♀ [lost]. Zetterstedt 1838: 790 [redescription]; Zetterstedt 1848: 2787 [as synonym of albimana Meigen ].

Heteroneura albimana Meigen 1830: 128 . Switzerland. Geneva Canton: Bernex, 415m. NT ♂ MHNG [designated by Lonsdale & Marshall 2007b: 129]. Curtis 1837: column 274 [checklist, Britain], 1838: plate 721 [designated as genus type, synonymy of scotica Curtis , illustrations of head, habitus, Britain], 1855: 6403 [bred from rotten willows]; Boie 1847: 331 [discussion of immature stages]; Zetterstedt 1848: 2790 [synonymy of geomyzina var. b Fallén and geomyzina var. c Zetterstedt, Sweden], 1855: 4817 [Sweden]; Curtis 1855: 4603 [notes on insect economy]; Rossi 1849: 78 [checklist, Austria]; Wulp & Snellen van Vollenhoven 1856: 114 [checklist, Netherlands]; Loew 1857: 51 [Germany], 1864b: 339 [synonymy of albimana var b Zetterstedt, redescription, discussion, Germany]; Schiner 1862: 38 [redescription, key, biology, Austria], 1864: 72 [catalogue, Europe]; Brauer 1883: 92 [larva in beetle burrows in wood, records of Vienna Museum]; Girschner 1888: 97 [comparison to alpina Loew (= geomyzinus Fallén )]; Becker 1902: 313 [examination of Meigen types], 1905: 38 [catalogue, Palaearctic]; Czerny 1903: 81 [redescription and biology]; Czizek 1910: 112 [catalogue, Germany]; Haliday 1933: 150 [England]; Lonsdale 2009a: 144 [proposed as type species]; Lonsdale et al. 2010: 240 [genes sequenced for phylogenetic analysis]; Lonsdale & Marshall 2010: 1042 [reproduction of figure from Soós (1987)]; ICZN 2010: 344 [conservation of Clusiodes View in CoL in Clusiidae View in CoL by setting aside all previous type fixations and by designating Heteroneura albimana Meigen, 1830 as type species of Heteroneura Fallén 1823 ].

Heteroneura nubila Meigen 1830: 127 View in CoL . Europe [not given]. ST ♂ MNHNP [lost, probably destroyed]. Rossi 1849: 78 [checklist, Austria]; Wilson 1855: 257 [description, records from Austria and Europe]; Loew 1857: 53 [ Germany], 1864b: 343 [as junior synonym of Heteroneura ruficollis Meigen 1830 ]; Becker 1902: 313 [examination of Meigen material], 1905: 39 [catalogue, Palaearctic]; Czerny 1903: 76 [redescription].

Heteronevra nubile . Macquart 1835: 587 [illustration of head, France].

Heteronevra albimana . Macquart 1835: 588 [ France].

Heteroneura scotica Curtis 1837 : column 274 [nomen nudum, Britain]; Curtis 1838: pl. 101 [as synonym of albimana Meigen, Britain ].

Agromyza geomyzina . Misidentification. Zetterstedt 1838: 790 [ Sweden].

Agromyza geomyzina var c Zetterstedt 1838: 790. Sweden. ST ♀ [lost - not found in ZIN]. Zetterstedt 1848: 2787 [as synonym of albimana Meigen ].

Heteronevra Nubila. Blanchard 1840: 628 [designated as type of Heteroneura Fallén 1823 , France].

Heteroneura albimana var b Zetterstedt 1852: 4817. Sweden. Lund. ST ♀ [lost]. Loew 1864b: 339 [as synonym of albimana Meigen ].

Heteroneura albimana var. pallidior Loew 1857: 52 [as “ albimana var. α”]. Germany. Type information unknown. Loew 1864b: 340 [includes albimana sensu Meigen (1830) , Zetterstedt (1848) and Schiner (1862), synonymy of albimana var. α Loew]; Czerny 1903: 81 [suggests synonymy of varieties]; Lonsdale & Marshall 2007b: 129 [as synonym of albimana Meigen ].

Heteroneura albimana var. obscurior Loew 1857: 52 [as “ albimana var. β”]. Germany. Type information unknown. Loew 1864b: 339 [includes albimana var. b Fallén and pictipes Zetterstedt sensu Schiner (1862) View in CoL , synonymy of albimana var. β Loew]; Czerny 1903: 81 [suggests synonymy of varieties]; Lonsdale & Marshall 2007b: 129 [as synonym of albimana Meigen ].

Heteroneura pictipes View in CoL . Misidentification. Schiner 1862: 37 [Moravia].

Heteroneura albimanus View in CoL . Aldrich 1905: 570 [gender disagreement, catalogue, North America].

Clusiodes (Clusiodes) albimana . Johnson 1913b: 98 [generic combination, Nearctic records - misidentification of johnsoni Malloch View in CoL ]; Melander & Argo 1924: 15 [includes forms pallidior Loew and obscurior Loew , photo, notes type likely destroyed]; Frey 1928c: 109 [catalogue, Finland], 1960: 26 (in part) [catalogue, world]; Kloet & Hincks 1945: 394 [checklist, Britain]; Trojan 1962: 25 [key, illustrations of male genitalia, Poland]; Chandler 1978: 264 [ Ireland]; Hackman 1980: 148 [checklist, Finland]; Stubbs 1982: 92 [Britain]; Greve & Midtgaard 1986: 89 [ Norway].

Clusiodes albimana . Wahlgren 1917: 187 [diagnosis, key, photo]; Czerny 1928: 8 [redescription, illustrations of thorax, wing, head and male genitalia]; Stackelberg 1933: 381 [key, European USSR], 1988: 505 [key, European USSR]; Séguy 1934: 352 [illustrations of head and male genitalia, photo of wing, France]; Richards 1934: 2 [puparia in rotting birch trunk]; Tuomikoski 1936: 184 [Finland]; Hennig 1952: 227 [immature stages]; Chandler 1973: 340 [associated with elm, England]; Withers 1985: 63 [bred from Fraxinus excelsior View in CoL , associated with Fagus sylvaticus , England]; Greve & Midtgaard 1986: 89 [Norway]; Ferrar 1987: 122 [immature stages, illustration of larva and puparium]; Zuska 1987: 235 [Czech Republic]; Smith 1989: 103 [reared from old logs and mossy tree stumps, illustrations of puparium, Britain]; Dunk 1996: 422 [catalogue, Bavaria]; Yagi & Sawada 2000: 41 [Mamaev collection holdings].

Clusiodes (Clusiodes) albimana form pallidior. Melander & Argo 1924: 15 [in part, misidentification in part of johnsoni Malloch View in CoL ].

Clusiodes (Clusiodes) albimana form obscurior . Melander & Argo 1924: 15 [in part].

Clusiodes (Clusiaria) nubila View in CoL . Melander & Argo 1924: 21 [synonymy, generic combination, distribution, comparison to ruficollis Meigen ]; Czerny 1928: 7 [redescription, central Europe]; Frey 1960: 26 [catalogue, world]; Trojan 1962: 22 [key, Poland]; Soós 1984: 12 [catalogue, Palaearctic]; Schumann et al. 1999: 144 [checklist, Germany].

Clusiodes nubila View in CoL . Séguy 1934: 352 [illustration of wing, France]; Ceianu 1989: 29 [expected in Romania].

Clusiodes (Columbiella) albimanus View in CoL . Steyskal 1965: 807 [catalogue, America north of Mexico].

Clusioides (Clusioides) albimana . Soós 1981: 99 [ Hungary, key, illustration of habitus, male terminalia].

Clusioides (Columbiella) nubila . Soós 1981: 102 [possible occurrence in Hungary, key].

Clusiodes (Clusiodes) albimanus View in CoL . Soós 1984: 12 [catalogue, Palaearctic (Europe, including Ukraine)]; Roháček 1995: 126 [biology and habitat, male behaviour, Czech and Slovak Republics]; Schumann et al. 1999: 144 [checklist, Germany]; Beuk 2002: 271 [checklist, Netherlands]; Greve 2005: 3 [ Norway].

Clusiodes (Clusiodes) albima View in CoL . Misspelling of albimana . Greve & Midtgaard 1986: 89.

Clusiodes albimanus View in CoL . Krištofík & Zuska 1986: 165 [ Slovakia]; Soós 1987: 855 [illustration of head]; Ceianu 1989: 29 [adults in boggy forests, catalogue, Romania]; Woznica 1991: 208 [checklist, Poland]; DeBruyn 1991: 146 [catalogue, Belgium]; Poole 1996: 126 [checklist, Nearctic]; Barrientos et al. 1996: 118 [biology, Spain]; Merz 1998: 276 [checklist, Switzerland]; Roháček & Barták 2001: 338 [checklist, Czech Republic]; Carles-Tolrá 2002: 155 [catalogue, Spain]; Roháček et al. 2005: 279 [catalogue, habitat, Czech Republic]; Barták & Kubík 2005: 279 [ Czech Republic]; Lonsdale & Marshall 2007b: 129 (synonymy of nubila Meigen and varieties pallidior and obscurior , maps, key, notes, neotype, phylogeny, illustrations of genitalia, photo, new records ( France, Greece, Netherlands, Russia, Switzerland)]; Roháček 2008: 154 [ Italy]; 2009: 218 [ Slovakia], 2012: 99 [central Slovakia], 2013: 130 [central Slovakia]; Nilsson 2009: 5 [ Sweden]; Rotheray & Horsfield 2013: 403 [illustrations of immatures, photo of puparium; rearing data— Alnus View in CoL , Betula View in CoL , Fagus View in CoL , Fraxinus View in CoL , Populus View in CoL , Salix ; England, Finland, Scotland]; Kahanpää 2014: 288 [checklist, Finland]; Roháček et al. 2016: 158 [ Portugal].

Clusiodes (Clusiodes) albimanus View in CoL . Roháček 1997: 75 [checklist, Czech and Slovak republics]; Schacht 2004: 276 [ Germany].

americanus Malloch. View in CoL NE: Northeast North America (Ontario to Quebec and Illinois to South Carolina).

Clusiodes americana Malloch 1922: 48 . USA. Maryland: Prince Georges Co., Beltsville. HT ♂ unknown [female allotype with “ Type ” label in USNM].

Clusiodes (Clusiodes) americana . Melander & Argo 1924: 16 [redescription, new records (New York, Pennsylvania, Virginia)]; Johnson 1925a: 245 [catalogue, New England (Maine, New Hampshire, Vermont)]; Frey 1960: 26 [catalogue, world].

Clusiodes (Columbiella) americanus View in CoL . Steyskal 1965: 807 [catalogue, America north of Mexico].

Clusiodes americanus View in CoL . Poole 1996: 126 [checklist, Nearctic]; Caloren & Marshall 1998: 271 [redescription, illustrations of wing and male and female genitalia, photo, new records (northeastern North America)]; Marshall, 2006: 510 [photo]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis].

apicalis (Zetterstedt) . PA: Europe, western Russia.

Heteroneura geomyzina var. apicalis Zetterstedt 1848: 2789 . Denmark. “ Dania ”. Type information unknown [ MZLU].

Heteroneura apicalis View in CoL . Czerny 1903: 79 [new status, redescription, illustrations of head and wing]; Becker 1905: 39 [catalogue, Palaearctic].

Clusiodes apicalis View in CoL . Wahlgren 1917: 187 [diagnosis, key]; Malloch 1918: 6 [Nearctic record - likely ater Melander & Argo View in CoL ]; Stackelberg 1933: 381 [key, European USSR], 1988: 505 [key, European USSR]; Tuomikoski 1933: 18 [ Finland], 1936: 183 [ Finland]; Séguy, 1934: 352 [ France]; Greve et al. 1987: 78 [ Norway]; Ceianu 1989: 29 [catalogue, Romania]; Poole 1996: 126 [checklist, Nearctic]; Dunk 1996: 422 [catalogue, Bavaria]; Yagi & Sawada 2000: 41 [Mamaev collection holdings]; Roháček 2005: 326 [listed as endangered in Czech Republic]; Lonsdale & Marshall 2007b: 135 [key, map, synonymy, notes, phylogeny, illustrations of genitalia, new records ( Finland, Norway, Russia)]; Rotheray & Horsfield 2013: 404 [illustrations of immatures; rearing data— Alnus View in CoL , Betula View in CoL , Populus View in CoL ; Finland, Russia, Scotland]; Kahanpää 2014: 288 [checklist, Finland].

Clusiodes (Clusiaria) apicalis View in CoL . Melander & Argo 1924: 17 [misidentification—likely ater Melander & Argo View in CoL , redescription, Idaho]; Czerny 1928: 6 [redescription, Europe]; Frey 1928c: 109 [ Finland, catalogue], 1960: 26 [catalogue, world]; Soós 1943: 72 [ Hungary], 1984: 12 [catalogue, Palaearctic ( Denmark, Finland, Britain, northern European USSR)]; Collin 1953b: 139 [ England]; Trojan 1962: 24 [key, illustrations of head and wing, Poland]; Hackman 1980: 148 [checklist, Finland]; Stubbs 1982: 92 [Britain]; Greve & Midtgaard 1986: 89 [biology, Betula verrucosa View in CoL and Salix spp. as likely hosts, Norway]; Roháček 1995: 128 [Czeck Republic]; Schumann et al. 1999: 144 [checklist, Germany]; Greve 2005: 5 [ Norway].

Clusiodes (Clusiodes) apicalis View in CoL . Steyskal 1965: 807 [catalogue, America north of Mexico].

Clusioides (Columbiella) apicalis . Soós 1981: 102 [possible occurrence in Hungary, key, illustration of wing].

Clusiodes (Clusiaria) pictipes View in CoL . Misidentification. Máca 1982: 71 [misidentification det. Roháček (1995), thorax with Trombidium cf. mediterraneum (Berlese) mites attached].

apiculatus Malloch. View in CoL NE: Northeastern North America (Illinois to Nova Scotia and Tennessee to South Carolina).

Clusiodes (Columbiella) apiculata Malloch 1922: 49 . USA. New Hampshire: Hillsborough Co., Milford. HT ♂ MCZ [Type No. MCZT 27014]. Melander & Argo 1924: 23 [redescription, North Carolina]; Johnson 1925a: 246 [catalogue, Vermont].

Clusiodes (Columbiella) nigripalpis View in CoL . Misidentification in part. Malloch 1922: 49 [misidentification of paratypes, det. Caloren & Marshall (1998)].

Clusiodes apiculata . Brimley 1938: 378 [North Carolina];

Clusiodes (Clusiodes) apiculata . Frey 1960: 26 [catalogue, world].

Clusiodes (Columbiella) apiculatus View in CoL . Steyskal 1965: 807 [catalogue, America north of Mexico].

Clusiodes apiculatus View in CoL . Poole 1996: 126 [checklist, Nearctic]; Caloren & Marshall 1998: 269 [redescription, photo, illustrations of male and female genitalia, new records (northeastern North America)]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis].

ater Melander & Argo. View in CoL NE: Widespread in North America north of Mexico (outside of the southeastern USA).

Clusiodes (Clusiaria) atra Melander & Argo 1924: 18 View in CoL . USA. Massachusetts: Franklin Co., Greenfield. HT ♂ USNM. Johnson 1925a: 245 [catalogue, New England]; Frey 1960: 26 [catalogue, world].

Clusiodes (Clusiaria) nitida var. scutellata Melander & Argo 1924: 20 View in CoL . USA. Colorado: Boulder Co., Boulder. HT ♀ USNM [Cat. No. 26264]. Caloren & Marshall 1998: 291 [as “misidentification”, synonymy with atra Melander & Argo View in CoL ].

Clusiodes atra View in CoL . Johannsen 1928: 844 [catalogue, New York].

Clusiodes (Clusiodes) ater View in CoL . Steyskal 1965: 807 [catalogue, America north of Mexico].

Clusiodes (Clusiodes) nitidus var. scutellatus . Steyskal 1965: 807 [catalogue, America north of Mexico].

Clusiodes scutellatus . Poole 1996: 126 [checklist, Nearctic, as synonym of nitidus Melander & Argo ].

Clusiodes ater View in CoL . Poole 1996: 126 [checklist, Nearctic]; Caloren & Marshall 1998: 291 [redescription, phylogeny, photo, illustrations of wing and male and female genitalia, new records (North America)]; Marshall, 2006: 510 [photo]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis, photo]; Lonsdale et al. 2010: 240 [genes sequenced for phylogenetic analysis].

caestus Caloren & Marshall. View in CoL NE: Western North America (Alberta, New Mexico, Utah), Nova Scotia and Ontario.

Clusiodes caestus Caloren & Marshall 1998: 289 View in CoL . Canada. Ontario: Fathom Five National Park, Cove Island. HT ♂ DEBU. Caloren & Marshall 1998: 289 [illustrations of wing and male and female genitalia]; Lonsdale & Marshall 2007b: 146 [map, discussion of C. melanostomus complex, phylogenetic analysis].

caledonicus (Collin) View in CoL . PA: Europe, western Russia.

Heteroneura caledonica Collin 1912: 106 [illustration of male external genitalia]. United Kingdom. Scotland: Inverness; England: Golspie. ST 9♂ 22♀ BMNH, CUMZ, OUMNH.

Clusiodes (Clusiodes) caledonica . Melander & Argo 1924: 17 [generic combination, summary of original description]; Czerny 1928: 9 [redescription, reillustration from original description]; Kloet & Hincks 1945: 394 [checklist, Britain]; Frey 1960: 26 [catalogue, world]; Trojan 1962: 26 [key, illustration of male genitalia, Poland]; Hackman 1980: 148 [checklist, Finland]; Stubbs 1982: 92 [Britain]; Soós 1984: 12 [catalogue, Palaearctic (Britain, Germany, Austria)]; Greve & Midtgaard 1986: 90 [ Norway]; Greve & Midtgaard 1986: 90 [ Norway]; Roháček 1995: 128 [biology, Czech and Slovak Republics]; Schumann et al. 1999: 144 [checklist, Germany]; Beuk 2002: 271 [checklist, Netherlands]; Greve 2005: 4 [ Norway].

Clusiodes caledonica . Tuomikoski 1936: 184 [reared from rotten aspen, adults on birch logs, illustrations of male genitalia, Finland]; Speight & Cogan 1979: 402 [Ireland]; Stackelberg 1988: 505 [key, European USSR].

Clusioides (Clusioides) caledonica . Soós 1981: 100 [possible occurrence in Hungary, key, illustrations of wing, male terminalia].

Clusiodes caledonicus View in CoL . Ceianu 1989: 29 [expected in Romania]; Merz 1998: 276 [checklist, Switzerland]; Roháček & Barták 2001: 338 [checklist, Czech Republic]; Roháček et al. 2005: 280 [catalogue (considered endangered in Czech Republic), habitat, Czech Republic]; Roháček 2005: 326 [listed as endangered in Czech Republic], 2009: 218 [ Slovakia], 2012: 98 [central Slovakia], 2013: 128 [central Slovakia]; Barták & Kubík 2005: 280 [ Czech Republic]; Lonsdale & Marshall 2007b: 131 [synonymy, map, key, phylogeny, notes, illustrations of genitalia, new records ( Finland, Greece, Italy, Switzerland, Sweden, Spain)]; Nilsson 2009: 5 [ Sweden]; Rotheray & Horsfield 2013: 404 [illustrations of immatures, photo of head skeleton in puparium; rearing data ( Pinus View in CoL ), Scotland]; Kahanpää 2014: 288 [checklist, Finland]; Chandler 2015: 212 [ England]; Roháček et al. 2016: 158 [ Portugal].

Clusiodes calcedonicus . Misspelling of caledonicus View in CoL . DeBruyn 1991: 146 [ Belgium].

Clusiodes (Clusiodes) caledonicus View in CoL . Roháček 1997: 75 [checklist, Czech and Slovak republics].

clandestinus Caloren & Marshall. NE: Northeast North America (Ontario to Nova Scotia and Illinois to North Carolina).

Clusiodes clandestinus Caloren & Marshall 1998: 277 View in CoL . Canada. Ontario: Nipissing District, Algonquin Provincial Park, Swan Lake Research Station. HT ♂ DEBU. Caloren & Marshall 1998: 277 [illustrations of wing and male and female genitalia, on fresh maple log, dogwood leaves and hemlock log]; Marshall, 2006: 510 [photo, female ovipositing in crack in dead wood]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis]; Lonsdale et al. 2010: 240 [genes sequenced for phylogenetic analysis].

coconino Caloren & Marshall. View in CoL NE: USA (Arizona).

Clusiodes coconino Caloren & Marshall 1998: 280 View in CoL . USA. Arizona: Coconino Co., 10 mi NW Flagstaff, San Francisco Mountains. HT ♂ DEBU. Caloren & Marshall 1998: 280 [illustrations of male genitalia]; Lonsdale & Marshall 2007b: 146 [map, discussion of C. melanostomus complex, phylogenetic analysis].

discostylus Sueyoshi. View in CoL PA: Japan.

Hendelia beckeri View in CoL . Misidentification in part. Sueyoshi et al. 2003: 182 [ Japan].

Clusiodes discostylus Sueyoshi 2006: 7 View in CoL . Japan. Hokkaido: Shikaribetsu, Shikaoi Town. HT ♂ BLKU. Sueyoshi 2006: 7 [illustrations of head and male and female genitalia, photo of wing], 2011: 8 [Japan, pictoral key], 2014: 618 [catalogue, Japan]; Lonsdale & Marshall 2007b: 132 [key, map, synonymy, phylogeny, illustrations of genitalia].

eremnos Caloren & Marshall. View in CoL NE: Western North America (Alberta, British Columbia, Oregon, Colorado, Idaho, Montana, Washington).

Clusiodes eremnos Caloren & Marshall 1998: 282 View in CoL . USA. Oregon: Baker Co., 9mi W of Unity, 4500′. HT ♂ CNC. Caloren & Marshall 1998: 282 [illustrations of male genitalia]; Lonsdale & Marshall 2007b: 146 [map, discussion of C. melanostomus complex, phylogenetic analysis].

flaveolus Mamaev. View in CoL PA: Russia.

Clusiodes flaveola Mamaev 1974: 115 View in CoL . Russia. Far East: Primorskiy Krai. Ussury Res. HT ♂ ZMM [Registration No. Di0160]. Mamaev 1974: 115 [illustration of surstylus]; Yagi & Sawada 2000: 41 [Mamaev collection holdings]. Lonsdale & Marshall 2007b: 129 [lapsus]; Ozerov 2010: 145 [types in ZMM].

Clusiodes flaveolus View in CoL . Lonsdale & Marshall 2007b: 137 [key, map, synonymy, notes, phylogeny, Russia].

freyi Tuomikoski. PA: Finland, Russia.

Clusiodes (Clusiaria) Freyi Tuomikoski 1933: 16 . Finland. Padasjoki, Lohja. HT ♂ UZMH [Type No. 4846].

Clusiodes freyi View in CoL . Tuomikoski 1936: 183 [ Finland]; Stackelberg 1988: 505 [key, European USSR]; Yagi & Sawada 2000: 41 [Mamaev collection holdings, questionable identification]; Lonsdale & Marshall 2007b: 139 [key, map, synonymy, notes, phylogeny, illustrations of genitalia, Finland, Russia]; Rotheray & Horsfield 2013: 404 [illustrations of immatures; rearing data— Betula View in CoL , Picea View in CoL ; Finland, Russia]; Kahanpää 2014: 288 [checklist, Finland].

Clusiodes (Clusiaria) freyi View in CoL . Frey 1960: 26 [catalogue, world]; Hackman 1980: 148 [checklist, Finland]; Soós 1984: 12 [catalogue, Palaearctic ( Finland, European Russia)].

gentilis (Collin) . PA: Europe (not Finland), western and central Russia.

Heteroneura gentilis Collin. 1912: 107 View in CoL . United Kingdom. England: Hampshire, New Forest, Lyndhurst Road. LT ♂ BMNH (designated by Lonsdale & Marshall 2007b: 132)]. Collin 1912: 107 [illustration of male external genitalia].

Clusiodes (Clusiodes) gentilis View in CoL . Czerny 1928: 9 [redescription, reillustration from original description]; Melander & Argo 1924: 17 [summary of original description]; Frey 1928c: 109 [catalogue, Finland], 1960: 26 [catalogue, world]; Kloet & Hincks 1945: 394 [checklist, Britain]; Trojan 1962: 26 [key, illustrations of male genitalia and larval cephalopharyngeal skeleton, Poland]; Chandler 1978: 264 [ Ireland]; Stubbs 1982: 92 [Britain]; Soós 1984: 12 [catalogue, Palaearctic (Britain, Poland, Finland, northern European USSR)]; Beuk 2002: 271 [checklist, Netherlands].

Clusiodes gentilis View in CoL . Stackelberg 1933: 381 [key, European USSR], 1988: 505 [key, European USSR]; Richards, 1934: 2 [puparia found in rotten birch]; Séguy 1934: 352 [illustrations of puparium and larval cephalopharyngeal skeleton, France]; Hennig 1952: 227 [immature stages]; Withers 1985: 63 [ England, reared from Salix sp. in bog and rotting Betula View in CoL sp., associated with Fagus sylvaticus ]; Ferrar 1987: 122 [immature stages, illustration of larva]; Smith 1989: 103 [bred from log, illustration of larval mouthparts, Britain]; Ceianu 1989: 29 [expected in Romania]; Woznica 1991: 208 [checklist, Poland]; DeBruyn 1991: 146 [catalogue, Belgium]; Yagi & Sawada 2000: 41 [Mamaev collection holdings]; Carles-Tolrá 2002: 155 [catalogue, Spain]; Roháček 2004: 250 [ Cyprus], 2013: 130 [central Slovakia]; Lonsdale & Marshall 2007b: 132 [synonymy, key, map, phylogeny, lectotype designation, illustrations of genitalia, new records ( England, Netherlands, Russia, Switzerland)]; Rotheray & Horsfield 2013: 404 [illustrations of immatures; rearing data— Betula View in CoL , Fagus View in CoL , Pinus View in CoL , Salix , Ulmus View in CoL ; England, Scotland]; Kahanpää 2014: 288 [reiterates Tuomikoski (1936) that previous Finnish records of gentilis View in CoL were misidentifications of caledonicus View in CoL ]; Roháček et al. 2016: 160 [colour variation, Portugal].

Clusioides (Clusioides) gentilis . Soós 1981: 100 [possible occurrence in Hungary, key, illustrations of wing, male terminalia].

geomyzinus (Fallén) . PA: Europe, Russia.

Heteroneura geomyzina Fallén 1823: 2 . Sweden. “Smolandia”. ST ♂ ♀ UZMC. Meigen 1830: 129 [Europe]; Zetterstedt 1848: 2787 [larvae and puparia subcortical in conifers, Norway, Sweden, Finland], 1852: 4816 [ Sweden]; Loew 1857: 54 [ Germany], 1864b: 345 [redescription, Germany]; Schiner 1862: 38 [redescription], 1864: 72 [catalogue, Europe]; Siebke 1877: 181 [catalogue, Norway]; Brauer 1883: 92 [records of the Vienna Museum]; Becker 1902: 313 [discussion of type], 1905: 39 [catalogue, Palaearctic]; Czerny 1903: 77 [redescription, synonymy of alpina Loew , Sweden, Norway, Denmark, Germany, Austria, Switzerland, Italy, illustrations of head and wing], 1928: 7 [redescription]; Czizek 1910: 112 [catalogue, Germany];

Agromyza geomyzina . Zetterstedt 1838: 790 [generic combination].

Heteroneura alpina Loew 1857: 53 . Germany [not given]. HT ♀, depository unknown. Loew, 1864b: 344 [discussion, Germany]; Girschner 1888: 97 [comparison to albimana Meigen ]; Czerny 1903: 77 [as synonym of geomyzina Fallén ].

Clusiodes geomyzina . Johnson 1913b: 97 [generic combination, Nearctic record likely orbitalis Malloch View in CoL ]; Wahlgren 1917: 186 [diagnosis, key]; Malloch 1918: 6 [synonymy, key]; Stackelberg 1933: 381 [key, European USSR], 1988: 505 [key, European USSR]; Séguy 1934: 352 [France]; Tuomikoski 1936: 183 [adults on decayed deciduous and coniferous logs (particularly birch and aspen), Finland]; Hennig 1952: 227 [immature stages, illustration of puparium]; Máca 1985: 397 [Czech Republic]; Ferrar 1987: 122 [illustration of puparium]; Zuska 1987: 235 [possibly present in Czech Republic]; Dunk 1996: 422 [catalogue, Bavaria]; Yagi & Sawada 2000: 41 [Mamaev collection holdings].

Clusiodes (Clusiaria) geomyzinus View in CoL . Melander & Argo 1924: 18 (in part) [misidentification—possibly orbitalis Malloch View in CoL or nitidus Melander & Argo View in CoL , redescription]; Frey 1928c: 108 [catalogue, Finland]; Tuomikoski 1933: 15 [Finland]; Soós 1984: 12 [catalogue, Palaearctic (Europe, Ukraine, northern European Russia)]; Roháček 1995: 128 [biology, habitat, male agonistic behaviour, Czech and Slovak Republics], 1997: 75 [checklist, Czech and Slovak republics], 1999: 291 [Czech Republic]; Poole 1996: 126 [checklist, Nearctic, [misidentification—possibly orbitalis Malloch View in CoL or nitidus Melander & Argo View in CoL ]; Schumann et al. 1999: 144 [checklist, Germany]; Beuk 2002: 271 [checklist, Netherlands]; Greve 2005: 6 [Norway].

Clusiodes (Clusiaria) geomyzina . Johnson 1925a: 245 [Nearctic records likely misidentifications of orbitalis Malloch View in CoL ]; Czerny, 1928: 7 [redescription, Europe]; Soós 1943: 72 [ Hungary]; Kloet & Hincks 1945: 394 [checklist, Britain]; Frey 1960: 26 [catalogue, world]; Trojan 1962: 22 [key, illustration of wing and puparium, Poland]; Hackman 1980: 148 [checklist, Finland]; Stubbs 1982: 92 [Britain]; Greve & Midtgaard 1986: 90 [habitat, Norway].

Clusiodes (Clusiodes) geomyzinus View in CoL . Steyskal 1965: 807 [catalogue, America north of Mexico - likely Clusiodes orbitalis View in CoL ].

Clusioides (Columbiella) geomyzina . Soós 1981: 102 [possible occurrence in Hungary, key, illustration of wing].

Clusiodes geomyzinus View in CoL . Ceianu 1989: 29 [catalogue, Romania]; Woznica 1991: 208 [checklist, Poland]; Merz 1998: 276 [checklist, Switzerland]; Roháček & Barták 2001: 338 [checklist, Czech Republic]; Carles-Tolrá & Ventura 2001: 12 [ Iberia]; Carles-Tolrá 2002: 155 [catalogue, Spain]; Roháček et al. 2005: 280 [catalogue, habitat, Czech Republic]; Barták & Kubík 2005: 280 [ Czech Republic]; Lonsdale & Marshall 2007b: 140 [key, biology, map, synonymy, notes, phylogeny, illustrations of genitalia, new records ( Finland, Russia, Spain, Sweden, Switzerland)]; Roháček 2008: 154 [ Italy], 2009: 218 [ Slovakia], 2012: 99 [central Slovakia]; Nilsson 2009: 5 [ Sweden]; Rotheray & Horsfield 2013: 404 [illustration of immature; photo of puparium; rearing data— Betula View in CoL (?), Picea View in CoL , Pinus View in CoL ; Finland, Russia]; Rotheray & Wilkinson 2013: 1356 [ Scotland]; Kahanpää 2014: 288 [checklist, Finland].

gracilolobus Caloren & Marshall. View in CoL NE: Alabama, Florida, Georgia, Kentucky, Maryland, Virginia.

Clusiodes gracilolobus Caloren & Marshall 1998: 282 View in CoL . USA. Maryland: Montgomery Co., Rockville. HT ♂ USNM. Caloren & Marshall 1998: 282 [photo and illustrations of male genitalia]; Lonsdale & Marshall 2007b: 146 [notes, map, discussion of C. melanostomus complex, phylogenetic analysis, photo, Alabama, Kentucky].

johnsoni Malloch. View in CoL NE: Widespread in North America south of Canadian Territories.

Heteroneura albimana . Misidentification. Osten Sacken 1878: 197 [catalogue, North America].

Clusiodes (Clusiodes) albimana . Misidentification. Johnson 1913b: 98 [eastern United States], 1925a: 245 [catalogue, Vermont]; Frey 1960: 26 (in part) [catalogue, world].

Clusiodes albimana . Misidentification. Malloch 1918: 6 [Ontario]; Johannsen 1928: 844 [catalogue, New York].

Clusiodes (Columbiella) johnsoni Malloch 1922: 49 View in CoL . USA. Maine: Kennebec Co., Mt. Desert, Echo Lake. HT ♂ MCZ [Type No. MCZT 27015]. Melander & Argo 1924: 23 [redescription, new records (Ontario, Maryland, New York, Virginia)]; Johnson 1925a: 246 [catalogue, New England (Maine, Massachutsetts, Vermont)]; Steyskal 1965: 807 [catalogue, America north of Mexico].

Clusiodes (Columbiella) nigripalpis Malloch 1922: 49 View in CoL . USA. Maryland: Glen Echo. HT ♂ USNM . Johnson 1925a: 246 [catalogue, New England (Maine, Massachusetts, New Hampshire, Vermont), misidentification in part of apiculatus Malloch View in CoL ].

Clusiodes albimanus var. pallidior . Misidentification in part. Melander & Argo 1924: 16 [photo, new records (Idaho, Washington)].

Clusiodes (Columbiella) johnsoni View in CoL form nigripalpis View in CoL . Melander & Argo 1924: 23 [new status]. Caloren & Marshall 1998: 274 [as synonym of johnsoni Malloch View in CoL ].

Clusiodes johnsoni View in CoL . Johannsen 1928: 844 [catalogue, New York]; Poole 1996: 126 [checklist, Nearctic]; Caloren & Marshall 1998: 274 [redescription, illustrations of wing and male and female genitalia, new records (North America)]; Lonsdale & Marshall 2007b: 146 [phylogenetic analysis, illustrations of puparium and head, photo], 2010: 1045 [illustration of male genitalia]; Lonsdale et al. 2010: 240 [genes sequenced for phylogenetic analysis].

Clusiodes (Clusiodes) johnsoni View in CoL . Frey 1960: 26 [catalogue, world].

Clusiodes nigripalpis View in CoL . Poole 1996: 126 [as synonym of johnsoni , checklist, Nearctic].

latus Sasakawa. OR: Laos.

Hendelia latus Sasakawa 2011b: 21 . Laos. Vientian: Ban Van Eue, 750m. HT ♂, BPBM [Type No. BPBM 17354]. Sasakawa 2011b: 21 [illustrations of male genitalia].

leptapodemus Caloren & Marshall. View in CoL NE: New Mexico, Arizona, Nevada.

Clusiodes leptapodemus Caloren & Marshall 1998: 280 View in CoL . USA. New Mexico: Socorro Co., South Baldy Peak, 20mi W of Socorro, alpine meadow, 10,400′. HT ♂ DEBU. Caloren & Marshall 1998: 280 [illustrations of male and female genitalia]; Lonsdale & Marshall 2007b: 146 [map, discussion of C. melanostomus complex, phylogenetic analysis].

leucopeza Frey. View in CoL OR: Burma.

Clusiodes (Clusiodes) leucopeza Frey 1960: 18 View in CoL . Burma. Kambaiti, 2000 m. ST 1♂ 2♀ UZMH. Frey 1960: 18 [catalogue, world]; Sasakawa 1977: 236 [catalogue, Orient], 1987: 101 [key].

Clusiodes leucopeza View in CoL . Lonsdale & Marshall 2007b: 132 [key, map, synonymy, phylogeny, notes].

melanostomus (Loew) . NE: Northern North America (uncommon in southern United States).

Heteroneura melanostoma Loew 1864a: 260 . USA. New Hampshire. LT 1♂ MCZ [accidental designation of lectotype by assumption of holotype (IZCN (1999) Art. 74.6) by Caloren & Marshall (1998); 1 ST originally described]. Osten Sacken 1878: 198 [catalogue, North America]; Czerny 1903: 77 [quotes description]; Aldrich 1905: 570 [catalogue, North America].

Heteroneura flavifascies Slosson 1897: 239 View in CoL . USA. New Hampshire: Coos Co., Mt. Washington. Slosson 1897: 239 [nomen nudum, attributed to Coquillett]; Poole 1996: 126 [checklist, Nearctic, as syn. of melanostoma Loew attributed to Coquillett].

Clusiodes melanostoma . Johnson 1913b: 98 [generic combination, new records (Quebec, Maine, Massachusetts, New Hampshire, Vermont)], 1925b: 95 [Harris collection]; Malloch 1918: 5 [Illinois]; Griffiths 1972: 214 [male dissected for discussion of phylogenetic relationships]; Poole 1996: 126 [checklist, Nearctic].

Clusiodes (Clusiaria) duplicata Malloch 1922: 49 View in CoL . USA. Maryland: Glen Echo. HT ♂ USNM. Melander & Argo 1924: 19 [as synonym of melanostoma Loew ].

Clusiodes (Clusiaria) melanostoma . Melander & Argo 1924: 19 [redescription, synonymy of duplicata Malloch View in CoL , photo, new records (Illinois, Maryland, New Hampshire, New York, Virginia, Washington)]; Johnson 1925a: 245 [catalogue, New England (Maine, Massachusetts, New Hampshire, Vermont)]; Frey 1960: 26 [catalogue, world].

Clusiodes (Clusiaria) ruficollis View in CoL . Misidentification. Melander & Argo 1924: 22 [redescription, possibly melanostomus Loew ].

Clusiodes melanostomus View in CoL . Johannsen 1928: 844 [catalogue, New York]; Caloren & Marshall 1998: 279 [redescription, illustrations of male and female genitalia, new records (northern North America), designation of LT]; Lonsdale & Marshall 2007b: 146 [notes, map, illustration of puparium, discussion of C. melanostomus View in CoL complex, phylogenetic analysis]; Lonsdale et al. 2010: 240 [member of “ Clusiodes melanostomus View in CoL complex” genetically sequenced for phylogenetic analysis].

Clusiodes (Clusiodes) melanostoma . Steyskal 1965: 807 [catalogue, America north of Mexico].

Clusiodes duplicata View in CoL . Poole 1996: 126 [as synonym of melanostomus , checklist, Nearctic].

microcerus Stackelberg. PA: Finland, Russia.

Clusiodes (Clusiodes) microcera Stackelberg, 1955: 329 View in CoL . Russia. Leningrad Region: Luzhnitsky Rayon, Tolmachevo. HT ♂ ZIN. Frey 1960: 26 [catalogue, world]; Stackelberg 1988: 505 [key, European USSR].

Clusiodes microcera View in CoL . Mamaev 1987: 114 [Russian Far East].

Clusiodes microcerca . Misspelling. Yagi & Sawada 2000: 40 [Mamaev collection holdings].

Clusiodes microcerus View in CoL . Lonsdale & Marshall 2007b: 134 [key, map, synonymy, notes, phylogeny, illustrations of genitalia, Russia]; Rotheray & Horsfield 2013: 405 [illustrations of immatures; rearing data— Betula View in CoL ; Finland]; Kahanpää 2014: 288 [checklist, Finland].

niger Melander & Argo. View in CoL NE: Georgia, New Mexico.

Clusiodes (Clusiodes) nigra View in CoL . Melander & Argo 1924: 17. USA. New Mexico: Lincoln Co., Rio Ruidoso, White Mts., ~6500′. HT ♂ USNM [Cat. No. 26262]. Frey, 1960: 26 [catalogue, world].

Clusiodes (Columbiella) niger View in CoL . Steyskal 1965: 807 [catalogue, America north of Mexico].

Clusiodes niger View in CoL . Poole 1996: 126 [checklist, Nearctic]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis].

Clusiodes nigrus . Caloren & Marshall 1998: 273 [redescription, illustrations of wing and male genitalia].

nitidus Melander & Argo. NE: New Mexico, Arizona, Mexico.

Clusiodes (Clusiaria) nitida Melander & Argo 1924: 20 View in CoL [with photo]. USA. New Mexico: Rio Arriba Co., Hill above Beulah. HT ♀ USNM [Cat. No. 26263]. Frey 1960: 26 [catalogue, world].

Clusiodes (Clusiodes) nitidus View in CoL . Steyskal 1965: 807 [catalogue, America north of Mexico].

Clusiodes nitidus View in CoL . Poole 1996: 126 [checklist, Nearctic]; Caloren & Marshall 1998: 284 [redescription, illustrations of wing and male and female genitalia, new records (Arizona, Mexico)]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis].

orbitalis Malloch. View in CoL NE: North America (uncommon in southern USA).

Clusiodes (Columbiella) orbitalis Malloch 1922: 50 View in CoL . USA. Maine: Hancock Co., Bar Harbor. HT ♀ MCZ [Type No. MCZT 27016 ].

Clusiodes (Clusiaria) orbitalis View in CoL . Melander & Argo 1924: 21 [summary of original description]; Johnson 1925a: 245 [catalogue, Maine]; Frey 1960: 26 [catalogue, world].

Clusiodes (Clusiodes) orbitalis View in CoL . Steyskal 1965: 807 [catalogue, America north of Mexico].

Clusiodes orbitalis View in CoL . Poole 1996: 126 [checklist, Nearctic]; Caloren & Marshall 1998: 287 [redescription, illustrations of wing and male and female genitalia, new records (North America)]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis]; Lonsdale et al. 2010: 240 [genes sequenced for phylogenetic analysis].

pictipes (Zetterstedt) View in CoL . PA: Europe, Russia.

Heteroneura pictipes Zetterstedt 1852: 4816 View in CoL . Sweden. “Scania”. ST ♂♀ MZLU. Schiner 1864: 72 [catalogue, Europe]; Slosson 1897: 239 [checklist, New Hampshire - likely orbitalis Malloch View in CoL ]; Czerny 1903: 80 [redescription]; Aldrich 1905: 571 [catalogue, North America - likely orbitalis Malloch View in CoL ]; Becker 1905: 39 [catalogue, Palaearctic]; Czizek 1910: 112 [catalogue, Germany].

Clusiodes pictipes View in CoL . Johnson 1913b: 98 [generic combination, Nearctic record - likely orbitalis Malloch View in CoL ]; Wahlgren 1917: 187 [diagnosis, key]; Malloch 1918: 7 [doubtful identification—likely orbitalis Malloch]; Tuomikoski 1933 View in CoL : 18 [ Finland], 1936: 183 [ Finland, adults on rotting birch and aspen trunks]; Stackelberg 1988: 505 [key, European USSR]; DeBruyn 1991: 146 [catalogue, Belgium]; Poole 1996: 126 [checklist, Nearctic, likely misidentification from Malloch (1918)]; Roháček & Barták 2001: 338 [checklist, Czech Republic]; Roháček et al. 2005: 280 [catalogue, habitat, Czech Republic]; Barták & Kubík 2005: 280 [ Czech Republic]; Lonsdale & Marshall 2007b: 140 [key, map, synonymy, phylogeny, illustrations of genitalia, illustration of egg, new records ( Hungary, Slovakia, Russia, Ukraine)]; Roháček 2008: 154 [Italy—acceptance of record in Palm (1869)], 2012: 99 [central Slovakia], 2013: 128 [central Slovakia]; Kahanpää 2014: 288 [checklist, Finland].

Clusiodes (Clusiaria) pictipes View in CoL . Czerny 1928: 7 [redescription]; Melander & Argo, 1924: 21 [redescription, USA specimens of uncertain identity]; Johnson 1925a: 245 [Nearctic record—likely misidentification of orbitalis Malloch View in CoL ]; Czerny 1928: 7 [redescription]; Frey 1928c: 109 [catalogue, Finland], 1960: 26 [catalogue, world]; Hackman 1980: 148 [checklist, Finland]; Soós 1984: 12 [catalogue, Palaearctic (northern Europe and northern European Russia)]; Roháček 1995: 130 [biology, Czech and Slovak Republics], 1997: 75 [checklist, Czech and Slovak republics], 1999: 291 [ Czech Republic], 2009: 218 [ Slovakia]; Schumann et al. 1999: 144 [checklist, Germany]; Greve 2005: 7 [considered rare, Norway].

Clusiodes (Clusiodes) pictipes View in CoL . Steyskal 1965: 807 [catalogue, America north of Mexico].

ruficollis (Meigen) . PA: Europe, Russia.

Heteroneura ruficollis Meigen 1830: 128 View in CoL . Europe. ST? NMW [Winthem collection]. Zetterstedt 1848: 2789 [redescription, Sweden]; Loew 1857: 53 [ Germany], 1864b: 343 [synonymy of nubila Meigen and laterella Zetterstedt , Germany]; Schiner 1862: 38 [redescription, key, Austria], 1864: 72 [catalogue, Europe]; Becker 1902: 313 [examination of Meigen types], 1905: 39 [catalogue, Palaearctic]; Czerny, 1903: 75 [redescription, Germany, Sweden].

Heteroneura laterella Zetterstedt 1848: 2791 View in CoL . Sweden. “Smolandia”. ST ♂, depository unknown. Loew 1857: 52 [Germany]; Loew 1864b: 343 [as synonym of ruficollis Meigen ].

Clusiodes ruficollis View in CoL . Wahlgren 1917: 186 [diagnosis, key]; Stackelberg 1933: 381 [key, European USSR], 1988: 505 [key, European USSR]; Séguy 1934: 353 [ France, illustrations of puparium and male genitalia (genitalia in illustration those of Clusia flava View in CoL )]; Tuomikoski 1936: 183 [puparia in rotting alder and birch, Finland]; Hennig 1938: 125 [illustration of male genitalia], 1952: 227 [immature stages]; Vaňhara 1981: 30 [ Czech Republic]; Krištofík & Zuska 1986: 165 [ Slovakia]; Zuska 1987: 235 [ Czech Republic]; Ceianu 1989: 29 [expected in Romania]; Woznica 1991: 208 [checklist, Poland]; DeBruyn 1991: 146 [catalogue, Belgium]; Poole 1996: 126 [checklist, Nearctic, likely representing misidentified melanostoma record in Melander & Argo (1924)]; Merz 1998: 276 [checklist, Switzerland]; Yagi & Sawada 2000: 41 [Mamaev collection holdings]; Roháček & Barták 2001: 338 [checklist, Czech Republic]; Roháček et al. 2005: 280 [catalogue, habitat, Czech Republic]; Barták & Kubík 2005: 280 [ Czech Republic]; Lonsdale & Marshall 2007b: 141 [key, map, synonymy, notes, phylogeny, illustrations of genitalia, new records ( Estonia, Finland, Netherlands, Russia, Switzerland, Ukraine)]; Nilsson 2009: 5 [ Sweden]; Roháček 2009: 218 [ Slovakia], 2012: 99 [central Slovakia], 2013: 128 [central Slovakia]; Rotheray & Horsfield 2013: 405 [illustrations of immatures; rearing data— Betula View in CoL , Fagus View in CoL , Fraxinus View in CoL ; England, Scotland]; Kahanpää 2014: 288 [checklist, Finland].

Clusiodes (Clusiaria) ruficollis View in CoL . Czerny 1928: 7 [redescription]; Malloch 1922: 47 [key to subgenera and species, eastern USA—likely melanostomus Loew View in CoL ]; Melander & Argo 1924: 22 [redescription - likey melanostomus Loew View in CoL )]; Frey 1928c: 108 [catalogue, Finland], 1960: 26 [catalogue, world]; Tuomikoski 1933: 15 [key]; Soós 1943: 72 [ Hungary], 1984: 12 [catalogue, Palaearctic (central and northern Europe, northern European Russia)]; Trojan 1962: 23 [key, illustrations of head and leg, Poland]; Hackman 1980: 148 [checklist, Finland]; Withers 1985: 63 [reared from larvae and puparia found beneath bark of felled hornbeam ( Carpinus betulus View in CoL ), synonymy of facialis Collin View in CoL , illustrations of face, England]; Vaňhara 1986: 123 [ Czech Republic]; Greve & Midtgaard 1986: 90 [bred from Quercus View in CoL sp. and Populus tremula View in CoL , Norway]; Roháček 1995: 131 [biology, habitat, adults on stumps and fallen trunks of various deciduous trees, reared from puparia in decayed birch logs, infestation with mites, mating behaviour, Czech and Slovak Republics], 1997: 75 [checklist, Czech and Slovak republics], 1999: 291 [ Czech Republic]; Schumann et al. 1999: 144 [checklist, Germany]; Beuk 2002: 271 [checklist, Netherlands]; Schacht 2004: 276 [ Germany]; Greve 2005: 8 [ Norway].

Clusiodes (Clusiaria) facialis View in CoL . Collin 1953a: 139. United Kingdom. England: Cambridgeshire, Chippenham Fen; Sutherland: Golspie; Cheshire: Wilmslow, Rostherne, Cotterhill Clough. Various locations. ST ♂♀, depositories unknown. Collin 1953a: 139 [on decayed fallen trunk]. Frey 1960: 26 [catalogue, world]; Stubbs 1982: 92 [Britain]; Soós 1984: 12 [catalogue, Palaearctic]. Withers 1985: 63 [as synonym of ruficollis Meigen View in CoL ].

Clusiodes (Clusiodes) ruficollis View in CoL . Steyskal 1965: 807 [catalogue, America north of Mexico].

Clusioides (Columbiella) ruficollis . Soós 1981: 101 [ Hungary, key].

spinos Caloren & Marshall. NE: Tennessee, New Hampshire, Vermont.

Clusiodes spinos Caloren & Marshall, 1998: 284 View in CoL . USA. Tennessee: Sevier Co., Great Smoky Mountains National Park, Arch Rock. HT ♂ USNM. Caloren & Marshall 1998: 284 [illustrations of male genitalia]; Lonsdale & Marshall 2007b: 146 [map, discussion of C. melanostomus View in CoL complex, phylogenetic analysis].

terminalis Melander & Argo. View in CoL NE: Ontario, New Hampshire.

Clusiodes (Clusiaria) terminalis Melander & Argo 1924: 22 View in CoL . USA. New Hampshire: White Mountains, Morrison. HT ♂ USNM [Cat. No. 26265]. Melander & Argo 1924: 22 [with photo]; Johnson 1925a: 246 [catalogue, New England]; Frey 1960: 26 [catalogue, world].

Clusiodes (Clusiodes) terminalis View in CoL . Steyskal 1965: 807 [catalogue, America north of Mexico].

Clusiodes terminalis View in CoL . Poole 1996: 126 [checklist, Nearctic]; Caloren & Marshall 1998: 293 [redescription, illustrations of wing and male genitalia, new record (Ontario)]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis].

tobi Sueyoshi. View in CoL PA: Japan.

Clusiodes tobi Sueyoshi 2006: 10 View in CoL . Japan. Hokkaido: Shikaribetsu, Shikaoi Town. HT ♂ BLKU. Sueyoshi 2006: 10 [illustrations of head and male and female genitalia, photo of wing], 2011: 8 [Japan, pictoral key], 2012: 26 [Japan], 2014: 618 [catalogue, Japan]; Lonsdale & Marshall 2007b: 142 [key, map, synonymy, notes, phylogeny, illustrations of genitalia].

tuomikoskii Mamaev. View in CoL PA: Russia (Far East).

Clusiodes tuomikoskii Mamaev 1974: 114 View in CoL . Russia. Far East: Primorsky Krai, Khasansky District, Kedrovaja Pad Nature Reserve. HT ♂ [with puparium] MNHA [not in ZIN, as stated in original publication, examined by author]. Mamaev 1974: 114 [illustration of surstylus]; Yagi & Sawada 2000: 41 [Mamaev collection holdings]. Lonsdale & Marshall 2007b: 142 [key, map, synonymy, phylogeny, illustrations of genitalia].

verticalis (Collin) View in CoL . PA: Europe, Israel, western Russia.

Heteroneura verticalis Collin 1912: 107 View in CoL . United Kingdom. England: Various localities. ST 3♂ [♀?] BMNH [additional depository(ies) unknown]. Collin 1912: 107 [illustration of male external genitalia].

Clusiodes (Columbiella) verticalis View in CoL . Melander & Argo 1924: 24 [synonymy, generic combination, comparison to albimanus Meigen View in CoL ]; Czerny 1928: 9 [redescription, reillustration]; Frey 1928c: 109 [ Finland, catalogue], 1960: 26 [catalogue, world]; Kloet & Hincks 1945: 394 [checklist, Britain]; Trojan 1962: 26 [key, illustration of male genitalia, Poland]; Chandler 1978: 264 [ Ireland, reared from rotten wood]; Stubbs 1982: 92 [Britain]; Greve 1983: 55 [habitat, Norway], 2005: 8 [ Norway]; Soós 1984: 12 [catalogue, Palaearctic]; Withers 1985: 63 [reared from rotten birch, illustration of posterior end of puparium, Britain]; Greve & Midtgaard 1986: 91 [ Norway]; Stackelberg 1988: 505 [key, European USSR]; Smith 1989: 103 [reared from old grass heap, England]; Roháček 1995: 133 [biology, Czech and Slovak Republics], 1997: 75 [checklist, Czech and Slovak republics]; Beuk 2002: 271 [checklist, Netherlands]; Schacht 2004: 276 [ Germany].

Clusiodes verticalis View in CoL . Andersson 1971: 234 [ Sweden]; Máca 1985: 398 [adult at fermenting beer in sawdust dump, Czech Republic]; Ceianu 1989: 29 [expected in Romania]; DeBruyn 1991: 146 [catalogue, Belgium]; Barrientos et al. 1996: 120 [biology, Spain]; Merz 1998: 276 [checklist, Switzerland]; Carles-Tolrá 2002: 155 [catalogue, Spain]; Lonsdale & Marshall 2007b: 134 [key, map, synonymy, notes, phylogeny, illustrations of genitalia, new records ( Finland, France, Israel, Italy, Switzerland)]; Roháček 2008: 154 [ Italy]; 2009: 218 [ Slovakia], 2013: 128 [central Slovakia]; Lonsdale et al. 2010: 240 [genes sequenced for phylogenetic analysis]; Rotheray & Horsfield 2013: 405 [illustrations of immatures; rearing data— Alnus View in CoL , Betula View in CoL , Fagus View in CoL , Fraxinus View in CoL , Pinus View in CoL , Salix ; England, Scotland]; Kahanpää 2014: 288 [checklist, Finland]; Carles-Tolrá & Andrade 2011: 489 [ Portugal]; Roháček et al. 2016: 160 [ Portugal].

Clusioides (Clusioides) verticalis . Soós 1981: 98 [ Hungary, key, associated with fungus Lactarius piperatus , illustrations of male and female terminalia, wing].

USNM

Smithsonian Institution, National Museum of Natural History

MZLU

Lund University

CUMZ

Chulalongkorn University Museum of Natural History

UZMH

University Museum (Zoology)

MCZ

Museum of Comparative Zoology

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Diptera

Family

Clusiidae

Loc

Clusiodes

Owen Lonsdale 2017
2017
Loc

Hendelia latus

Sasakawa 2011: 21
Sasakawa 2011: 21
2011
Loc

Clusiodes flaveolus

Lonsdale 2007: 137
2007
Loc

Clusiodes leucopeza

Lonsdale 2007: 132
2007
Loc

Clusiodes microcerus

Kahanpaa 2014: 288
Rotheray 2013: 405
Lonsdale 2007: 134
2007
Loc

Clusiodes discostylus

Lonsdale 2007: 132
Sueyoshi 2006: 7
Sueyoshi 2006: 7
2006
Loc

Clusiodes tobi

Lonsdale 2007: 142
Sueyoshi 2006: 10
Sueyoshi 2006: 10
2006
Loc

Hendelia beckeri

Sueyoshi 2003: 182
2003
Loc

Clusiodes caestus

Lonsdale 2007: 146
Caloren 1998: 289
Caloren 1998: 289
1998
Loc

Clusiodes clandestinus

Lonsdale 2007: 146
Caloren 1998: 277
Caloren 1998: 277
1998
Loc

Clusiodes coconino

Lonsdale 2007: 146
Caloren 1998: 280
Caloren 1998: 280
1998
Loc

Clusiodes eremnos

Lonsdale 2007: 146
Caloren 1998: 282
Caloren 1998: 282
1998
Loc

Clusiodes gracilolobus

Lonsdale 2007: 146
Caloren 1998: 282
Caloren 1998: 282
1998
Loc

Clusiodes leptapodemus

Lonsdale 2007: 146
Caloren 1998: 280
Caloren 1998: 280
1998
Loc

Clusiodes nigrus

Caloren 1998: 273
1998
Loc

Clusiodes spinos

Lonsdale 2007: 146
Caloren 1998: 284
Caloren 1998: 284
1998
Loc

Clusiodes (Clusiodes) albimanus

Schacht 2004: 276
Rohacek 1997: 75
1997
Loc

Clusiodes (Clusiodes) caledonicus

Rohacek 1997: 75
1997
Loc

Clusiodes americanus

Lonsdale 2007: 146
Caloren 1998: 271
Poole 1996: 126
1996
Loc

Clusiodes apiculatus

Lonsdale 2007: 146
Caloren 1998: 269
Poole 1996: 126
1996
Loc

Clusiodes scutellatus

Poole 1996: 126
1996
Loc

Clusiodes nigripalpis

Poole 1996: 126
1996
Loc

Clusiodes duplicata

Poole 1996: 126
1996
Loc

Clusiodes niger

Lonsdale 2007: 146
Poole 1996: 126
1996
Loc

Clusiodes nitidus

Lonsdale 2007: 146
Caloren 1998: 284
Poole 1996: 126
1996
Loc

Clusiodes orbitalis

Lonsdale 2007: 146
Caloren 1998: 287
Poole 1996: 126
1996
Loc

Clusiodes terminalis

Lonsdale 2007: 146
Caloren 1998: 293
Poole 1996: 126
1996
Loc

Clusiodes calcedonicus

DeBruyn 1991: 146
1991
Loc

Clusiodes caledonicus

Chandler 2015: 212
Kahanpaa 2014: 288
Rotheray 2013: 404
Nilsson 2009: 5
Lonsdale 2007: 131
Rohacek 2001: 338
Merz 1998: 276
Ceianu 1989: 29
1989
Loc

Clusiodes geomyzinus

Kahanpaa 2014: 288
Rotheray 2013: 404
Rotheray 2013: 1356
Nilsson 2009: 5
Rohacek 2008: 154
Lonsdale 2007: 140
Carles-Tolra 2002: 155
Rohacek 2001: 338
Carles-Tolra 2001: 12
Merz 1998: 276
Woznica 1991: 208
Ceianu 1989: 29
1989
Loc

Clusiodes microcera

Mamaev 1987: 114
1987
Loc

Clusiodes (Clusiodes) albima

Greve 1986: 89
1986
Loc

Clusiodes albimanus

Kahanpaa 2014: 288
Rotheray 2013: 403
Nilsson 2009: 5
Rohacek 2008: 154
Lonsdale 2007: 129
Carles-Tolra 2002: 155
Rohacek 2001: 338
Merz 1998: 276
Poole 1996: 126
Barrientos 1996: 118
Woznica 1991: 208
DeBruyn 1991: 146
Ceianu 1989: 29
Soos 1987: 855
Kristofik 1986: 165
1986
Loc

Clusiodes (Clusiodes) albimanus

Greve 2005: 3
Beuk 2002: 271
Schumann 1999: 144
Rohacek 1995: 126
Soos 1984: 12
1984
Loc

Clusiodes (Clusiaria) pictipes

Maca 1982: 71
1982
Loc

Clusioides (Clusioides) albimana

Soos 1981: 99
1981
Loc

Clusioides (Columbiella) nubila

Soos 1981: 102
1981
Loc

Clusioides (Columbiella) apicalis

Soos 1981: 102
1981
Loc

Clusioides (Clusioides) caledonica

Soos 1981: 100
1981
Loc

Clusioides (Clusioides) gentilis

Soos 1981: 100
1981
Loc

Clusioides (Columbiella) geomyzina

Soos 1981: 102
1981
Loc

Clusioides (Columbiella) ruficollis

Soos 1981: 101
1981
Loc

Clusioides (Clusioides) verticalis

Soos 1981: 98
1981
Loc

Clusiodes flaveola

Ozerov 2010: 145
Lonsdale 2007: 129
Mamaev 1974: 115
Mamaev 1974: 115
1974
Loc

Clusiodes tuomikoskii

Lonsdale 2007: 142
Mamaev 1974: 114
Mamaev 1974: 114
1974
Loc

Clusiodes verticalis

Kahanpaa 2014: 288
Rotheray 2013: 405
Carles-Tolra 2011: 489
Rohacek 2008: 154
Lonsdale 2007: 134
Carles-Tolra 2002: 155
Merz 1998: 276
Barrientos 1996: 120
DeBruyn 1991: 146
Ceianu 1989: 29
Maca 1985: 398
Andersson 1971: 234
1971
Loc

Clusiodes (Columbiella) albimanus

Steyskal 1965: 807
1965
Loc

Clusiodes (Columbiella) americanus

Steyskal 1965: 807
1965
Loc

Clusiodes (Clusiodes) apicalis

Steyskal 1965: 807
1965
Loc

Clusiodes (Columbiella) apiculatus

Steyskal 1965: 807
1965
Loc

Clusiodes (Clusiodes) ater

Steyskal 1965: 807
1965
Loc

Clusiodes (Clusiodes) nitidus var. scutellatus

Steyskal 1965: 807
1965
Loc

Clusiodes (Clusiodes) geomyzinus

Steyskal 1965: 807
1965
Loc

Clusiodes (Clusiodes) melanostoma

Steyskal 1965: 807
1965
Loc

Clusiodes (Columbiella) niger

Steyskal 1965: 807
1965
Loc

Clusiodes (Clusiodes) nitidus

Steyskal 1965: 807
1965
Loc

Clusiodes (Clusiodes) orbitalis

Steyskal 1965: 807
1965
Loc

Clusiodes (Clusiodes) pictipes

Steyskal 1965: 807
1965
Loc

Clusiodes (Clusiodes) ruficollis

Steyskal 1965: 807
1965
Loc

Clusiodes (Clusiodes) terminalis

Steyskal 1965: 807
1965
Loc

Clusiodes (Clusiodes) apiculata

Frey 1960: 26
1960
Loc

Clusiodes (Clusiaria) freyi

Soos 1984: 12
Hackman 1980: 148
Frey 1960: 26
1960
Loc

Clusiodes (Clusiodes) johnsoni

Frey 1960: 26
1960
Loc

Clusiodes (Clusiodes) leucopeza

Sasakawa 1977: 236
Frey 1960: 18
Frey 1960: 18
1960
Loc

Clusiodes (Clusiodes) microcera

Stackelberg 1988: 505
Frey 1960: 26
Stackelberg 1955: 329
1955
Loc

Clusiodes (Clusiaria) facialis

Withers 1985: 63
Soos 1984: 12
Stubbs 1982: 92
Frey 1960: 26
Collin 1953: 139
Collin 1953: 139
1953
Loc

Clusiodes apiculata

Brimley 1938: 378
1938
Loc

Clusiodes caledonica

Stackelberg 1988: 505
Speight 1979: 402
Tuomikoski 1936: 184
1936
Loc

Clusiodes freyi

Kahanpaa 2014: 288
Rotheray 2013: 404
Lonsdale 2007: 139
Stackelberg 1988: 505
Tuomikoski 1936: 183
1936
Loc

Clusioides

Soos 1981: 97
Craighead 1950: 541
Curran 1934: 363
1934
Loc

Clusiodes nubila

Ceianu 1989: 29
Seguy 1934: 352
1934
Loc

Clusiodes (Clusiaria) Freyi Tuomikoski 1933 : 16

Tuomikoski 1933: 16
1933
Loc

Clusiodes gentilis

Kahanpaa 2014: 288
Rotheray 2013: 404
Lonsdale 2007: 132
Rohacek 2004: 250
Carles-Tolra 2002: 155
Woznica 1991: 208
DeBruyn 1991: 146
Smith 1989: 103
Ceianu 1989: 29
Ferrar 1987: 122
Withers 1985: 63
Hennig 1952: 227
Richards 1934: 2
Seguy 1934: 352
Stackelberg 1933: 381
1933
Loc

Clusiodes atra

Johannsen 1928: 844
1928
Loc

Clusiodes (Clusiodes) gentilis

Beuk 2002: 271
Soos 1984: 12
Stubbs 1982: 92
Trojan 1962: 26
Kloet 1945: 394
Czerny 1928: 9
Frey 1928: 109
Melander 1924: 17
1928
Loc

Clusiodes johnsoni

Lonsdale 2007: 146
Caloren 1998: 274
Poole 1996: 126
Johannsen 1928: 844
1928
Loc

Clusiodes melanostomus

Lonsdale 2007: 146
Caloren 1998: 279
Johannsen 1928: 844
1928
Loc

Clusiodes (Clusiaria) pictipes

Greve 2005: 7
Schumann 1999: 144
Rohacek 1995: 130
Soos 1984: 12
Hackman 1980: 148
Czerny 1928: 7
Czerny 1928: 7
Frey 1928: 109
Johnson 1925: 245
Melander 1924: 21
1928
Loc

Clusiodes (Clusiaria) ruficollis

Greve 2005: 8
Schacht 2004: 276
Beuk 2002: 271
Schumann 1999: 144
Rohacek 1995: 131
Vanhara 1986: 123
Greve 1986: 90
Withers 1985: 63
Hackman 1980: 148
Trojan 1962: 23
Soos 1943: 72
Tuomikoski 1933: 15
Czerny 1928: 7
Frey 1928: 108
Melander 1924: 22
Malloch 1922: 47
1928
Loc

Clusiodes (Clusiaria) geomyzina

Greve 1986: 90
Stubbs 1982: 92
Hackman 1980: 148
Trojan 1962: 22
Frey 1960: 26
Kloet 1945: 394
Soos 1943: 72
Czerny 1928: 7
Johnson 1925: 245
1925
Loc

Clusiodes (Clusiodes) albimana

Melander 1924: 15
1924
Loc

Clusiodes (Clusiodes) albimana

Melander 1924: 15
1924
Loc

Clusiodes (Clusiaria) nubila

Schumann 1999: 144
Soos 1984: 12
Trojan 1962: 22
Frey 1960: 26
Czerny 1928: 7
Melander 1924: 21
1924
Loc

Clusiodes (Clusiodes) americana

Frey 1960: 26
Johnson 1925: 245
Melander 1924: 16
1924
Loc

Clusiodes (Clusiaria) apicalis

Greve 2005: 5
Schumann 1999: 144
Rohacek 1995: 128
Greve 1986: 89
Stubbs 1982: 92
Hackman 1980: 148
Trojan 1962: 24
Collin 1953: 139
Soos 1943: 72
Czerny 1928: 6
Frey 1928: 109
Melander 1924: 17
1924
Loc

Clusiodes (Clusiaria) atra

Frey 1960: 26
Johnson 1925: 245
Melander 1924: 18
1924
Loc

Clusiodes (Clusiaria) nitida var. scutellata

Caloren 1998: 291
Melander 1924: 20
1924
Loc

Clusiodes (Clusiodes) caledonica

Greve 2005: 4
Beuk 2002: 271
Schumann 1999: 144
Rohacek 1995: 128
Greve 1986: 90
Greve 1986: 90
Soos 1984: 12
Stubbs 1982: 92
Hackman 1980: 148
Trojan 1962: 26
Frey 1960: 26
Kloet 1945: 394
Czerny 1928: 9
Melander 1924: 17
1924
Loc

Clusiodes (Clusiaria) geomyzinus

Greve 2005: 6
Beuk 2002: 271
Schumann 1999: 144
Poole 1996: 126
Rohacek 1995: 128
Soos 1984: 12
Tuomikoski 1933: 15
Frey 1928: 108
Melander 1924: 18
1924
Loc

Clusiodes albimanus var. pallidior

Melander 1924: 16
1924
Loc

Clusiodes (Columbiella) johnsoni

Caloren 1998: 274
Melander 1924: 23
1924
Loc

Clusiodes (Clusiaria) melanostoma

Frey 1960: 26
Johnson 1925: 245
Melander 1924: 19
1924
Loc

Clusiodes (Clusiaria) ruficollis

Melander 1924: 22
1924
Loc

Clusiodes (Clusiaria) nitida

Frey 1960: 26
Melander 1924: 20
1924
Loc

Clusiodes (Clusiaria) orbitalis

Frey 1960: 26
Johnson 1925: 245
Melander 1924: 21
1924
Loc

Clusiodes (Clusiaria) terminalis

Frey 1960: 26
Johnson 1925: 246
Melander 1924: 22
Melander 1924: 22
1924
Loc

Clusiodes (Columbiella) verticalis

Schacht 2004: 276
Beuk 2002: 271
Rohacek 1995: 133
Smith 1989: 103
Stackelberg 1988: 505
Greve 1986: 91
Withers 1985: 63
Soos 1984: 12
Greve 1983: 55
Stubbs 1982: 92
Trojan 1962: 26
Kloet 1945: 394
Czerny 1928: 9
Frey 1928: 109
Melander 1924: 24
1924
Loc

Clusiaria Malloch 1922 : 49

Beuk 2002: 271
Schumann 1999: 144
Caloren 1998: 263
Sasakawa 1987: 101
Soos 1984: 12
Stubbs 1982: 89
Trojan 1962: 22
Frey 1960: 16
Kloet 1945: 394
Neave 1939: 770
Czerny 1928: 6
Johnson 1925: 245
Malloch 1922: 49
1922
Loc

Clusiodes americana

Malloch 1922: 48
1922
Loc

Clusiodes (Columbiella) apiculata

Johnson 1925: 246
Melander 1924: 23
Malloch 1922: 49
1922
Loc

Clusiodes (Columbiella) nigripalpis

Malloch 1922: 49
1922
Loc

Clusiodes (Columbiella) johnsoni

Steyskal 1965: 807
Johnson 1925: 246
Melander 1924: 23
Malloch 1922: 49
1922
Loc

Clusiodes (Columbiella) nigripalpis

Johnson 1925: 246
Malloch 1922: 49
1922
Loc

Clusiodes (Clusiaria) duplicata

Melander 1924: 19
Malloch 1922: 49
1922
Loc

Clusiodes (Columbiella) orbitalis

Malloch 1922: 50
1922
Loc

Clusiodes albimana

Johannsen 1928: 844
Malloch 1918: 6
1918
Loc

Clusiodes albimana

Dunk 1996: 422
Smith 1989: 103
Ferrar 1987: 122
Zuska 1987: 235
Greve 1986: 89
Withers 1985: 63
Chandler 1973: 340
Hennig 1952: 227
Tuomikoski 1936: 184
Seguy 1934: 352
Richards 1934: 2
Stackelberg 1933: 381
Czerny 1928: 8
Wahlgren 1917: 187
1917
Loc

Clusiodes apicalis

Kahanpaa 2014: 288
Rotheray 2013: 404
Lonsdale 2007: 135
Poole 1996: 126
Dunk 1996: 422
Ceianu 1989: 29
Greve 1987: 78
Seguy 1934: 352
Stackelberg 1933: 381
Tuomikoski 1933: 18
Malloch 1918: 6
Wahlgren 1917: 187
1917
Loc

Clusiodes ruficollis

Kahanpaa 2014: 288
Rotheray 2013: 405
Nilsson 2009: 5
Rohacek 2009: 218
Lonsdale 2007: 141
Rohacek 2001: 338
Merz 1998: 276
Poole 1996: 126
Woznica 1991: 208
DeBruyn 1991: 146
Ceianu 1989: 29
Zuska 1987: 235
Kristofik 1986: 165
Vanhara 1981: 30
Hennig 1938: 125
Tuomikoski 1936: 183
Seguy 1934: 353
Stackelberg 1933: 381
Wahlgren 1917: 186
1917
Loc

Clusiodes (Clusiodes) albimana

Greve 1986: 89
Stubbs 1982: 92
Hackman 1980: 148
Trojan 1962: 25
Kloet 1945: 394
Frey 1928: 109
Melander 1924: 15
Johnson 1913: 98
1913
Loc

Clusiodes geomyzina

Dunk 1996: 422
Ferrar 1987: 122
Zuska 1987: 235
Maca 1985: 397
Hennig 1952: 227
Tuomikoski 1936: 183
Seguy 1934: 352
Stackelberg 1933: 381
Malloch 1918: 6
Wahlgren 1917: 186
Johnson 1913: 97
1913
Loc

Clusiodes (Clusiodes) albimana

Frey 1960: 26
Johnson 1913: 98
1913
Loc

Clusiodes melanostoma

Poole 1996: 126
Griffiths 1972: 214
Malloch 1918: 5
Johnson 1913: 98
1913
Loc

Clusiodes pictipes

Kahanpaa 2014: 288
Rohacek 2008: 154
Lonsdale 2007: 140
Rohacek 2001: 338
Poole 1996: 126
DeBruyn 1991: 146
Stackelberg 1988: 505
Malloch 1933: 18
Malloch 1918: 7
Wahlgren 1917: 187
Johnson 1913: 98
1913
Loc

Heteroneura caledonica

Collin 1912: 106
1912
Loc

Heteroneura gentilis

Lonsdale 2007: 132
Collin 1912: 107
Collin 1912: 107
1912
Loc

Heteroneura verticalis

Collin 1912: 107
Collin 1912: 107
1912
Loc

Heteroneura albimanus

Aldrich 1905: 570
1905
Loc

Clusiodes

Evenhuis 2016: 70
Kahanpaa 2014: 288
Lonsdale 2007: 123
Sueyoshi 2006: 6
Papp 2006: 190
Greve 2005: 3
Beuk 2002: 271
Marshall 2000: 4
Caloren 1998: 263
Woznica 1991: 208
DeBruyn 1991: 146
Balthazar 1990: 416
Ferrar 1987: 121
Stubbs 1982: 89
Sasakawa 1977: 236
Cole 1969: 426
Steyskal 1965: 807
Soos 1964: 7
Trojan 1962: 22
Frey 1960: 18
McAlpine 1960: 81
Hennig 1952: 227
Kloet 1945: 394
Neave 1939: 770
Seguy 1934: 351
Stackelberg 1933: 381
Tuomikoski 1933: 15
Czerny 1928: 6
Schulze 1927: 730
Melander 1924: 13
Malloch 1918: 5
Wahlgren 1917: 186
Johnson 1913: 97
Coquillett 1910: 525
Coquillett 1904: 93
1904
Loc

Heteroneura apicalis

Becker 1905: 39
Czerny 1903: 79
1903
Loc

Heteroneura flavifascies

Poole 1996: 126
Slosson 1897: 239
Slosson 1897: 239
1897
Loc

Heteroneura albimana

Osten 1878: 197
1878
Loc

Heteroneura melanostoma

Aldrich 1905: 570
Czerny 1903: 77
Osten 1878: 198
Loew 1864: 260
1864
Loc

Heteroneura pictipes

Schiner 1862: 37
1862
Loc

Heteroneura albimana var. pallidior

Lonsdale 2007: 129
Czerny 1903: 81
Loew 1864: 340
Loew 1857: 52
1857
Loc

Heteroneura albimana var. obscurior

Lonsdale 2007: 129
Czerny 1903: 81
Loew 1864: 339
Loew 1857: 52
1857
Loc

Heteroneura alpina

Czerny 1903: 77
Girschner 1888: 97
Loew 1864: 344
Loew 1857: 53
1857
Loc

Heteroneura albimana

Loew 1864: 339
Zetterstedt 1852: 4817
1852
Loc

Heteroneura pictipes

Czizek 1910: 112
Aldrich 1905: 571
Becker 1905: 39
Czerny 1903: 80
Slosson 1897: 239
Schiner 1864: 72
Zetterstedt 1852: 4816
1852
Loc

Agromyza geomyzina

Zetterstedt 1848: 2787
1848
Loc

Heteroneura geomyzina var. apicalis

Zetterstedt 1848: 2789
1848
Loc

Heteroneura laterella Zetterstedt 1848 : 2791

Loew 1864: 343
Loew 1857: 52
Zetterstedt 1848: 2791
1848
Loc

Heteronevra Nubila. Blanchard 1840 : 628

Blanchard 1840: 628
1840
Loc

Heteronevra

Evenhuis 2016: 70
Schulze 1930: 1540
Scudder 1882: 161
Blanchard 1840: 628
Macquart 1835: 587
1835
Loc

Heteronevra nubile

Macquart 1835: 587
1835
Loc

Heteronevra albimana

Macquart 1835: 588
1835
Loc

Heteroneura albimana Meigen 1830 : 128

Lonsdale 2010: 1042
Lonsdale 2009: 144
Lonsdale 2007: 129
Czizek 1910: 112
Czerny 1903: 81
Becker 1902: 313
Girschner 1888: 97
Brauer 1883: 92
Schiner 1862: 38
Loew 1857: 51
Vollenhoven 1856: 114
Curtis 1855: 4603
Rossi 1849: 78
Zetterstedt 1848: 2790
Boie 1847: 331
Meigen 1830: 128
1830
Loc

Heteroneura nubila

Czerny 1903: 76
Becker 1902: 313
Loew 1857: 53
Wilson 1855: 257
Rossi 1849: 78
Meigen 1830: 127
1830
Loc

Heteroneura ruficollis

Czerny 1903: 75
Becker 1902: 313
Schiner 1862: 38
Loew 1857: 53
Zetterstedt 1848: 2789
Meigen 1830: 128
1830
Loc

Heteroneura Fallén 1823 : 2

Lonsdale 2009: 144
Neave 1939: 640
Neave 1939: 640
Schulze 1930: 1540
Hendel 1910: 309
Kertesz 1906: 320
Aldrich 1905: 571
Czerny 1903: 73
Becker 1902: 313
Strobl 1897: 27
Williston 1896: 386
Osten 1878: 197
Osten 1878: 197
Schiner 1862: 37
Wilson 1855: 256
Rossi 1849: 78
Agassiz 1846: 519
Westwood 1840: 152
Haliday 1838: 188
Meigen 1830: 126
Fallen 1823: 2
1823
Loc

Heteroneura geomyzina

Zetterstedt 1848: 2787
Fallen 1823: 3
1823
Loc

Heteroneura geomyzina Fallén 1823 : 2

Czizek 1910: 112
Czerny 1903: 77
Becker 1902: 313
Brauer 1883: 92
Schiner 1862: 38
Loew 1857: 54
Zetterstedt 1848: 2787
Meigen 1830: 129
Fallen 1823: 2
1823
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