Parapenetretus (Propenetretus) selaensis, A.S & D.W & A & H, 2006

A. S, D. W, A, W & H, E, 2006, New taxa of the tribe Patrobini (Coleoptera, Carabidae) from the East Himalayas, Linzer biologische Beiträge 38 (1), pp. 991-1008 : 998-999

Parapenetretus (Propenetretus) selaensis spec. nov. ( figs 2 , 4, 6, 8 , 14-18 , 20 )

T y p e m a t e r i a l: Holotype: (DW), NE India, W Arunachal Pradesh, Sela Pass, SE of Tawang, 27 o 30'N 92 o 06'E, 4200-4400 m, 29.-31.V.2004, R. Businský. Paratypes: 4, 2 (JB), 1 (DW), 1, 2 (AZ), 3, 2 (PB), same locality, together with holotype. 11, 17 (NHMB), 3, 4 (DW), 2, 4 (AZ), same locality but: 4400 m, 31.V.2004, L. Dembický.

D e s c r i p t i o n: Habitus as in fig. 2 . Body dark brown, shiny, antennae, mandibles, and legs brown, palpi brown to reddish-brown. Slender, with long appendages. Total length 9.7-11.2 mm.

Head ovate, 0.78-0.82 times wider than pronotum; eyes rather large, moderately convex; tempora about as long as eye diameter, moderately to faintly tumid; neck-constriction deep; frontal furrows rather deep, moderately divergent posteriad; surface mostly smooth, densely punctate in frontal furrows, neck-constriction finely and sparsely punctate; 2, very seldom 3 setiferous pore punctures present on each side, anterior one situated in supraorbital groove nearly at mid-eye level, median (present in 1 ex. only) between posterior margin of eye and neck constriction, and posterior, represented as a rather large setiferous foveole, beside neck-constriction; besides long anterodorsal seta antennomere 1 seldom bears 1-2 additional minute setae; tooth of mentum without peculiarities, bifid ( fig. 4 ), epilobes narrow, median area of mentum moderately elevated, lateral lobes separated from it by rather deep oblique furrows, submentum with 2 setae on each side.

Pronotum transverse, cordate, 1.25-1.38 times wider than long, weakly convex; front margin almost straight; sides widely rounded, weakly sinuate just before hind angles, with rather broad explanate margin; basal margin nearly straight to weakly rounded; front angles distinct, strongly projecting anteriad, rounded; hind angles usually obtuse, seldom rectangular, pointed, more or less distinctly denticulate, basal carina absent; anterior transverse impression rather deep to shallow, coarsely punctate; basal foveae rather deep, densely and coarsely punctured; disk faintly wrinkled to smooth, coarsely punctate in basal area and lateral gutters, median line obliterated at both extremities; 2 marginal setae (seldom 3), basal seta present in hind angles.

Front margin of metasternum with prominent swelling; pro- and metepisterna, lateral sides of metasternum densely and rather smoothly punctured, punctures being of medium size, mesosternum and mesepisterna covered by more or less dense, large, sharp punctures, lateral areas of sternite 1 (urite 3) rugous-punctured, other sternites almost smooth, finely wrinkled.

Elytra oblong-ovate, 1.53-1.59 times longer than wide and 1.41-1.52 times wider than pronotum, widest in the middle, weakly convex; sides somewhat rounded, narrowly bordered, margin gradually tapering behind; humeri rather narrow but prominent, rounded and indistinctly denticulate; striae deep and distinct, basally finely punctured; intervals weakly convex, interval 3 with 3-4 setiferous pore punctures adjoining stria 3; marginal series composed of 12-16 small setiferous pore punctures, rarefied in the middle; microsculpture composed of fine transverse meshes and wrinkles and very sparse and fine microscopic puncturation. Hind wings reduced.

Aedeagus ( figs 14, 15 ) sharply bent at base; apical lamella moderately curved ventrally (viewed laterally), rather sharply attenuating towards apex, which is narrowly rounded to nearly pointed (viewed dorsally), with long horn-shaped protuberance on the right side dorsally; armature of endophallus composed of large poorly sclerotized bilobed proximal copulatory piece, its distal ending gradually transforming to membranous folded structures; left paramere ( fig. 16 ) larger than right one ( fig. 17 ), their apical projections rather short but distinct, bearing 2-3 long apical and 0-3 smaller or minute subapical setae.

ventrite 9 as in fig. 18 .

Female reproductive tract as in fig. 20 ; distal sclerite of female reproductive tract large, of regular symmetric shape; bursal sclerite large, elongate, of complicated irregular shape, its length at maximum diameter ca. 1.1 mm.

E t y m o l o g y: The specific epithet refers to the name of locality where this species was collected, the Sela Pass.

Figures

Figs 1-2: Patrobini, general view. Indopatrobus bashtai spec. nov., paratype, (1). Parapenetretus selaensis spec. nov., paratype, (2).

Figs 3-8: Patrobini, details of external morphology (3-4 - mentum, dorsal view, 5-6 - lateral part of thorax, lateroventral view, 7-8 - female tergite 8, dorsal view, ac2 - middle coxal cavity, epm2 - mesepimeron, epm3 - metepimeron, eps2 - mesepisternum, eps3 - metepisternum, st2 - mesosternum, st3 - metasternum). Indopatrobus bashtai spec. nov., paratype, (3, 5, 7). Parapenetretus selaensis spec. nov., paratype, (4, 6, 8). Scale bar: 1 mm.

Figs 9-18: Patrobini, male genitalia (9, 14 - aedeagus, left lateral view, 10, 15 - aedeagus, dorsal view, 11, 16 - left paramere, left lateral view, 12, 17 - right paramere, right lateral view, 13, 18 - ventrite 9, dorsal view). Indopatrobus bashtai spec. nov., paratype, (9-13). Parapenetretus selaensis spec. nov., paratype, (14-18). Scale bar: 1 mm.

Figs 19-20: Patrobini, reproductive tract (bc - bursa copulatrix, bs - bursal sclerite, sg - spermathecal gland, sp - spermatheca). Indopatrobus bashtai spec. nov., paratype, (19). Parapenetretus selaensis spec. nov., paratype, (20). Scale bar: 1 mm.