Montenegrina skipetarica puskasi, Feher, Zoltan & Szekeres, Miklos, 2016

Feher, Zoltan & Szekeres, Miklos, 2016, Taxonomic revision of the rock-dwelling door snail genus Montenegrina Boettger, 1877 (Mollusca, Gastropoda, Clausiliidae), ZooKeys 599, pp. 1-137 : 89-91

Taxon classification Animalia Stylommatophora Clausiliidae

Montenegrina skipetarica puskasi ssp. n. Fig. 30A

Diagnosis.

Medium-size subspecies with very narrowly attached peristome, dorsolateral lunella and often uninterrupted plica superior with long anterior part.

Description.

The shell is medium- to large-sized, light brownish-corneous, somewhat tumid, of its 91/2 to 11 whorls the lower three are almost almost of the same width. The surface is smooth at the lower whorls but becomes indistinctly costate over the short apex. The weakly inflexed neck has dense, sharp ribs behind the peristome. The basal crest is weak, the peripheral one is well visible. The light brown, ovoid peristome has simple, narrowly attached (rarely somewhat detached) margin. The lamellae superior and spiralis overlap. The lamella inferior is emerged, well visible in front view. The broadly bent lamella subcolumellaris is not visible in frontal, only in slanted view at the aperture. The lunella is dorsolateral, broad, mostly fused to the basalis that is as long as the subclaustralis. The sulcalis is week. The anterior part of the plica superior is long, ends in a lump behind the aperture and often also reaches the lunella complex. Part of the clausilium plate is visible through the aperture.

Dimensions

(in mm). Holotype Hs: 18.9, Ws: 5.2, Ha: 4.8, Wa: 4.05; paratypes Hs: 16.8-19.7 (mean 18.7, SD 0.76), Ws: 4.5-5.4 (mean 4.9, SD 0.28), Ha: 4.4-5.3, Wa: 3.5-4.2 (n = 12).

Differential diagnosis.

Distinguishable from Montenegrina skipetarica ersekensis and Montenegrina skipetarica nobilis by the retracted, frontally not visible lamella subcolumellaris, and from all other Montenegrina skipetarica subspecies by the cylindical lower whors that are nearly of the same width. From the nearby-occurring and apparently closest related Montenegrina skipetarica gurelurensis ssp. n. differs by the smooth shell and attached peristome.

Type locality.

Albania, Dibrë District, Arras, 2 km from the Kukës to Peshkopi road toward Cidhnë, 460 m, 41.7391°N, 20.2970°E.

Type material.

Type locality, leg. TD, ZE, ZF, DM, 10.x.2005, holotype (HNHM 99448), paratypes (HNHM 99449/32, MMM-B01318/32, ER/32, SZ/4); type locality, leg. ZE, ZF, AH, DM, 12.iv.2006 (HNHM 99450/7, ER/7, HU/7); type locality, leg. ZE, ZF, JG, 1.vii.2015, (HNHM 99451/18, NHMW 111206/18, GR/18).

Other material.

Albania, Dibrë District, beneath Cidhnë, gorge of the Setë Stream at the lower hydroelectric station, 510 m, 41.7506°N, 20.2626°E, leg. TD, ZF, 10.x.2005 (HNHM 99452, MMM-B01319); 1 km W of Cidhnë, along the footpath to Gurë-Lurë, gorge of the Setë Stream, 680 m, 41.7518°N, 20.2611°E, leg. TD, ZE, ZF, DM, 10.x.2005 (HNHM 99453, MMM-B01320); same locality, leg. ZE, ZF, AH, DM, 12.iv.2006 (HNHM 99455); ca. 1.5 km W of Cidhnë, 41.7528°N, 20.2505°E, leg. PJ, TK, DM, GP, 8.x.2012, (HNHM 99457); ca. 2 km W of Cidhnë, 730-750 m, leg. ZE, DM, 10.x.2005, (HNHM 99454, MMM-B01321); 0.1-1.5 km W of Cidhnë, leg. ZE, ZF, JG, 1.vii.2015 (HNHM 99458, NHMW 111207); Mat district, Mali i Dejës, limestone hill of 1700 m height above the headwaters of the Lumi i Varoshit, 1540 m, 41.6746°N, 20.2112°E, leg. ZF, DM, ZU, 18.v.2010 (HNHM 99691).

Etymology.

The new taxon is named after Gellért Puskás, entomologist (HNHM), who collected invaluable mollusc material during his Balkan field trips.

Distribution.

Southern part of the Lura-Dejë mountain group in northern Albania. This taxon was found at some nearby localities along the Setë Stream (Fig. 31B).