Oedichirus longipennis KRAATZ

Rougemont, Guillaume de, 2018, New oriental Oedichirus (Staphylinidae, Paederinae, Pinophilini), Linzer biologische Beiträge 50 (1), pp. 461-536 : 483-487

Oedichirus longipennis KRAATZ (Figs 14)

Oedichirus longipennis KRAATZ, 1859: 154 .

Oedichirus idae SHARP, 1874: 76 nov.syn.

Oedichirus schultheissi FAUVEL, 1895: 217 nov.syn.

Oedichirus excellens CAMERON, 1925: 180 nov.syn.

Oedichirus longipennis CAMERON 1932: 26.

Oedichirus idae SHIBATA et al. 2013: 161.

Oedichirus longipennis SABATINELLI 2015.

M a t e r i a l s t u d i e d: ♀ Holotype: India or. / Holotypus / Oedichirus longipennis Krz. / Coll. Kraatz / DEI Münchenberg Col. – 07524 / Oedichirus longiceps [sic] Kr. / Oedichirus longipennis Kr. det. 2017 G. de Rougemont [DEI].

A d d i t i o n a l m a t e r i a l:

India: 1♂: W. Almora, Kumaon, U.P., India, H.G.C. / Oedichirus sp. / Oedichirus longipennis / G.C. Champion BM 1927-409 [NHML]; 1♂: W. Almora Div. Kumaon U.P. Mar. 1918, HGC / C.G. Champion coll. BM 1927-409 / Oedichirus longipennis Kr. det. 2015 G. de Rougemont [NHML]; 1♂: Haldwani Dist. Kumaon, HGC / C.G. Champion coll. BM 1927-409 [NHML]; 1♀: N-INDIA: Uttaranchal state, Nainital, China Peak env. 1900-2300m, 18- 19.7.2003 leg. Z. Kejva & M. Tryzna [NHMW]; 1♀: N-INDIA, 8/1986, Rishikesh, F. Werner / Pinophilini spec. DET. M. SCHÜLKE 19.. / Oedichirus spec. cf. longipennis Kr. det. M. Schülke 1996 / Sammlung M. Schülke Berlin / Oedichirus longipennis Kr , det. 2015 G. de Rougemont [CSB]; 1♀: INDIA: Him. Pr. Patikuhl Town, 28-29.v.1999, 3207.4’N 7708.8’E, 1200 m, Y. Marusik leg., FIELD MUS. NAT. HIST. / Oedichirus longipennis Kr. Det. 2016 G. de Rougemont [FMC]; 1♀: INDIA, 23 -24.VI.1995, Barpata Road, Assam, Werner leg. / Rougemont collection [CRO]; 4♂♂ & 2♀♀: S-INDIA, Karnataka state, Coorg distr. NE of Virajpet, 75º13’E 12º13’N, ca. 500 m, 4-8.vi.1999, Z. Kejval & M. Trysna leg. / Oedichirus longipennis det. 2016 G. de Rougemont [CJN]; 1♂: Shimoga [= Shivamogga], India / Oedichirus longipennis Kr. det. 2017 G. de Rougemont [MNHN]; 2♂♂: S- INDIA, Kerala state, Kallar env., 30 km NE of Trivandrum, valley of riv. Kallar 77º05’E 8º45’N, 7-13.v.1999, Z. Kejval & M. Trysna leg. / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [CJN].

Pakistan: 1 ex.: Pakistan, Islamabad, 33º43’N 73º03’E, Guido Sabatinelli lgt., Oedichirus longipennis Kraatz, J. Matejiček det.

Nepal: 1♀: NEPAL. Chaur , 600 m., XI.1987, P. Morvan / Oedichirus longipennis Kr. det. 1988 G. de Rougemont [ CRO] ; 1♂: NEPAL, 500 m. Bhairabsthan, XI.1987, P. Morvan [ CRO] ; 1 ex.: NEPAL: 300 m, Dolalghat i. xii.1982 / Lt. Col. M.G. Allen, Brit. Mus. 1983-254 / Oedichirus longipennis Kr. det. Willers 7.99 [ NHML] .

Sri Lanka : 1♀: CEYLON, Wilpattu Nat. P., 18.VII.79, G. de Rougemont / Oedichirus longipennis Kr. det. G. de Rougemont [ CRO] ; 1♂ & 1♀: SRI LANKA: Kandy , Peak View Motel (1800’) / 7-14-I- 1970 Davis & Rowe / OEDICHIRUS det. Newton 1993 / Oedichirus longipennis Kr. Det. 2006 G. de Rougemont [ FMC] ; 1♂: SRI LANKA: Kandy Distr. , 5 mi NW Mahiyangana Hasalak Irrigation Bungalow, 30-III-9-IV-1971, P. & P. Spangler / at black light / Oedichirus longipennis Kr. Det. 2016 G. de Rougemont [ FMC] ; 1♀: SRI LANKA, Galle Dist. , 11 mi E Udugama, Kanneliya jungle 12-X-1973 / Krombein, Karunaratne, Fernando & Fernandino [ FMC] .

Malaysia: 2♂♂: PENANG, 4.XI.90, Rougemont / Oedichirus longipennis Kr. det. 1990 G. de Rougemont [CRO]; 1 ex.: MALAYA, SELANGOR, F.M.S., Bukit Chinalcah July 18 th 1921 / Ex. F.M.S. Museum B.M. 1955-354 [NHML]; 1♂: MALAYSIA – Pahang / Johor, Endau-Rompin n. park 100m, Salendang 28.II-12.III.1995, M. Strbat & R. Hergovitz leg. / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [NHMW]; 1♀: MALAYSIA, W. Ipoh, 5 km of Tanjong Rambutan, 13-15.IV.2000, leg. M. Snižek / Oedichirus spec. det. M. Schülke 2015 / Oedichirus longipennis det. 2015 G. de Rougemont [CSB].

Singapore: 1 ex.: Res. W. 2/1/23 cut grass / Singapore C.J. Saunders, BM 1929-369 / Oeddichirus longipennis det. 2016 G. de Rougemont [NHML].

China: 1♂: CHINA, Yunnan, Xishuangbanna, Jinghong , I.1993, G. de Rougemont / Oedichirus longipennis Kr. det. 1993 G. de Rougemont [ CRO] ; 1 ex.: Shanghai [ Department of Biology , Shanghai Normal University] ; 1♂: Hong Kong, Wang Tong, Lantau , 7.X.2012, at light, leg. Paul Aston [Aston collection, Hong Kong] .

Taiwan: 1♀: TAIWAN: Nantou Hsien, Hwy 14, Fengnan 700m, 22.IV.90, A. Smetana (T17) / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [AMNH].

Thailand: 1♀: THAILAND: 10.2.1992, Ban Kiriwong, Khao Luang Nat. Park 170 m, P. Schwendinger / Oedichirus longipennis Kr. det. 2015 G. de Rougemont [MHNG]; 2♀♀: THAILAND, 240 km nw Bangkok, 110m. leg. Thielen / 25 km nw Lam Sek, Lichtfang [NHMW]; 1♀: THAILAND, Chiang M., NW Chiang Mai, Pai City, 29.4.1993, Patcholatko & Dembicky [NHMW]; 1♂ & 1♀: N-THAILAND, Nan, Lom Sak – Dan Sai, 17.- 19.5.1993, Patcholatko & Dembicky [NHMW]; 2♀♀: THAILAND, Nan Prov., Ban Bo Klua env., 13-26.v.2002, P. Průdek leg. [NHMW]; 1♀: Nan prov. Ban Huay Kon env., 27.v-10.vi.2002, P. P. Průdek leg. [NHMW]; 1♀: Thailand, Khao Kai 400 m, 12.1990 / Oedichirus spec. det. M. Schülke 2015 / Sammlung M. Schülke Berlin / Oedichirus longipennis Kr. det. 2015 G. de Rougemont [CSB]; 1♂: SAN SAI, CHIANG MAI, THAI, 1.V.1990, T. & N. ITO / Oedichirus longipennis Kraatz Det. TATEO ITO 1993 [CIK]; 1♀: MON ANGGET, CHIANG MAI, THAILAND, 28.IV1992, T. & N. ITO / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [CIK].

Laos: 1♂: Zentral-LAOS, 70 km NE Vientiane, Ban Phabat , 18º16.1N 102º10.9E (GPS), 150 m, 27.IV- 1.V.1997, leg. C. Holzschuh / Oedichirus longipennis Kr. det. 2015 G. de Rougemont [ CRO] ; 1♀: LAOS centr., Bolikhamsai prov., BAN NAPE, - Kaew Nua Pass , 18.4- 1.5.1998, alt. 600± 100m, N 18º22.3 N 105º09.1 E (GPS), E. Jendek & O. Sauša leg. / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [ CST] ; 1♀: N-LAOS, May 1999, Oudom Xai prov. , NAM MIANG riv., 30km S Muang Xai, 1200m., Lao coll. leg. / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [ CST] ; 2♀♀: LAOS centr., 27.IV-1.V.1997, 70 km NE Vientiane, BAN PHABAT env., 150 m, N 18º16.1, E 103º10.9, E. Jendek & O. Sauša leg. / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [ CIK] .

Vietnam: 1♂: S-VIETNAM, 40 km NW An Khe, Buon Luoi, 620-750 m, 14º10’N 108º30’E, 28.3- 12.4.1995, Patcholatko & Dembicky / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [NHMW].

Indonesia: 6♀♀: SULAWESI TENGAH, Nr. Morowali, Ranu River Area, 27.i-20.iv.1980 / at light / M.J.D. Brendell BM 1980-280 / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [NHML, 1 ex. in CRO]; 1♀: Indonesia, Sulawesi Teng., 5Km W of Bancea (Danau Poso), 900 m, 120.32.4E 1.59.3S, 19-24.vii.1999 / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [CST]; 1♀ [type of O. schueltheissi Fvl.]: N.O. Sumatra, Tebing-tinggi, Dr. Schultheiss / Schultheissi Fvl. / lewisius Shp. var. / Ex-Typis / Oedichirus longipennis Cam. New Synonymy des. 2015 G. de Rougemont [IRScNB]; 1♀ [type of O. excellens Cam.]: KL Kombuis Java-Zee, 27.I.1928 / [BM Paratype label] / Oedichirus excellens Cam. / M. Cameron Bequest 1955-147 / Oedichirus longipennis Kr. New Synonymy des. 2015 G. de Rougemont [NHML]; 1♀: C.J. Louwerens Java 84 m, Toeloengagoeng / Oe. ab. nigripennis Cam. / M. Cameron Bequest BM 1955-147 / Oedichirus longipennis Kr. det. 2015 G. de Rougemont [NHML]; 1♀: INDONESIA, E. Java, Baluran Nat. P. 50 m, Wonorejo, 24- 25.2.1994, Bolm leg. [SMNS].

Japan: 1 ex. (Type of O. idea Sharp): Mitzuyama, Nagasaki / Oedichirus longipennis Kr. New Synonymy Des. 2016 G. de Rougemont [ NHML] ; 1♀: (HONSHU) Hirakawa , Yamaguchi / 1.VII.1981, K. Yano et al., at light / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [ KUM] ; 1♂: OKINAWA Is., Okinawa Pref., Nago C., Ooura , 26.VII.1993, M. Kimura leg. [ KUM] ; 1♀: SHIRAHAMA IRIOMOTE Is. , 20.XII.1996, I. MATORA leg. / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [ CIK] ; 1♂: JAPAN, Ryukyus, Yaeyama Iss., MIYAKO Is. , 29.II.1997, I. MATOBA leg. / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [ CIK] ; 1♂: JAPAN Ryukyus, Mt. Urabu-dake, YONAGUNI Is. , 27.II.2007, I. TANAKA leg. / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [ CIK] ; 1♂: YOSHIWARA, ISHIGAKI Is. , OKINAWA, 10.III.2006, Isamu TANAKA / Ordichirus longipennis Kr. det. 2016 G. de Rougemont [ CIK] ; 1♂: JAPAN, Ryukyus, Naha 20-100m, OKINAWA Is., 28.VI.2002002, K. KUSANO leg. / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [ CIK] ; 1♀: JAPAN, Ryukyus, Aha Kunigami, OKINAWA Is. , 23.I.2008, S. IMASAKA leg. / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [ CIK] ; 1♀: Kamijijima Is., Yaeyam Iss. , Okinawa Pref., 20.VI.2011, Y. Kusui Leg. / Oedichirus longipennis Kr. det. 2016 G. de Rougemont [ CIK] .

D e s c r i p t i o n (of type): length: 7 mm; length of fore-body: 3.2; length of head: 0.75; breadth of head: 0.92; length of antenna: 1.65; length of pronotum: 1.1; breadth of pronotum: 0.9; length of elytron: 1.32; breadth of elytra: 1.3. Head black, pronotum red, elytra black, abdominal segments III-VI red, the following black; palpi and antennae testaceous, legs testaceous, knees infuscate. Dorsal surfaces devoid of microsculpture. Pubescence pale, fairly long and dense, erect on head, pronotum and abdomen, decumbent on elytra. Habitus: Figs 14h (ex. from Laos), 14hjp (type of O. ideae).

Head very transverse; post-ocular groove well marked by series of small punctures, but carina feeble, not forming an angle behind eyes; puncturation sparse, consisting of a number of very small punctures on frons, 5-6 larger punctures on vertex, 4-5 ocular punctures, the posterior one large, and transverse series half a dozen punctures just before base, in addition to the very small punctures in post-ocular furrow. Pronotum strongly elongate, the sides convergent in straight lines from anterior angles to base; lateral margins not bordered; disc with a pair of series of five punctures each preceded by a pair of smaller punctures set more closely together than discal series, and near the anterior margin a series of six punctures disposed in an arc, two large lateral punctures, and the usual small punctures along lateral margins. Elytra elongate, ample, widest at posterior 1/3 rd, the sides rounded between well marked humeral angles and acute posterior angles, the joint posterior margin coarctate; puncturation coarse and close, the interstices in anterior ¾ almost everywhere narrower than diameter of punctures. Puncturation of abdominal tergites arranged in three discrete transverse rows and an anterior transverse row adjacent to anterior row of keels.

Male: abdominal sternite VII unmodified; sternite VIII: Fig. 14s8; aedoeagus: Figs 14arl, av.

Female: sternite IX and vulvar plate (of a specimen from Japan): 14vp.

Variability: the extent of infuscation of the knees varies; in most specimens from Japan and the Ryukyu Islands (= O. idea SHARP), the legs are uniformly testaceous; elsewhere the knees are more or less extensively infuscate. In the female from Galle, Sri Lanka, the abdominal segments III-VI are very dark, almost black. The type of O. schultheissi FAUVEL does not differ in any way from typical O. longipennis ; FAUVEL seems to have been unaware of KRAATZ’ species, for he compared O. schultheissi to his own West Palearctic species O. walkeri. In BERNHAUER & SCHUBERT’ s catalogue (1912, 40: 202) the home of O. schultheissi (Sumatra) is erroneously given as Burma, probably because Fauvel’s article is entitled "Staphylinides nouveaux de l’Inde et de la Birmanie ". The specimen from "Kombuis" is the type of O. excellens CAMERON, but does not bear a CAMERON Type label. The specimen only differs from typical O. longipennis by being generally paler: the knees are less extensively infuscate than in typical specimens, and the centre of each elytron is broadly reddish.

O. longipennis is very close to O. lewisius SHARP, from which it is most evidently distinguished by its more densely punctate and larger elytra (distinctly longer than pronotum), differences in shape of the emargination of male 8 th sternite and of the anterior process of the ventral sclerite of the aedoeagus. The ranges of the two species overlap in southern Honshu, Kyushu and the Ryukyu Islands.

O. longipennis is the commonest and by far the most widespread species of the genus (distribution map, below) with a range that covers the entire Oriental Region except, as far as we know, the Philippines. It was previously recorded from northern and southern India and Pakistan, and, as O. lewisius , from Java (RATNA et. al. 2002) Shanghai ( LI et al. 2010) and Guangxi ( WANG 1990). O. ‘ idae ’ was described from Tokunoshima Island and recorded from Honshu and Kyushu. Rather surprisingly SHARP did not consider O. longipennis when he described O. ‘idea’, and CAMERON also omitted mention of Sharp’s two species when he redescribed O. longipennis (1932) ; it is more easily understandable that later Chinese and Indonesian authors looked to the far-eastern species when they recorded O. lewisius from China and Indonesia rather to the less well known O. longipennis described from India. O. longipennis is recorded here for the first time from Nepal, Ceylon, Thailand, Laos, Vietnam, China, Taiwan, Malaysia, Singapore and Celebes.

D i s t r i b u t i o n: Map 1 . Keys: 2, 3, 4.

Figures

Map 1: Distribution of Oedichirus longipennis (records from Honshu and Kyusyu omitted. Empty circles represent specimens not seen, recorded as O. lewisius).