Tonsilla subyanlingensis K. Liu & X. Xu, 2020

Liu, Ke-ke, Luo, Hui-pu, Xu, Xiang, Chen, Zhiwu & Xiao, Yong-hong, 2020, Description of two new species of Tonsilla Wang & Yin, 1992 with an updated key to species (Araneae, Agelenidae), ZooKeys 944, pp. 31-46 : 31

publication ID	https://dx.doi.org/10.3897/zookeys.944.48575
publication LSID	lsid:zoobank.org:pub:7FC65CCE-E408-4918-B238-073881E2961A
persistent identifier	https://treatment.plazi.org/id/64760314-489A-4A5C-AFA3-3D857B92ED14
taxon LSID	lsid:zoobank.org:act:64760314-489A-4A5C-AFA3-3D857B92ED14
treatment provided by	ZooKeys by Pensoft
scientific name	Tonsilla subyanlingensis K. Liu & X. Xu
status	sp. nov.

Treatment

Tonsilla subyanlingensis K. Liu & X. Xu sp. nov. Figures 4 View Figure 4 , 5 View Figure 5 , 6 View Figure 6 , 7 View Figure 7 , 8 View Figure 8

Type material.

Holotype ♂; China: Jiangxi Province, Ji’an City, Jinggangshan County Level City, Ciping Town, Wuzhi Peak Scenic Spot; 26°31'59.07"N, 114°08'28.47"E, 735 m; 2.X.2018; Ke-ke Liu et al. leg. Paratypes: 2 ♀; same data as holotype; 1 ♀; same locality, Dajing Village; 26°33'50.4"N, 114°07'26.4"E, 930 m; 19.X.2014; Ke-ke Liu et al. leg.; 1 ♀; same locality; 26°34'12.89"N, 114°07'41.87"E, 950 m; 30.IX.2018; Ke-ke Liu et al. leg.; 1 ♀; same locality, Jingzhushan Scenic Spot; 26°31'33.37"N, 114°06'30.34"E, 786 m; 1.X.2018; Ke-ke Liu et al. leg.

Etymology.

The specific name refers to its similarity to T. yanlingensis (Zhang, Yin & Kim, 2000); adjective.

Diagnosis.

Females of the new species closely resemble T. yanlingensis by the heart-shaped, large atrium and wide epigynal teeth, but can be distinguished by the spermathecae separated by less than 1/5 of their width (vs touching each other in T. yanlingensis ), long and broad copulatory ducts along with the spermathecae (vs very short in T. yanlingensis ), the slightly procurved spermathecal heads located at posterior part of spermatheca (Figs 5C, D View Figure 5 , 7H, I View Figure 7 ) (vs strong procurved spermathecal heads located at mid part of spermatheca in vulva), and the spermathecae slightly separated by less than 1/5 of their width (vs touching each other) ( Wang and Yin 1992; Zhu et al. 2017). The male of this species is similar to that of T. mopanensis and T. truculenta in having a long, broad, and furrowed basal lamella of conductor, but can be separated by the patellar apophysis which is relatively shorter than patellar (vs as long as patellar apophysis in T. mopanensis or longer in T. truculenta ) and the conductor with a long, broad curved dorsal apophysis (Figs 5A-C View Figure 5 , 7A-G View Figure 7 , 8A-C View Figure 8 ) (vs long, narrowed in T. mopanensis ; short, strong in T. truculenta ).

Description.

Male (Holotype). Habitus as in Figures 4A, B View Figure 4 , 5A, B View Figure 5 . Total length 11.25. Carapace (Fig. 5A View Figure 5 ) 6.01 long, 4.44 wide, anteriorly narrowed to between 0.6 and 0.7 its maximum width. Eye sizes and interdistances: AME 0.20; ALE 0.25; PME 0.24; PLE 0.25; AME-AME 0.10; AME-ALE 0.18; PME-PME 0.08; ALE-ALE 0.81; PME-PLE 0.36; PLE-PLE 1.16; ALE-PLE 0.10, AME-PME 0.16; AME-PLE 0.45. MOA: 0.60 long; 0.49 front width, 0.50 back width. Chelicerae (Fig. 5B View Figure 5 ) with 2 promarginal teeth (proximal smaller) and 2 retromarginal teeth (proximal larger). Leg (Fig. 5A, B View Figure 5 ) measurements: I 17.84 (4.88, 1.95, 4.46, 4.30, 2.25); II 15.78 (4.25, 1.90, 3.74, 3.74, 2.15); III 13.36 (3.62, 1.81, 2.82, 3.40, 1.71); IV 17.27 (4.87, 1.92, 3.69, 4.77, 2.02). Spination : I fe 120, pa 001, ti 055, mt 033, II fe 000, ti 044, mt 033,. Femur I with 6 ventral cusps. Leg measurements (Fig. 5A, B View Figure 5 ): I 13.15 (4.17, 1.85, 3.26, 2.54, 1.33); II 12.90 (3.62, 1.18, 2.90, 2.83, 1.71); III 11.00 (3.01, 1.72, 2.14, 2.75, 1.38); IV 14.12 (3.84, 1.87 3.45, 3.70, 1.32). Spination : I fe 210, ti 043, mt 055; II fe 210, ti 043, mt 065; III fe 121, pa 011, ti 433, mt 655, ta 011; IV fe 100, ti 342, mt 753, ta 012. Pedicel 0.32. Abdomen (Fig. 5A, B View Figure 5 ) 5.24 long, 3.68 wide.

Carapace dark brown. Chelicerae dark brown. Endites and labium dark yellow-brown. Sternum and legs yellow-brown. Abdomen dark brown with 5 pairs of yellow-brown spots on posterior half.

Palp (Figs 5C-E View Figure 5 , 7A-G View Figure 7 , 8A-C View Figure 8 ). Femur more than 2 times longer than patella. Patellar apophysis slightly shorter than patella, thumb shaped. Tibia with wide ventrolateral apophysis and long retrolateral apophysis, the former extending beyond tibia, strongly sclerotized; the later slightly shorter than ventrolateral, and twice thinner, apex bent ventrally to the tip of ventrolateral apophysis, forming a right angle with its axis. Cymbium, approximately 3 times longer than wide, cymbial furrow less than 2/3 of the cymbial length, approximately 1/3 of cymbial width in retrolateral view. Median apophysis spoon-shaped, located near the base of embolus; conductor, slightly curved, with a long, broad and furrowed basal lamella and a large, twisted, sclerotized dorsal apophysis; embolus long and broad, originates at 6 o’clock position, coiled around the margin of cymbium and posteriorly embedded in the furrow of conductor.

Female (Paratype). Habitus as in Figures 4C, D View Figure 4 , 6A, B View Figure 6 . Total length 13.21. Carapace (Fig. 6A View Figure 6 ) 6.66 long, 4.33 wide, anteriorly narrowed to between 0.7 and 0.8 its maximum width. Eye sizes and interdistances: AME 0.20; ALE 0.24; PME 0.22; PLE 0.24; AME-AME 0.10; AME-ALE 0.18; PME-PME 0.20; ALE-ALE 0.92; PME-PLE 0.41; PLE-PLE 1.46; ALE-PLE 0.14, AME-PME 0.16; AME-PLE 0.45. MOA: 0.68 long; 0.60 front width, 0.70 back width. Chelicerae (Fig. 6A, B View Figure 6 ) with 3 promarginal teeth (proximal smallest, median largest) and 3 retromarginal teeth (proximal largest). Leg measurements (Fig. 6A, B View Figure 6 ): I 13.15 (4.17, 1.85, 3.26, 2.54, 1.33); II 12.90 (3.62, 1.18, 2.90, 2.83, 1.71); III 11.00 (3.01, 1.72, 2.14, 2.75, 1.38); IV 14.12 (3.84, 1.87 3.45, 3.70, 1.32). Spination : I fe 402, ti 004, mt 044; II fe 122, pa 011, ti 035, mt 065; III fe 122, pa 111, ti 055, mt 622, ta 011; IV fe 122, pa 011, ti 055, mt 866, ta 011. Pedicel 0.46. Abdomen (Fig. 6B, C View Figure 6 ) 6.10 long, 4.01 wide.

Lighter than male. Abdomen, dorsally with four indistinct yellow-brown chevron-like stripes on posterior half.

Epigyne (Figs 6C, D View Figure 6 , 7H, I View Figure 7 , 8D, E View Figure 8 ). Atrium with a transverse depression, broad, more than 2 times longer than its length, heart-shaped, anterior margin near the apex of teeth, posterior part relatively broad. Copulatory openings located at postero-lateral of the atrium. Epigynal teeth flat, separated by less than their length, apex slightly converging. Copulatory ducts, originating laterally, extending forward along spermathecae, then back, but located at lateral part of spermathecae. Spermathecae egg-shaped, clearly separated by less than 1/5 their width. Spermathecal heads relatively broad, short, posteriorly located, curved laterally. Fertilization ducts located at the posterior part of the spermathecae.

Distribution.

Known only from the type locality in Jiangxi Province, China (Fig. 9 View Figure 9 ).