Senaspis haemorrhoa (Gerstaecker, 1871)

Meyer, Marc De, Goergen, Georg & Jordaens, Kurt, 2020, Taxonomic revision of the Afrotropical hover fly genus Senaspis Macquart (Diptera, Syrphidae), ZooKeys 1003, pp. 83-160 : 83

Senaspis haemorrhoa (Gerstaecker, 1871) Figs 5 , 28-31 , 57 , 73 , 85 , 95 , 106

Plagiocera haemorrhoa Gerstaecker, 1871: 363.

Dolichomerus griseifacies Bezzi, 1908: 381. Syn. nov.

Differential diagnosis.

A species with a distinct medial dark brown macula on the wing (Fig. 73 ). It can be differentiated from other Senaspis species with a distinct wing macula by the coloration of the abdomen: largely brown with the posterior terga conspicuously orange to orange-red and with red pile (Fig. 95 ) (without red pile in S. dentipes (Fig. 91 ); red coloration and pile more extensive in S. xanthorrhoea (Fig. 100 )).

Type.

Plagiocera haemorrhoa Gerstaecker: Syntypes, male, female [number unknown], TANZANIA, Wanga [not examined, institutional depository unknown].

Examined material.

Dolichomerus griseifacies Bezzi: Lectotype (hereby designated), male, " Senaspis // Dolichomerus haemorrhoa / det. C. Kassebeer 1996" " Dolichomerus // Dolichomerus griseifacies // ♂ n.sp." "Cf. Ann. Soc. Ent. Belg // Vol. 52 (1908) p. 381" “Ex-typis” "M. Bezzi det., 1908: // Dolichomerus // griseifascies Bezzi" "Moliro // Mars-Mai 95 //J. Duvivier" “♂” ”LECTOTYPUS” [KBIN]. Paralectotype, male, "Cf. Ann. Soc. Ent. Belg // Vol. 52 (1908) p. 381" “Ex-typis” "M. Bezzi det., 1908: // Dolichomerus // griseifascies Bezzi" "Moliro // Mars-Mai 95 //J. Duvivier" “♂” "PARA- // LECTOTYPUS" [KBIN]. Paralectotype, male, "Cf. Ann. Soc. Ent. Belg // Vol. 52 (1908) p. 381" “Ex-typis” "M. Bezzi det., 1908: // Dolichomerus // griseifascies Bezzi" "Moliro // Mars-Mai 95 //J. Duvivier" “♂” "PARA- // LECTOTYPUS" [KBIN].

Other material.

Angola • 2♂♂; 12 mi SW Luimbale; 20-21 Mar. 1972; NHMUK • 2♀♀; Luanda; Jun. 1957-Jul. 1958; G. Heinrich leg.; CNC • 1♀; Roçada; 30 Feb. 1972; NHMUK. Benin • 1♀; Cotonou; 3 Jun. 1914; W.A. Lamborn leg.; NHMUK • 7♂♂; Porto Novo; 7 Mar. 2018; K. Jordaens leg.; KMMA • 1♂ 1♀; Sémé; Nov. 2019; G. Goergen leg.; IITA. Botswana • 1♂; 20 km W Chonzi; 28 Aug. 1983; C. Stockmann leg.; NMSA • 1♀; Molepolole District; 3 Mar. 1963; T. Oatley leg.; NMSA • 1♀; Serowe; 14 Apr. 1983; P. Forchhammer leg.; NMSA. Burundi • 2♂♂; Bubanza, Gihanga Hill, Ruzizi; 23 Nov. 1951; F.J. François leg.; KBIN • 1♂; Bujumbura; 26 Feb. 1979; J. Decelle leg.; KMMA • 1♂ 1♀; Bujumbura; 21 Feb. 2017; G. Goergen leg.; IITA • 2♀♀; Kanyinya; Dec. 1947-Jan. 948; Dames de Marie leg.; KMMA • 1♂; Kitega; 3 Dec. 1950; F.J. François leg.; KBIN • 1♂; same collection data as for preceding; 1 May 1952 • 4♂♂ 3♀♀; same collection data as for preceding; 19 May 1955 • 1♀; same collection data as for preceding; 12 Jun. 1957 • 1♀; 11 Dec. 1962; M. Fontaine leg.; KMMA • 1♂; Murehe; 19 [?] 2013; S. Girukwishaka leg.; OBPE • 1♂; Rusizi N P; 26 [?] 2013; S. Girukwishaka leg.; OBPE • 1♀; Musuma, Bututsi; 29 Nov. 1950; F.J. François leg.; KBIN • 1♂ 9♀♀; Nyakibande, Mumirwa; 24 Apr. 2019; E. Sinzinkayo leg.; OBPE • 4♂♂ 9♀♀; same collection data as for preceding; 14 Jul. 2019 • 1♀; Nyambuye; Mumirwa; 18 Mar. 2019; E. Sinzinkayo leg.; OBPE • 1♂ 18; same collection data as for preceding; May 2019 • 6♀♀; same collection data as for preceding; 18 Sep. 2019 • 1♀; Rumonge; 20 Apr. 1948; • 1♀; same collection data as for preceding; 19-20 Jun. 1948 • 1♂; same collection data as for preceding; Jul. 1948 • 1♀; same collection data as for preceding; May 1949 • 1♀; 19 Feb. 1950; all F.J. François leg.; KBIN • 5♂♂ 20♀♀; Rwegura, Kibira NP; 31 May 2018; E. Sinzinkayo leg.; OBPE • 1♀; Rusinga NP, secteur Delta; 18 May 2018; E. Sinzinkayo leg.; OBPE • 2♂♂ 10♀♀; same collection data as for preceding; 19 May 2018 • 1♂; same collection data as for preceding; 20 May 2018 • 1♂ 4♀♀; same collection data as for preceding; 24 May 2018 • 1♀; Usumbura; Sep. 1961; G. Pierrard leg.; KMMA. Comoros • 1♂ 1♀; Le Galawa, Grande Comore; 22 Apr.-5 May 1991; K.M. Guichard leg.; NHMUK • 1♀; M’Vouni, Grande Comore; 23 Apr.-3 May 1991; K.M. Guichard leg.; NHMUK • 1♂ 1♀; Moroni, Grande Comore; Malet leg.; MNHN • 1♀; Grande Comore; 1899; MNHN. Democratic Republic Of The Congo • 3♀♀; Albertville [= Kalemie]; 14 Aug. 1953; J. Verbeke leg.; KBIN • 1♀; same collection data as for preceding; 17 Aug. 1953 • 1♀; same collection data as for preceding; 24 Aug. 1953 • 1♀; same collection data as for preceding; Jul. 1955; H.E. Bomans leg.; KBIN • 1♂; same collection data as for preceding; Dec. 1918; R. Mayné leg.; KMMA • 1♂; same collection data as for preceding; 1-20 Jan. 1919 • 1♂; same collection data as for preceding; 7 Feb. 1933; L. Burgeon leg.; KMMA • 1♀; Bendera; Oct. 1958; N. Leleup leg.; KMMA • 1♀; Baudouinville [= Kirungu]; May 1953; H. Bomans leg.; KMMA • 1♂; Bukama; 18 Apr. 1911; Bequaert leg.; KMMA • 1♂; same collection data as for preceding; 4 Jun. 1911 • 1♀; 3 Aug. 1923; Ch. Seydel leg.; KMMA • 1♀; Buta; 1949; R.F. Hutsebaut leg.; KMMA • 3♂♂; Chamunyonge; Lake Edward; 26 Sep. 1953; F.J. François leg.; KBIN • 2♂♂ 4♀♀; Costermansville [= Bukavu]; 1948; P.H. Vercammen leg.; KMMA • 1♂; Costermansville [= Bukavu]; 13 May 1949; H. Bomans leg.; KMMA • 1♀; Elisabethville [= Lubumbashi]; May 1964; E. Coussement leg.; KBIN • 2♂♂; Elisabethville [= Lubumbashi]; M. Bequaert leg.; KBIN • 1♂; Elisabethville [= Lubumbashi]; Nov. 1919; KMMA • 6♂♂; Elisabethville [= Lubumbashi]; 19 Sep. 1919; J. Ghesquière leg.; KMMA • 1♀; Elisabethville [= Lubumbashi]; 6 May 1920; M. Bequaert leg.; KMMA • 1♀; same collection data as for preceding; 3 Jun. 1920 • 2♂♂; same collection data as for preceding; 1 Jan. 1921 • 1♀; same collection data as for preceding; 5 Jan. 1921 • 1♂; same collection data as for preceding; 7 Jan. 1921 • 1♀; same collection data as for preceding; 16 Jan. 1921 • 1♂; same collection data as for preceding; 14 Feb. 1921 • 1♂; same collection data as for preceding; 1 Jul. 1923 • 1♂; same collection data as for preceding; 2 May 1923 • 1♀; same collection data as for preceding; Nov. 1929 • 1♀; same collection data as for preceding; Dec. 1939 • 1♀; same collection data as for preceding; Dec. 1928; Ch. Seydel leg. • 1♀; 1928-1929; same collection data as for preceding; P. Quarré leg. • 1♂; Elisabethville/Lubumbashi; 5 Jan. 1920; M. Bequaert leg.; NHMUK • 1♂; same collection data as for preceding; 23 Dec. 1920 • 1♀; same collection data as for preceding; 5 Jan. 1921 • 1♀; Goma; 1 May 1953; KBIN • 1♂; Goma; 18 Nov. 1952; J. Verbeke leg.; KBIN • 1♀; same collection data as for preceding; 10-15 May 1953 • 1♀; Ibembo; Uelé; Oct. 1949; R.F. Hutsebaut; KMMA • 1♀; same collection data as for preceding; 20 May 1950 • 1♀; same collection data as for preceding; Apr. 1952 • 1♀; same collection data as for preceding; 1953 • 1♀; Jadotville [= Likasi]; 18 Aug. 1965; E. Coussement leg.; KBIN • 1♀; Jampwe; 1912; Van den Heuvel leg.; KMMA • 1♀; Kabinda, Lomami; KBIN • 1♀; Mt Kabobo, Hte Kiymbi; Sep. 1958; N. Leleup leg.; KMMA • 1♀; Kakanda, Mutaka; 1955; R.P. Th de Caters leg.; KMMA • 1♀; Kamaniola; 1 Feb. 1927; J. Bequaert leg.; KMMA • 1♂; Kambove, Katanga; 11 Jun. 1907; S.A. Neave leg.; NHMUK • 1♀; Kasongo; Sep. 1959; P.L.G. Benoit leg.; KMMA • 4♀♀; Pons leg.; KMMA • 1♀; Kiambi; 27 Apr. 1931; G.F. de Witte leg.; KMMA • 1♂; Kigoma; Jul. 1918; R. Mayné leg.; KMMA • 1♀; same collection data as for preceding; Sep. 1918 • 1♂; Kindu; 15 Nov. 1911; Rodhain leg.; KMMA • 1♀; Kindu (300 km from) Russo; KMMA • 1♂; Kivu; Rodhain leg.; KMMA • 2♂♂ 1♀; Kolwezi; Lualaba; 1-6 Nov. 1952; L. Gilbert leg.; KMMA • 1♀; same collection data as for preceding; 13 Nov. 1952 • 1♀; same collection data as for preceding; 1 Feb. 1953 • 1♀; same collection data as for preceding; 5 Feb. 1953 • 1♀; Kongolo; 6 Feb. 1974; R. Baker leg.; NHMUK • 1♀; Kulu-Mwanza; Katanga; May 1927; A. Bayet leg.; KMMA • 1♀; Kunda; 23 Nov. 1910; Bequaert leg.; KMMA • 1♂; Lubongola; 1939; Hautmann leg.; KMMA • 1♀; same collection data as for preceding; May 1939 • 1♂; Lubumbashi; 5 Jan. 1921; M. Bequaert leg.; KMMA • 1♀; Lubumbashi; 2 Jan. 1968; A.B. Stam leg.; KMMA • 1♂; Lubutu; 22 Jan. 1915; J. Bequaert leg.; KMMA • 1♀; Luluabourg [= Kananga], Kabure; 1937; Soeurs du Carmel leg.; KMMA • 1♀; Luputa, Lomami; Sep. 1934; Bouvier leg.; KMMA • 1♀; same collection data as for preceding; Nov. 1934 • 1♀; same collection data as for preceding; 1935 • 1♀; Lusinga, P.N. Upemba; 10 Apr. 1947; G.F. de Witte leg.; KBIN • 1♀; same collection data as for preceding; 24 Apr. 1947; KMMA • 1♀; same collection data as for preceding; 26 Apr. 1947 • 1♂; Luvungi, North Kivu; 12 Dec. 1932; L. Burgeon leg.; KMMA • 4♂♂; Malondo, Katanga; Ch. Seydel leg.; KMMA • 1♂; Matale; Jul. 1939; Hautmann leg.; KMMA • 1♀; Moba, Tanganyika; Jun. 1931; G.F. de Witte leg.; KMMA • 1♀; Moba, Tanganyika; Jun. 1953; H. Bomans leg.; KMMA • 2♀♀; same collection data as for preceding; Aug.-Oct. 1953 • 2♀♀; Mpala, Tanganyika; H. Bomans leg.; KMMA • 1♀; Nyangwe; 15 Nov. 1910; Bequaert leg.; KMMA • 1♂ 1♀; Nyangwe; Apr.-May 1918; R. Mayné leg.; KMMA • 1♀; Nyonga, Katanga; 18 May 1925; G.F. de Witte leg.; KMMA • 1♀; Paulis; 29 Oct. 1959; M. Fontaine leg.; KMMA • 1♂ 1♀; Stanleyville [= Kisangani]; Apr. 1915; Lang and Chapin leg.; CNC • 1♀; same collection data as for preceding; Feb. 1915; KMMA • 17♂♂ 28♀♀; same collection data as for preceding; Mar. 1915 • 2♀♀; same collection data as for preceding; 1 Apr. 1915 • 1♂ 1♀; same collection data as for preceding; 2 Apr. 1915 • 1♀; same collection data as for preceding; 4 Apr. 1915 • 1♀; same collection data as for preceding; 5 Apr. 1915 • 2♂♂ 1♀; same collection data as for preceding; 6 Apr. 1915 • 2♂♂ 8♀♀; same collection data as for preceding; 7 Apr. 1915 • 4♀♀; same collection data as for preceding; 14 Apr. 1915 • 1♀; same collection data as for preceding; Apr. 1915 • 2♂♂ 3♀♀; same collection data as for preceding; 7 Jul. 1915 • 1♀; same collection data as for preceding; 1915; J. Bequaert leg.; KMMA • 1♀; same collection data as for preceding; 5 May 1926; H. Schouteden leg. • 1♀; same collection data as for preceding; 13 Aug. 1928; A. Collart leg. • 1♀; same collection data as for preceding; 6 Sep. 1928 • 1♂; same collection data as for preceding; 1929; J. Muller leg.; KBIN • 1♀; same collection data as for preceding; 28 Apr. 1932; J. Vrydagh leg.; KMMA • 1♀; Uvira; Jul. 1927; Ch. Seydel leg.; KMMA • 1♀; Uvira; Aug. 1949; G. Marlier leg.; KMMA • 1♀; Vieux-Kasongo; Dec. 1910; Pons leg.; KMMA. Ethiopia • 1♂; Abijata Shala National Park, Oromia; 1-10 Oct. 2012; A. Pauly leg.; KMMA • 1♂ 1♀; Senkele National Park, Oromia; 17 Oct. 2012; A. Pauly leg.; KMMA. Guinea Bissau • 2♀♀; Bolama; Jun.-Dec. 1899; L. Fea leg.; MCSNG. Kenya • 2♀♀; Bungoma, Western Province; 29 Oct. 1968; J. Field leg.; CDFA • 1♀; Funzi Island; 9 Jul. 2012; ICIPE • 1♂; Gede National Park, Malindi; 1 May 1973; H. Falke leg.; CNC • 1♀; Kibwezi; 6 Jan. 1926; W. Feather leg.; NHMUK • 7♀♀; same collection data as for preceding; 24 Dec. 1927 • 4♀♀; same collection data as for preceding; 24 Jan. 1928 • 1♀; same collection data as for preceding; 9 Feb. 1928 • 1♀; Kisumu; Nov. 1932; V.G.L. van Someren leg.; NHMUK • 2♀♀; Lake Mpeketoni near Kipini; 4-5 Mar. 1912; S.A. Neave leg.; NHMUK • 1♀; Loyia Turkana; 20 Aug. 1954; NHMUK • 1♀; Magadi; Jul. 1941; V.G.L. van Someren leg.; NHMUK • 1♀; Mombasa; Jul. 1904; Ch. Alluaud leg.; MNHN • 1♀; Nairobi; 14 Dec. 1951; L.C. Edwards leg.; NHMUK • 1♀; Nanyuki; Jul. 2010; M. Roberts leg.; ICIPE • 1♂; New Runda; 22 Apr. 2012; F. Haas leg.; ICIPE • 2♀♀; Rabai; May 1928; V.G.L. van Someren leg.; NHMUK • 2♂♂; same collection data as for preceding; Jul. 1928 • 1♂; Tsavo East; 6 Jan. 1989; A.u.G. Rautenstrauch leg.; CDFA • 1♀; Uchweni Forest; 1-3 Mar. 1912; S.A. Neave leg.; NHMUK • 1♂ 2♀♀; Ziwani; Aug. 1947; V.G.L. van Someren leg.; NHMUK. Malawi • 1♀; 30 km NW of Dedza; 15 Mar. 1987; J. and A. Londt leg.; NMSA • 3♀♀; Kasungu; 9 Dec. 1980; B. Stuckenberg and J. Londt leg.; NMSA • 1♂; Livingstonia; 3 May 1939; R.C. Wood leg.; NHMUK • 1♀; Mulanje Mountain Forest Reserve; 12-15 Nov. 2016; K. Jordaens leg.; KMMA • 1♂; Ruo; 4 Feb. 1916; R.C. Wood leg.; NHMUK • 1♂ 1♀; Viphya Mountains, Chikangawa; 5-8 Dec. 1980; B. Stuckenberg and J. Londt leg.; NMSA. Mozambique • 1♀; Chiramba, Zambeze; 1929; P. Lesne leg.; MNHN • 1♀; Lourenço Marques [= Maputo]; 1909; J. de O.S. de Azevedo leg.; NHMUK • 2♂♂ 2♀♀; Lourenço Marques [= Maputo]; Sep.-Dec. 1913; H.A. Junod leg.; NHMUK • 1♂; same collection data as for preceding; Jan.-Mar. 1914 • 1♀; Luabo, Lower Zambesi River; Jun.-Jul. 1957; CNC • 1♀; same collection data as for preceding; Aug. 1957 • 2♀♀; Luabo; U. Stuckenberg leg.; NMSA • 2♂♂; Manga, Beira Region; 27 Jul. 1929; P. Lesne leg.; MNHN • 1♂; Revoué Valley, Andrada envir.; Apr.-May 1905; G. Vasse leg.; MNHN • 1♀; Rikatla; H.A. Junod; NMSA • 1♀; Siluwe Hills, W. of Beira; 3 Jun. 1964; D. Cookson leg.; NMSA • 1♀; Tumbine Mountains, Milange District; Apr. 1958; B. and P. Stuckenberg leg.; NMSA. Namibia • 3♀♀; Kombat; 1-6 Apr. 1972; NHMUK • 1♀; between Omaruru and Wilhelmstad; 3 Apr. 1998; F.W. and S.K. Gess leg.; AMGS • 1♀; WNW of Omatjete; 15 Mar. 2004; F.W. and S.K. Gess leg.; AMGS • 1♂; Oranjemund, 28 km from checkpoint on road to Sendelingsdrift; 25 Sep. 1997; F.W. and S.K. Gess leg.; AMGS • 1♀; Rietfontein, 23 mi W SW Grootfontein; 3 Apr. 1972; NHMUK • 1♀; 24 km SE Stampriet, Gross Nabas; 30 Mar. 2000; F.W. and S.K. Gess leg.; AMGS • 1♀; 71 km East Stampriet; 23 Mar. 2000; F.W. and S.K. Gess leg.; AMGS • 1♂; 71 km West Witvlei; 17 Mar. 1984; J. Londt and B. Stuckenberg leg.; NMSA. Nigeria • 1♀; 23 km W Lagos; 11 May 1975; J. Riley leg.; NHMUK. Republic Of The Congo • 1♀; Brazzaville env.; 1907; E. Roubaud and A. Weiss leg.; MNHN. Rwanda • 1♀; Astrida [= Butare]; 22 Feb. 1953; P. Basilewsky leg.; KMMA • 1♀; same collection data as for preceding; 5 Mar. 1955; P. Elsen leg.; • 1♀; same collection data as for preceding; 15 Mar. 1955 • 1♀; Karissimbi; Dec. 1925; H. Schouteden leg.; KMMA. São Tomé and Príncipe • 3♂♂ 2♀♀; São Tomé, Ribeira Palma; Aug. 1900; L. Fea leg.; MCSNG • 3♀♀; Príncipe, Roca Inf. D. Henrique; Mar. 1901; L. Fea leg.; MCSNG • 1♀; 1900; A. Negreiros leg.; MNHN • 1♀; 1919-1921; H.J. Snell leg.; NHMUK • 2♂♂ 1♀; 1920; H. Navel leg.; MNHN • 1♀; 19 Sep. 1921; H.J. Snell leg.; NHMUK • 1♀; no data; H. De Saeger and Prince Leopold leg.; KMMA. Somalia • 1♀; Mogadishu; 20 Feb. 1953; NHMUK. South Africa • 1♀; Amatikulu Nature Reserve; 14 Jun. 2011; J. Londt and T. Dikow leg.; NMSA • 4♀♀; Ben Lavin Nature Reserve; 13 Feb. 2005; J. Londt and T. Dikow leg.; NMSA • 1♂; Bluff, Durban; 8 Mar. 1925; NMSA • 1♀; Blyderivierspoortdam Nature Reserve; 25-26 Oct. 1984; C.D. Eardley leg.; NMSA • 1♀; Empangeni; 7 Apr. 1989; P. Reavell leg.; NMSA • 1♀; False Bay; 24 Feb.-4 Mar. 1990; A.J. Weaving leg.; NMSA • 1♀; Gillits; 24 Feb. 1957; NMSA • 1♀; Ingwavuma; 10 Dec. 1963; B. and P. Stuckenberg leg.; NMSA • 1♀; Irene; 24 May 1971; C.K. Brain leg.; NMSA • 1♂; Letaba Camp, Kruger National Park; 14-18 Nov. 1961; Vari and Rorke leg.; NMSA • 2♀♀; 37 km N Louis Trichardt, Limpopo; Jan. 1975; B. Stuckenberg leg.; NMSA • 2♂♂ 4♀♀; Mafikeng Game Reserve, Kolobe drinking pond; 16 Mar. 2003; J.Londt leg.; NMSA • 1♀; Mafikeng Game Reserve, Noka picnic site; 17 Mar. 2003; J. Londt leg.; NMSA • 1♀; Malvern; Natal; Jun. 1897; G.A.K. Marshall leg.; NHMUK • 1♀; Mntunzini, Garland Farm, Natal; 1 Aug. 1985; P. Atkinson leg.; NMSA • 1♀; Modimolle, Limpopo; 20 Nov. 2017; A. Vujic and E. Rättel leg.; MZH • 1♂ 2♀♀; Mpenjati Nature Reserve; 22-23 Jan. 2011; J. and A. Londt leg.; NMSA • 1♀; Ndumu Game Reserve, staff house; 15-17 Feb. 2011; J. Londt leg.; NMSA • 1♀; Ndumu Game Reserve, rest camp; 23-29 Nov. 1977; D. Brothers and J. Guillarmod leg.; NMSA • 1♀; Port St John Pondoland; 1-15 Apr. 1924; B.E. Turner leg.; NHMUK • 1♀; 5 km E Sabie, Bergvliet Road; 24 Feb. 1971; B. Stuckenberg leg.; NMSA • 1♂ 1♀; San Lameer resort area; 11-12 Feb. 2012; J. and A. Londt leg.; NMSA • 1♀; Shelly Beach; 29 May 1984; D. Wheeler leg.; NMSA • 1♀; 4 km E Skukuza, Sabie River, Kruger National Park; 4 Jan. 1974; B. and P. Stuckenberg leg.; NMSA • 1♀; Tshifhire, Venda; Jul. 1983; Neluheni leg.; NMSA • 1♀; Umbilo; 21 Mar. 1915; L. Bevis leg.; NMSA • 2♂♂ 1♀; Umtamvuna Nature Reserve; 14 Jan. 1981; J. Londt leg.; NMSA • 1♂; same collection data as for preceding; 29 Oct. 1990; A. Whittington leg. • 1♀; van Reenen’s Pass, Ladysmith, Kwa-Zulu Natal; 23 Feb. 2016; G. Ståhls and E. Rättel leg.; MZH • 1♀; Vernon Crookes Nature Reserve; 16 Nov. 2003; G.B.P. Davies leg.; NMSA. Tanzania • 1♀; Amani; 1957; J.G. Halcrow leg.; NHMUK • 1♀; Chenzema, Uluguru Mountains; 2-22 Jul. 1971; L. Berger; N. Leleup and J. Debecker leg.; KMMA • 1♂; Dar es Salaam; Apr. 1961; G. Heinrich leg.; CNC • 1♀; same collection data as for preceding; 12 Sep. 1961 • 1♀; same collection data as for preceding; 14 Sep. 1961 • 11♂♂ 1♀; same collection data as for preceding; Oct. 1961 • 1♂; same collection data as for preceding; 20 Oct. 1961 • 2♂♂; Dodoma; Nov. 2008; G. Goergen leg.; IITA • 2♂♂; same collection data as for preceding; 27 Nov. 2008 • 8♂♂; Iringa-Morogoro; 23 Nov. 2008; G. Goergen leg.; IITA • 1♀; Kunduchi; Jul. 1973; CDFA • 1♀; Lake Manyara; Jun.-Aug. 1937; B. Cooper leg.; NHMUK • 1♀; Matumbi Highlands; 25 Nov. 1989; W.R.B. Hynd leg.; NHMUK • 1♀; Morogoro; 6 May 1925; A.H. Ritchie leg.; NHMUK • 1♂ 1♀; same collection data as for preceding; 20 May 1925 • 2♀♀; same collection data as for preceding; 10 Jun. 1925 • 1♀; nr Morogoro, Uluguru Mountains; Jan. 1962; CNC • 1♂; Mungula Gorge, Bunduki, Uluguru Mts; P. Basilewsky and N. Leleup leg.; KMMA • 1♂; Musoma; 29 Nov. 2008; G. Goergen leg.; IITA • 2♂♂; Mwanza; 30 Nov. 2008; G. Goergen leg.; IITA • 1♂; Njombe; 23 Oct. 1951; W. Peters leg.; NHMUK • 1♀; same collection data as for preceding; 9 Dec. 1952 • 1♂; same collection data as for preceding; 23 Oct. 1957 • 1♂; Old Shinyanga; 26 Mar. 1953; E. Burtt leg.; NHMUK • 1♂; same collection data as for preceding; 28 Mar. 1953 • 1♂; same collection data as for preceding; 10 Apr. 1953 • 1♂; same collection data as for preceding; 11 Apr. 1953 • 1♂; Rukwa Valley; 16 Sep. 1952; NHMUK • 2♀♀; Tanga; Apr.-May 1950; R.C.H. Sweeney leg.; NHMUK • 3♂♂; Ukara Island; Lake Victoria; 1953; A. Smith leg.; NHMUK • 1♀; Zanzibar; [no date]; ex coll Bigot; NHMUK • 2♀♀; Zanzibar; C. Cooke leg.; CNC • 1♂; Zanzibar; 17 May 1911; W.M. Aders leg.; NHMUK • 1♂; Zanzibar; Jan.-Feb. 1925; H.J. Snell leg.; NHMUK. Togo • 1♂; Kloto; 3 Mar. 2018; K. Jordaens leg.; KMMA. Uganda • 1♂ 1♀; Budongo Forest, nr Lake Albert; 1 Apr. 1972; E.B. Babyetagara leg.; CNC • 1♂ 1♀; Entebbe; May 1906; M. de Rothschild leg.; MNHN • 1♀; Entebbe; 20 Mar. 1911; C.C. Gowdey leg.; NHMUK • 1♀; same collection data as for preceding; 17 Aug. 1911 • 2♂♂ 10♀♀; same collection data as for preceding; 21 Aug. 1911 • 1♀; same collection data as for preceding; 1-11 Sep. 1911 • 2♂♂ 1♀; same collection data as for preceding; 7-9 May 1912 • 1♀; same collection data as for preceding; 27 May 1912 • 5♂♂ 3♀♀; same collection data as for preceding; 3 Sep. 1912 • 2♀♀; same collection data as for preceding; 16 Oct. 1912 • 1♀; same collection data as for preceding; 14 Nov. 1912 • 1♂; same collection data as for preceding; 18-20 Nov. 1912 • 2♀♀; same collection data as for preceding; 3-4 Dec. 1912 • 1♂ 1♀; same collection data as for preceding; 12-13 Dec. 1912 • 1♀; same collection data as for preceding; 31 Dec. 1912 • 1♀; same collection data as for preceding; 20 Mar. 1913 • 5♀♀; same collection data as for preceding; 1913 • 1♀; same collection data as for preceding; 6-13 May 1912; C.A. Wiggins leg. • 1♀; Entebbe; 1 Feb. 1972; H. Falke leg.; CNC • 2♂♂; same collection data as for preceding; 20 Nov. 1972-22 Jan. 1973 • 2♂♂; same collection data as for preceding; 25-28 Jan. 1973 • 1♀; 7 mi N Entebbe; 31 Mar. 1972; H. Falke leg.; CNC • 3♂♂ 3♀♀; near Entebbe; 23-31 Jan. 1973; H. Falke leg.; CNC • 1♂; Kayonza Forst, Kigezi; 1 Sep. 1972; H. Falke leg.; CNC • 1♀; Masindi; Aug. 1945; P.A. Buxton leg.; NHMUK • 1♀; Ruwenzori; 17 May 1911; C.C. Gowdey leg.; NHMUK • 1♀; Sukulu; 26 Jan. 1954; NHMUK • 1♀; Sukulu; 9 Apr. 1961; E. Burtt leg.; NHMUK • 2♂♂ 1♀; same collection data as for preceding; 26 Jan. 1962 • 1♀; same collection data as for preceding; 31 Jan. 1962 • 1♂; Tero; 14 Apr. 1911; C.C. Gowdey leg.; NHMUK • 1♂; Tororo; 30 Jan. 1954; NHMUK. Zambia • 2♂♂; Abercorn; 17 Jan. 1951; F.O. Albrecht leg.; NHMUK • 1♀; same collection data as for preceding; 12 Feb. 1951 • 1♂; same collection data as for preceding; 26 Feb. 1951 • 1♀; Jan. 1966; H. Mathes leg.; KMMA • 1♀; Broken Hill; 1 May 1912; E.A. Copeman leg.; NHMUK • 1♀; same collection data as for preceding; 29 May 1912 • 1♀; Mwinilunga; 9 Apr. 1914; F.V. Bruce Miller leg.; NHMUK • 1♀; Ndola; 31 Jan. 1980; T. Grout leg.; NMSA • 1♀; Nyamkolo; 9 May 1930; A. Loveridge leg.; CNC. Zimbabwe • 1♀; Banket; Dec. 1950; A. Carnegie; NMSA • 1♀; Bazely Bridge, Mutare; 14 Apr. 1964; D. Cookson leg.; NMSA • 1♀; Bomponi; 13 May 1963; D. Cookson leg.; NMSA • 1♂; same collection data as for preceding; 16 Jun. 1965 • 1♀; Chigudu Farm, Mvurwi; 14 Jan. 1985; A.J. Weaving leg.; NMSA • 1♂; Melsetter District; 23 Nov. 1993; E. Bruce-Miller leg.; AMGS • 1♂; Salisbury [= Harare]; 1 Jan. 1952; J.M. Brown leg.; NMSA • 1♀; same collection data as for preceding; 22 Apr. 1978; P.E. Hulley leg. • 1♀; Vumba Mt; March 1938; NHMUK • 1♀; East Vumba; 1 Jun. 1964 • 1♂; North Vumba; 27 Mar. 1964; D. Cookson leg.; NMSA • 1♀; same collection data as for preceding; 16 Nov. 1964.

Description.

Body length: 11.0-15.8 mm. Wing length: 8.0-11.0 mm.

Male (Fig. 5 ). Head (Figs 28 , 29 ). Eye bare; holoptic, eye contiguity for distance equal to or slightly longer than length of ocellar triangle, facets dorsally slightly larger, at most twice as large in diameter, as ventral ones. Frons black to black-brown; weakly subshiny, with dense greyish pollinosity throughout; with dispersed medium long straw-yellow pile. Face black to black-brown; weakly subshiny, with dense greyish pollinosity, facial tubercle weakly shining black, posterolateral margin of mouth shining brownish; in parts with dispersed long pale pile; facial tubercle strongly pronounced. Gena colour and pollinosity as ventral lateral margin of face; with short to long pale pile. Occiput black-brown, covered with dully grey pollinosity; with dispersed pale pile, except dorsally where pile black adjacent to vertex. Antennal segments black-brown, basoflagellomere apically more brownish, arista pale yellow.

Thorax (Fig. 57 ). Scutum mainly subshiny black; sparse grey pollinose, more densely so along margins and transverse suture; pilosity variable, usually predominantly pale yellow, sometimes more or less interspersed with short black pile. Scutellum apical margin weakly rounded, distinctly marginated, slightly more than twice as wide as long; pale yellow, anterior margin narrowly black-brown; with short pale pile. Pleura ground colour black-brown, sparsely greyish pollinose; covered with dispersed long pale pile except on meron, dorsomedial anepimeron, ventral part of katepimeron, anterior part of katepisternum and anterior anepisternum.

Legs. Dark brown to reddish brown; with short dark pile, along posterior margin of pro- and mesofemora with longer pale pile except apically, metafemur predominantly long pale pile in basal two-thirds. Metaleg (Fig. 85 ), femur moderately thickened, with one distinct ventral swelling in apical fifth, a second less developed thickening proximal of this; tibia thickened and slightly curved, pile along ventral margin more dense.

Wing (Fig. 73 ). Largely hyaline, only with very faint yellowish brown tinge; with distinct dark brown macula running from anterior margin and covering most of stigma, parts of cell r1 and r2+3, distal part of cell br and basal part of r4+5, decreasing in intensity on anterior part of cell dm, macula well demarcated apically but decreasing gradually in colour basally. Calypters white; with fringe of white pile. Cell r1 closed, petiole usually approx. equal to height of base of stigma, rarely shorter. Vein R4+5 sinuate and short appendiculate, rarely appendix missing.

Abdomen (Fig. 95 ). Uniformly subshiny brown to reddish brown; tergum IV, postabdomen and sometimes posterior margin of tergum III conspicuously orange to orange-red; with short dark pile, except tergum I completely, anterior half of tergum II and anterior fourth of tergum III with longer silvery to pale yellow pile; lateral margins with pale pile; orange areas with pale orange pile. Male genitalia as in Fig. 106 .

Female. As male except for the following character states: Eye dichoptic (Figs 30 , 31 ), facets equal to subequal in size all over. Frons subshiny black to black-brown, in medial part with greyish pollinosity, pilosity shorter and more silvery pale except in front of ocellar triangle where short and black. Thorax, pilosity usually more extensively black, usually as fasciae. Abdomen pale pilosity along lateral margins missing; pale pilosity in tergum II at most in anterior third, tergum III predominantly with black pile. In general, thoracic and abdominal pale pilosity more silvery than in male.

Distribution.

Angola, Benin, Botswana, Burundi, Comoros, Democratic Republic of the Congo, Ethiopia, Guinea Bissau, Kenya, Malawi, Mozambique, Namibia, Nigeria, Republic of the Congo, Rwanda, São Tomé and Príncipe, Somalia, South Africa, Tanzania, Togo, Uganda, Zambia and Zimbabwe.

Comments.

The type material of S. haemorrhoa (both male and female specimens according to the original description) could not be traced.

Senaspis griseifacies was proposed as a species by Bezzi (1908), who provided a short description. Bezzi (1912) included Senaspis griseifacies and Senaspis haemorrhoa Gerstaecker in his key for the genus Protylocera (page 416) and provided a more detailed description of S. griseifacies . He differentiated the latter from S. haemorrhoa mainly on characters of the wing (anterior margin basally more hyaline in S. griseifacies , infuscated in S. haemorrhoa ) and the abdomen (terga II and III along posterior margin yellow in S. haemorrhoa , dark in S. griseifacies ). Examination of long series of S. haemorrhoa and of type material of S. griseifacies has shown these characters to be variable. In addition, Bezzi (1912) contradicted the abdominal character partially in his detailed redescription where he indicated that in female specimens of S. griseifacies the “è spesso rosso anche l’orlo posterior del terzo segment" (partially red along posterior margin of the third segment). Both species are considered conspecific and thus S. griseifacies is considered junior synonym of S. haemorrhoa . Hervé-Bazin (1914) already alluded to this when indicating that S. griseifacies should be considered perhaps as a mere variety of S. haemorrhoa . An identification label ' Senaspis haemorrhoa ' of 1996 by C. Kassebeer on one of the types of S. griseifacies at the KBIN collection also alludes to the synonymy but apparently was never formally published.

Figures

Figures 5, 6. Senaspis species, habitus, lateral view 5 S. haemorrhoa (Gerstaecker) (♂) 6 S. melanthysana (Speiser) (♂).

Figures 28 - 31. Head, frontal and lateral view S. haemorrhoa (Gerstaecker) (28, 29 ♂ 30, 31 ♀).

Figures 56 - 59. Senaspis species, thorax, dorsal view 56 S. flaviceps Macquart (♂) 57 S. haemorrhoa (Gerstaecker) (♂) 58 S. melanthysana (Speiser) (♂) 59 S. nigrita (Bigot) (♂).

Figures 68 - 73. Senaspis species, right wing 68 S. elliotii Austen (♂) 69 - 71 S. elliotii Austen close-up apical end 72 S. flaviceps Macquart (♂) 73 S. haemorrhoa (Gerstaecker) (♂).

Figures 80 - 85. Senaspis species, hind leg, anterior view 80 S. dentipes (Macquart) (♂) 81 S. dibapha (Walker) (♂) 82 S. elliotii Austen (♂) 83 S. flaviceps Macquart (♂) 84 S. flaviceps Macquart (♀) 85 S. haemorrhoa (Gerstaecker) (♂).

Figures 91 - 96. Senaspis species, abdomen, dorsal view 91 S. dentipes (Macquart) (♂) 92 S. dibapha (Walker) (♂) 93 S. elliotii Austen (♂) 94 S. flaviceps Macquart (♂) 95 S. haemorrhoa (Gerstaecker) (♂) 96 S. melanthysana (Speiser) (♂).

Figures 102 - 113. Senaspis species, male terminalia 102 - 108, 110, 112 dorsal view 109, 111, 113 lateral view 102 S. dentipes (Macquart) 103 S. dibapha (Walker) 104 S. elliotii Austen 105 S. flaviceps Macquart 106 S. haemorrhoa (Gerstaecker) 107 S. nigrita (Bigot) 108, 109 S. melanthysana (Speiser) 110, 111 S. umbrifera (Walker) (holotype) 112, 113 S. nr umbrifera (Walker).

Figures 97 - 101. Senaspis species, abdomen, dorsal view 97 S. nigrita (Bigot) (♂) 98 S. umbrifera (Walker) (♂ holotype) 99 S. nr umbrifera (Walker) (♂) 100 S. xanthorrhoea (Bezzi) (♀) 101 S. pennata (Herve-Bazin) (♀).