Tizatetrichia panamensis Harris and Armitage, 2019

Harris, Steven C. & Armitage, Brian J., 2019, The Trichoptera of Panama X. The Quebrada Rambala drainage, with description of 19 new species of microcaddisflies (Trichoptera: Hydroptilidae), Insecta Mundi 707 (707), pp. 1-54 : 18

Tizatetrichia panamensis Harris and Armitage , new species

Fig. 23

Diagnosis. This new species is difficult to place, as it lacks one of the characters used by Harris et al. 2002, to designate the genus, that of the venter of segment X being well-developed and bearing heavy spines. However, the inferior appendages are longer than segment X and thin, and the subgenital plate is reduced, which is characteristic of the genus. This is the second species to be placed in Tizatetrichia , and it is possible that these are aberrant members of the closely related genus Bredinia . However, both T. costaricensis and the new species lack the elongate thickened seta from the posterior of the inferior appendage which is typical for Bredinia . The structure of the phallus in the new species is also more similar to that seen in T. costaricensis , being deeply divided posteriorly, and lacking a medial, protruding ejaculatory duct which is typical of Bredinia . We have collected a third species belonging to this genus which strengthens our argument, but females for Tizatetrichia remain unknown and could provide additional information when discovered.

Male. Length 1.3–1.5 mm. Head, wings and 18 segmented antennae typical for genus, as are the legs and thoracic segments, brown in alcohol. Abdominal segments VII annular with pair of posterolateral apodemes. Segment VIII annular, incised ventrally on posterior margin. Segment IX rectanguloid in lateral view, rounded ventrally and tapering dorsally, anteriorly narrowing to elongate apodemes which extend into segment VI; ventrally with mesal notch; dorsally deeply incised on posterior margin, laterally with sclerotized margins. Segment X reduced laterally to thin lobe, with dorsal notch posteriorly; dorsally membranous with posterior incision. Inferior appendages in lateral view narrow, tapering distally and bearing stout spike; in ventral view, thin and elongate, rectanguloid posteriorly, fused basally and bearing numerous short spines, apically with short, stout spike Subgential plate not noticeable and seemingly represented by a flat plate which is incised posteriorly. Phallus tubular, in dorsal view divided apically into pair of long, narrow processes each tipped with numerous spines, long lateral spine subapically, mesally with cluster of short spikes and lateral processes; in lateral view ending in several long, spiny processes, mesally with narrow sclerite just posterior to cluster of sharp spikes.

Female. Unknown.

Type material. Holotype, male— Panama, Bocas del Toro Province, Quebrada Rambala, near Rambala Jungle Lodge , 3.74 km SSE Rambala, 8.91543°N and 82.15527°W, 120 m, Malaise trap, February 6–12, 2017, E. Carlson . Paratypes. ibid., November 15–20, 2016, 1 male ( SCH).

Etymology. Named for the country of Panama, whence the species is described.

Figures

Figure 23. Tizatetrichia panamensis, sp. n., Male genitalia. A) Lateral view. B) Ventral view. C) Dorsal view. D) Phallus, lateral view. E) Phallus, dorsal view.