Colossobolus, Wesener & Enghoff & Sierwald, 2009

Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra, 2009, Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda), ZooKeys 19 (19), pp. 1-128 : 23-24

publication ID	https://doi.org/ 10.3897/zookeys.19.221
publication LSID	lsid:zoobank.org:pub:C473F9F6-1AE7-4B3F-B17F-CA1C2709010C
DOI	https://doi.org/10.5281/zenodo.3791382
persistent identifier	https://treatment.plazi.org/id/1772122E-7811-FFD1-FF01-3A44AB0DEFAB
treatment provided by	Plazi
scientific name	Colossobolus
status

Treatment

Key to the Colossobolus View in CoL species:

1 Telopodite of posterior gonopod basally with a finger-shaped coxite branch ( Figs 9C View Figure 9 , 14A View Figure 14 , 15F View Figure 15 )................................................................................... 2

– Telopodite of posterior gonopod lacking coxite branch ( Figs 16F View Figure 16 , 19A View Figure 19 , 20G, 21E View Figure 21 ) .......................................................................................................... 4

2 Lateral branch of posterior gonopod telopodite curved towards coxite branch, almost forming a circle with latter ( Figs 9H, I View Figure 9 ). Coxite of anterior gonopod in anterior view with a short, wide, well-rounded process ( Fig. 9D View Figure 9 )............. ...................................................................................... C. semicyclus View in CoL sp. n.

– Lateral branch of posterior gonopod telopodite straight ( Fig. 14A View Figure 14 ). Coxite of anterior gonopod in anterior view with a short pointed process ( Fig. 13C View Figure 13 )........... 3

3 Tarsi whitish ( Fig. 13B View Figure 13 ). Collum completely red ( Fig. 13A View Figure 13 ). Coxite branch of posterior gonopod short, half as long as lateral branch. Lateral branch irregularly curved, becoming apically thicker ( Figs 14A, B View Figure 14 ). Retrorse process absent from mesal margin of telopodite process of anterior gonopod ( Fig. 13D View Figure 13 )..... ................................................................................ C. oblongopedus View in CoL sp. n.

– Legs completely red ( Fig. 15B View Figure 15 ). Collum at margins red, median part dark ( Fig. 15A View Figure 15 ). Coxite branch of posterior gonopod long, as long as lateral branch. Lateral branch regularly curved ( Fig. 15F View Figure 15 ). Mesal margin of telopodite process of anterior gonopod protruding into a very short, but pointed retrorse process ( Fig. 15D View Figure 15 ) .............................................. C. giganteus View in CoL sp. n.

4 Telopodites of posterior gonopods oriented face-to-face to one another. Main branch of telopodite overlapping juxtaposed counterpart ( Figs 16E, F View Figure 16 ). Small, 85–100 mm long. Colour of body rings dark brownish ( Fig. 16B View Figure 16 ).... ............................................................................................ C. minor View in CoL sp. n.

– Telopodites of posterior gonopods oriented parallel to one another. Telopodite main branches not overlapping ( Figs 19A, B View Figure 19 ). Body large,> 120 mm. Colour of body rings always partly red....................................................... 5

5 Mesal margin of telopodite process of anterior gonopod widely projecting, almost protruding up to lateral margin ( Fig. 18F View Figure 18 ). Coxite process extraordinary long, but wide and well-rounded, longer than sternite, but far shorter than process of telopodite ( Fig. 18E View Figure 18 ). Telopodite of posterior gonopod not U-shaped. Lateral branch weakly developed, very slender ( Fig. 19A View Figure 19 ). Telopodite main branch separated into two parts, lateral part with large and long membranes, mesal part short and sclerotized. Telopodite lateral branch and both parts of main branch orientated parallel to one another ( Figs 19A, B View Figure 19 ) ...................... C. litoralis View in CoL sp. n.

– Mesal margin of telopodite process of anterior gonopod sharp, but not projecting ( Fig. 20F View Figure 20 ). Coxite process present, but short and well-rounded ( Fig. 20A View Figure 20 ). Telopodite of posterior gonopod U-shaped ( Fig. 20E View Figure 20 ). Lateral and main branches apically of similar width. Telopodite main branch not divided into two parts, branch basally sclerotized, apically membranous ( Figs 20E, G View Figure 20 )........................ 6

6 Preanal process apically sharp-edged, slightly shorter than anal valves ( Fig. 20C View Figure 20 ). Main branch of posterior gonopod telopodite of equal width all along ( Fig. 20E View Figure 20 ), longer than lateral branch ( Fig. 20G View Figure 20 ) ............ C. aculeatus View in CoL sp. n.

– Preanal process apically rounded, shorter than anal valves ( Fig. 21B View Figure 21 ). Main branch of posterior gonopod telopodite basally very slender, but with a wide, swollen tip ( Fig. 21E View Figure 21 ), shorter than lateral branch ( Fig. 21F View Figure 21 ) ....................... ............................................................................ C. pseudoaculeatus View in CoL sp. n.