Grishin, Zhang & Cong & Shen & Grishin, 2022

Zhang, Jing, Cong, Qian, Shen, Jinhui & Grishin, Nick V., 2022, Taxonomic changes suggested by the genomic analysis of Hesperiidae (Lepidoptera), Insecta Mundi 2022 (921), pp. 1-135 : 48-49

Tixe Grishin , new subgenus

http://zoobank.org/ 6A1F5AB0-013D-460A-9E0C-2A740784317F

Type species. Cobalus quadrata Herrich-Schäffer, 1869 .

Definition. Our genomic tree reveals that a number of species currently placed in Tisias Godman, 1901 (type species Proteides myna Mabille, 1889 ) are not monophyletic with it and instead form a clade sister to Xeniades Godman, 1900 (type species Papilio orchamus Cramer, 1777 ), which also includes Cravera de Jong, 1983 (type species Cravera rara de Jong, 1983 ) as a subgenus ( Fig. 11 ). This non- Tisias clade is not prominent enough to be a genus of its own, but together with Xeniades sensu stricto and Cravera , they form a prominent clade that we define as the genus Xeniades , with the non- Tisias clade as its new subgenus. It keys to K.20.1 or K. 20.2 in Evans (1955). Distinguished from its relatives by the following combination of characters: forewing discal cell spots staggered in males, upper spot is not connected to the lower spot (connected or nearly so in Tisias ); forewing with brands; body robust, palpi quadrantic, third segment short and stout, antennae half of costa in length, apiculus long, half of the club, mid-tibiae with spines; gnathos arms broadly separated and visible in dorsal view protruding on the sides of uncus. In DNA, a combination of the following base pairs is diagnostic: aly1146.55.6:C75T, aly1139.48.3:G142A, aly1432.13.2:A93G, aly144.18.1:A185G, and aly536.195.3:A187C.

Etymology. The name is a feminine noun in the nominative singular, a fusion of previous and presently suggested genus names for these species: Ti [sias] + xe [niades].

Species included. The type species, Tisias rinda Evans, 1955 and Tisias putumayo Constantino and Salazar, 2013 .

Parent taxon. Genus Xeniades Godman, 1900 .

Figures

Figure 11. Genomic tree of Hesperiina and relatives. See Fig. 1 legend for notations.