Grishin, Zhang & Cong & Shen & Grishin, 2022
publication ID |
https://doi.org/ 10.5281/zenodo.6392056 |
persistent identifier |
https://treatment.plazi.org/id/183DE44C-FFC0-FFB6-AFF9-FECBFD5EC2E0 |
treatment provided by |
Felipe |
scientific name |
Grishin |
status |
subgen. nov. |
Nausia Grishin , new subgenus
http://zoobank.org/ 2DF8540E-15F3-41E2-8F17-C537DB2F20E4
Type species. Oenus [sic] nausiphanes Schaus, 1913 .
Definition. Currently in Tigasis Godman, 1900 View in CoL (type species Tigasis zalates Godman, 1900 ) but not monophyletic with it, and instead is sister to the clade consisting of subgenera Mnasicles Godman, 1901 View in CoL (type species Mnasicles geta Godman, 1901 View in CoL ) and Remella Hemming, 1939 View in CoL (type species Hesperia remus Fabricius, 1798 View in CoL ) with subgenus Styriodes Schaus, 1913 View in CoL (type species Styriodes lyco Schaus, 1913 View in CoL ) originating right before, therefore is a subgenus of its own ( Fig. 13 View Figure 13 ). Keys to J. 45.11 in Evans (1955). Similar to subgenus Remella in ventral hindwing pattern with a broad diffuse central cream band, but differs in that both sides of the band blend gradually with the basal or marginal brown areas (basal area sharply outlined in Remella ); antennae checkered; stigma more prominent than in relatives, wider, less curved than in Remella , and lined with areas of gray scales outwards; forewing costal margin straight, not concave as in subgenus Styriodes Schaus, 1913 View in CoL (type species Styriodes lyco Schaus, 1913 View in CoL ) hindwing tornus in males more extended than in other subgenera; valva broad, only a third longer than its height, costa slightly convex, harpe rounded terminally, unturned, appressed to rounded ampulla, separated from it by a small notch, aedeagus terminally with a keel on both sides. In DNA, a combination of the following base pairs is diagnostic: aly216.57.2:G803A, aly499.78.3:A371T, aly 1249.8.1:C1504A, aly 1249.8.1:A1513C, and aly1603.54.2:G937T.
Etymology. The name is a feminine noun in the nominative singular, formed from the type species name.
Species included. Only the type species.
Parent taxon. Genus Mnasicles Godman, 1901 .
Cobalus asella Herrich-Schäffer, 1869 is a junior subjective synonym of Amblyscirtes alternata (Grote and Robinson, 1867)
Placed by Evans (1955) as a junior subjective synonym of Amblyscirtes vialis (W. H. Edwards, 1862) (type locality USA: Illinois, Mercer Co.), Cobalus asella Herrich-Schäffer, 1869 (type locality not specified) has not changed its status since. A single syntype of C. asella that we found and sampled (NVG-18042H03) in the ZMHB collection, groups with Amblyscirtes (Amblyscirtes) alternata (Grote and Robinson, 1867) (type locality USA: Georgia) in the genomic trees ( Fig. 13 View Figure 13 ) and phenotypically is consistent with A. alternata , not A. vialis , due to very short apiculus (extended in A. vialis ) and more rounded wings. The specimen NVG-18042H03 is a syntype of C. asella because it bears the labels typical of Herrich-Schäffer, 1869 types: old darkening elongated handwritten label with the word “asella” on it and printed “Coll. H.–Sch” label, and agrees with the original description. Due to genetic and phenotypic similarities, we propose that Cobalus asella Herrich-Schäffer, 1869 is a junior subjective synonym of Amblyscirtes alternata (Grote and Robinson, 1867) .
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