Polytrichophora mimbres, Mathis, Wayne N. & Zatwarnicki, Tadeusz, 2012

Mathis, Wayne N. & Zatwarnicki, Tadeusz, 2012, A revision of the new world species of Polytrichophora Cresson and Facitrichophora, new genus (Diptera, Ephydridae), ZooKeys 231, pp. 1-116 : 53-54

Polytrichophora mimbres sp. n. Figs 8993

Diagnosis.

This species is distinguished from congeners by the following combination of characters: Small to moderately small shore flies, body length 1.85-2.25 mm.

Head: Frons dull to very faintly subshiny, tan with moderate investment of gray to whitish gray microtomentum; fronto-orbits narrow, gray to whitish gray. Antenna mostly yellow; basal flagellomere tannish yellow apicodorsally; arista with 5 dorsal rays. Face at narrowest point about equal to combined length of pedicel and basal flagellomere; face more densely microtomentose than frons, microtomentum faintly shiny, mostly gray to whitish gray; parafacial and gena becoming slightly more whitish to silvery white than face; parafacial becoming very slightly wider ventrally with little ventral dilation; gena moderately short, height less than height of basal flagellomere; gena-to-eye ratio 0.11-0.13.

Thorax: Mesonotum mostly dull to faintly subshiny, moderately densely microtomentose, mostly gray to tannish gray, completely gray laterally and anteriorly; pleural area mostly gray. Anterior margin of wing lacking spine-like setae; costal vein ratio 0.86-0.88; M vein ratio 0.50-0.56. Legs, except for tarsi but including most of coxae, gray; tibiae yellowish at apices; forefemur lacking a row of 9-10 short, stout setulae along apical half of anteroventral surface; forefemur with a row of 4-5, of moderately well-developed, evenly spaced posteroventral setae on apical half, each seta less than width of femur; tibiae mostly gray, only apices yellowish; basal tarsomeres yellow, apical 1-2 becoming slightly darker, mostly tannish yellow.

Abdomen: Tergites dull, gray, medially with some grayish green, metallic luster; tergite 5 of male pointed apically, lacking row of 6-10, distinctly larger setae along extreme posterior margin with posterodorsal orientation. Male terminalia (Figs 89-92): Epandrium in posterior view (Fig. 89) generally elliptical, relatively uniformly setulose, dorsal margin rounded, both ventral extensions relatively wide, bluntly pointed, slightly turned medially, in lateral view (Fig. 90) with ventral apex acutely pointed, posterior margin shallowly concave; cercus large, conspicuous; aedeagus tubular but with distinct basiphallus and more membranous, more narrowly developed distiphallus, basiphallus tubular, distiphallus curved, irregularly tapered to narrow apex, in ventral view (Fig. 91) narrowly elongate with lateral margins very shallowly concave; phallapodeme in lateral view (Fig. 92) roughly triangular with extended keel truncate apically, in ventral view (Fig. 91) bluntly clavate at both apices; gonite in lateral view (Fig. 92) narrowly elongate, shallowly curved, in ventral view (Fig. 91) angularly arched, slightly wider toward base of aedeagus; hypandrium in lateral view (Fig. 92) elongate but relatively wide, in ventral view (Fig. 91) with anterior margin deeply and broadly rounded, posterior margin as lateral, parallel-sided projections.

Type material.

The holotype male is labeled "USA. N[EW] M[EXICO]. Grant: Mimbres River (32°43.8'N, 107°52'W; 1665m), 22 Aug 2009, W. N. Mathis/USNM ENT 00285965 [plastic bar code label]/HOLOTYPE ♂ Polytrichophora mimbres Mathis & Zatwarnicki, USNM [red]." The holotype is double mounted (minuten pin in a block of plastic), is in excellent condition, and is deposited in the USNM. Forty-two paratypes (37♂, 5♀; USNM) are from the same locality as the holotype with dates from 1-22 Aug 2008, 2009.

Type locality.

United States. New Mexico. Grant: Mimbres River (New Mexico Highway 61 & Royal John Mine Road; 32°43.8'N, 107°52'W; 1665 m).

Other specimens examined.

Nearctic. UNITED STATES. COLORADO. Rio Grande: South Fork (37°40.2'N, 106°38.4'W; 2440 m; Malaise trap), 20 Jun 1972, W. W. Wirth (1♀; USNM).

NEVADA. Clark: Las Vegas Wash (36°2.3'N, 114°58.8'W), 10-11 May 2004, D. and W. N. Mathis (4♂, 7♀; USNM); Las Vegas Wash (36°5.2'N, 114°58.9'W), 10-13 May 2004, D. Mathis (1♀; USNM).

NEW MEXICO. Catron: Gila River (33°13.6'N, 106°15.1'W; 1750 m), 15 Aug 2007, D. and W. N. Mathis (1♂; USNM). Grant: Bill Evans Lake (32°52.1'N, 108°34.5'W; 1416 m), 14 Aug 2007, D. and W. N. Mathis (4♂; USNM); Silver City (Big Ditch; 32°46.4'N, 108°16.5'W; 1790 m), 14 Aug 2007, D. and W. N. Mathis (2♂; USNM). Sandoval: La Cueva (Junction of Highways 126 & 4; 35°52'N, 106°38.4'W; 2342 m), 14 Jun 2011, D. and W. N. Mathis (3♂, 1♀; USNM).

TEXAS. Blanco: Miller Creek (30°15.2'N, 98°31.7'W; 410 m), 3 Jun 2004, W. N. Mathis (2♂; USNM). Kinney: Del Rio (30 km E; Pinto Creek; 29°20'N, 100°31.6'W), 5 Jun 2004, W. N. Mathis (12♂, 9♀; USNM). San Patricio: Mathis (6.5 km S; Nueces River; 28°02.8'N, 97°51.8'W; 18 m), 5 Jun 2004, W. N. Mathis (1♂; USNM). Travis: Austin (Zilker Park; 30°15.8'N, 97°46.3'W), 2 Jun 2004, W. N. Mathis (2♂; USNM). Uvalde: Garner State Park (29°35.3'N, 99°44.3'W), 3 Apr 1955, W. W. Wirth (1♂; USNM).

UTAH. Emery: Green River (Green River; 38°59.6'N, 110°08.5'W; 1240 m), 31 Jul 2007, D. and W. N. Mathis (1♂; USNM); Green River (3.3 km N; 39°01.7'N, 110°09.7'W; 1253 m), 30 Jul-5 Aug 1992, 2007, D. and W. N. Mathis (10♂; USNM); San Rafael River (22.5 km SW Green River; 38°55.7'N, 110°24.5'W; 1270 m), 31 Jul 2007, D. and W. N. Mathis (5♂, 1♀; USNM). Garfield: Willow Tank-Hurricane Wash (37°23.2'N, 111°08'W; stock tank), 22 May 2001, D. and W. N. Mathis (1♂, 1♀; USNM). Grand: Thompson Spring (8.9 km N Thompson Springs; 39°02.3'N, 109°43.4'W; 1740 m), 16 Aug 2008, D. and W. N. Mathis (1♂; USNM). Salt Lake: Draper (40°31.6'N, 111°55.1'W; Jordan River; 1320 m), 10 May 2007, D. and W. N. Mathis (1♂; USNM). Utah: Thistle (40°0.4'N, 111°29.7'W; 1530 m), 11 May 2007, D. and W. N. Mathis (1♂; USNM).

Distribution

(Fig. 93). Nearctic: United States (Colorado, Nevada, New Mexico, Texas, Utah).

Etymology.

The species epithet, mimbres, is a noun in apposition and refers to the river in southern New Mexico where the type series was collected.

Remarks.

Among species of the orbitalis group, this species is distinguished by characters of the male terminalia as follows: Area between anterior arms of the hypandrium is partially sclerotized; the ventral epandrial processes are relatively wide, bluntly pointed, slightly turned medially (Fig. 89); the posteroventral edge of the epandrium in lateral view is very shallowly concave, and the taper at the apex of is more abrupt (Fig. 90).