Tylopus species

Nguyen, Anh D., 2012, Tylopus Millipedes In Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini), With Descriptions Of Five New Species, Raffles Bulletin of Zoology 60 (2), pp. 289-311 : 308-310

publication ID

https://doi.org/ 10.5281/zenodo.5347271

persistent identifier

https://treatment.plazi.org/id/191D7C72-CD31-E80E-FF0C-9538FA21FB4F

treatment provided by

Tatiana

scientific name

Tylopus species
status

 

Key to Tylopus species known from Vietnam (male)

This key was made based on examined material and literature ( Attems, 1937, 1953; Golovatch, 1984; Korsós & Golovatch, 1989; Golovatch & Enghoff, 1993; Likhitrakarn et al., 2010).

1. Sterna except sternum 5 without any modifications ............. 2

– All sterna with modifications .............................................. 17

2. Sternum 5 with more than two processes between coxae 4. Tip of gonopod with process k ( Figs. 8D View Fig , 15B View Fig ) ......................... 3

– Sternum 5 with one or two processes between coxae 4. Tip of gonopod without process k ................................................... 4

3. Both gonopod process h and spine z absent ( Fig. 15 View Fig ). ........... ................ T. strongylosomoides (Korsós & Golovatch, 1989) View in CoL

– Gonopod process h present, thin, long and finger-shaped; spine z absent ( Figs. 7C,D View Fig , 8A–D View Fig ) ....... T. golovatchi View in CoL , new species

4. Paraterga pink. Metatergal surface smooth. Gonopod process h present, short; process z absent ( Figs. 5E,F View Fig , 6A–C View Fig ) ............ ................................................ T. roseiparaterga View in CoL , new species

– Paraterga not pink. Metatergal surface rugulose or rugose. Gonopod process h present; spine z present or absent ........ 5

5. Gonopod spine z present ....................................................... 6

– Gonopod spine z absent ...................................................... 13

6. Gonopod spine z strong, horizontal, coarsely serrated along distal margin .......................................................................... 7

– Gonopod spine z erect or bent mesad, not serrated along distal margin .................................................................................... 9

7. Spine z only coarsely serrated along 2/3 distal margin. Process h short, hook-shaped or strongly spiniform ( Figs. 11D,E View Fig , 12A,B View Fig ) ................................................................................... 8

– Spine z coarsely serrated along whole distal margin. Process h with two branches, one spiniform, the other a serrated lamina ( Figs. 1E,F View Fig , 2A,B View Fig ) ....................... T. phanluongi View in CoL , new species

8. Metatergal transverse sulcus present from body ring 5. Metatergum 18 with only a single row of 2+2 setae on anterior half. (North Vietnam) ............. T. hilaroides Golovatch, 1984 View in CoL

– Metaterga transverse sulcus present from body ring 4. Metatergum 18 with two transverse rows of setae: 2+2 before and 3+3 after sulcus (southern part of Central Vietnam) ........ ......................................................... T. hilaris ( Attems, 1937) View in CoL

9. Gonopod with three more or less spiny processes including process h and spine z near base of lamina l. Male legs without tubercles ...................................... T. mutilatus ( Attems, 1953) View in CoL

– Gonopod with only process h and spine z near base of lamina l. Male legs with or without tubercles ................................ 10

10. Gonopod lamina l normal. Spine z very small, dentiform. Process h large, lamellate, sigmoid .................................................. 11

– Gonopod lamina l well-developed, distal part produced into long, erect spine z. Process h erect, spiniform without evidence of torsion ............................................................................. 12

11 Spine z bent mesad. Lobe m strongly developed, spiniform ( Fig. 13 View Fig ) ..................................... T. nodulipes ( Attems, 1953) View in CoL

– Spine z normal. Lobe m modestly developed, somewhat lamellate, not spiniform ........ T. granulatus Golovatch, 1984 View in CoL

12. Body shorter than 20 mm. Large tubercles present on male legs, especially femora 6, 9 and 10 with proximal finger-shaped tubercles crowned with a bunch of setae. Gonopod process h obviously longer than spine z ( Fig. 17A,B View Fig ) ............................ ........................................................ T. topali Golovatch, 1984 View in CoL

– Body longer than 20 mm. Microtubercles present on male legs ( Fig. 10F View Fig ). Femora 6, 9, 10 without tubercles crowned with a bunch of setae. Gonopod process h obviously shorter than spine z ( Figs. 9D,E View Fig , 10A–E View Fig ) .................. T. sapaensis View in CoL , new species

13. Metatergal transverse sulcus present from body ring 4, but fully developed and reaching base of paraterga from body ring 5 .............................................................................................. 14

– Metatergal transverse sulcus present from body ring 5 ..... 15

14. Process h with an outgrowth at middle ( Fig. 16 View Fig ) .................... ................................................. T. procurvus Golovatch, 1984 View in CoL

– Process h without an outgrowth at middle ( Fig. 14 View Fig ) ............... .................................................. T. crassipes Golovatch, 1984 View in CoL

15. Body small, width <1.6 mm. Gonopod process h short, spiniform ......................................... T. sigma ( Attems, 1953) View in CoL

– Body large, width> 1.6 mm. Gonopod process h well-developed, not spiniform, somewhat lamellate ..................................... 16

16. Midbody metaterga with a single row of 2+2 setae after transverse sulcus. Process h sub-transverse, coiled with an additional spine at base ............. T. magicus Golovatch, 1984 View in CoL

– Midbody metaterga with two rows of 2+2 and 3+3 setae before and after transverse sulcus, respectively. Process h suberect, without any additional spine .................................................... ............................. T. tamdaoensis Korsós & Golovatch, 1989 View in CoL

17. Metaterga strongly rugose with two rows of 2+2 and 2+2 setiferous knobs before and after transverse sulcus. Each sternum with four projections ( Fig. 3C View Fig ). Gonopod simple with a long, slightly twisted process h. Solenophore suberect, coiled, and as high as process h ( Figs. 3E,F View Fig , 4A–C View Fig ) ................................. ...................................................... T. spinisterna View in CoL , new species

– Metaterga rugose, with two rows of 2+2 and 3+3 setiferous tubercles. Each sternum with a pair of small spines at base of posterior legs.Gonopod process h poorly developed. Solenophore strongly coiled, completely sheathing solenomere................... ................................................. T. maculatus Golovatch, 1984 View in CoL

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