Euconnus (Cladoconnus) motschulskii Motschulsky

Hlaváč, Peter & Stevanović, Miroslav, 2013, A review of the subgenus Cladoconnus Reitter of the genus Euconnus Thomson (Coleoptera: Staphylinidae: Scydmaeninae) from the Balkan Peninsula, Turkey and Caucasus, Zootaxa 3646 (4), pp. 401-425 : 419-420

Euconnus (Cladoconnus) motschulskii Motschulsky

( Figs. 1–6 )

Scydmaenus motschulskii Motschulsky, 1837: 119 . Type data: unknown

Euconnus motschulskyi Sturm : Croissandeau, 1898: 119; Castellini, 2006: 90.

Euconnus (s.str.) motschulskyi Sturm : Ganglbauer, 1899: 48; Reitter, 1882: 574.

Euconnus (Cladoconnus) motschulsky Sturm : Karaman, 1974: 134, Fig. 16 .

Euconnus dubius J. Sahlberg, 1913: 46 . Type locality: not given, synonymy in Davies, 2004: 207.

Euconnus europaeus Csiki, 1919: 44 . A new name for Euconnus kiesenwetteri Reitter, 1881: 574 , not Kiesenwetter, 1851, unjustified replacement name, synonymy in Davies, 2004: 207.

Euconnus mothoulskii Sturm, 1838: 25 (misspeling).

Euconnus rufescens Ganglbauer, 1899: 48 . Type locality: not given, synonymized by Davies, 2004: 26.

Euconnus skalitzkyi Croissandeau, 1893: 113 . Type locality: unknown, synonymized by Franz, 1964: 160.

Scydmaenus major Motschulsky, 1851: 598 , syn. nov. Type locality: vallée de Nanos, Birnbaumer-Wald [in the enviroment of] „la grotte de Luck" [Volčja jama, Nanos Mts., Slovenia].

Material examined (118 3, 30 Ƥ): AUSTRIA: 1 3: Karawanken, Hochstuhl. (HNHM). 2 3: Austria mer. 900 m, Ostkarawanken, Jovan Berg, 16.8.2000, J. Jelínek leg. (JJCC). SLOVENIA: 2 3: Podklanec, 26.vi.2004, J. Vávra lgt. (JVOC). 1 3: Sajevce env, Krakovski Gozd forest, Kostanjevica na Krki env, 2–3. vii. 2004, J. Vávra lgt. (JVOC). 4 3: Sajevce env, Krakovski Gozd forest, 28.vi. 2003, J. Vávra lgt. (JVOC). 6 3: Luče env, Pečovnik, 6.v.2001, J. Vávra lgt. (JVOC, PHKS). 4 3: Ocizla, 7.v.2001, J. Vávra lgt. (JVOC, PHKS). 3 3: Podgorski kras mts, Poljane, 14.vii.2000, J. Vávra lgt. (JVOC, PHKS). 4 3: Stojna mts., Kočevje env, 900m, 6–7.vii.2002, Vávra lgt. (JVOC). 7 3, 2 Ƥ: Julische Alpen, gesibt aus Moos und Laub, 550 m, 1.viii.1969, S. Horvatovich lgt. (HNHM). 2 3: Slovenija-VM34, Karawanken-Stol, 2000m, 0 6.07.1999, leg. Broder. (JBKS). CROATIA: 1 3: Istria, Buzet env., 23.vii.2010, sifting, P. Hlaváč lgt. (PHKS). 1 3: Ravna Gora, Peščenica Vinička, Plitvička Špilja, 25.v.2007, R. Ozimec lgt. (PHKS). 1 3: Istra: Limski kanal, 2.iv.2010, P. Hlaváč, sifting in forest near sea. (PHKS). 4 3, 2 Ƥ: Sarajevo, Igman Planina, 5.xi.1929, Dr. J. Fodor, lgt. (HNHM). 1 3: E Croatia, Moslavčka Gora hills, N Kutina, 13.5.2004. leg. P. Białooki. (PJWP). MONTENEGRO: 2 3: Orjen, Biela Gora, 11.6.2005, Stevanoviċ lgt. (PHKS). BOSNIA AND HERZEGOVINA: 1 3: 19km N-Konjič (E73: Sarajevo-Jalanica) Ivan sedlo, Fagetum, leaf litter, 900m, 19.x.2005, Hlaváč lgt. (PHKS). 4 3, 2 Ƥ: Sarajevo, Igman Planina, 5.xi.1929, Dr. J. Fodor, lgt. (HNHM). 1 3: Visegrad, Dabrum, 11.x.1935, Dr. J. Fodor, lgt. (HNHM). 4 3: Igman plan, 13.x.1929, Dr. J. Fodor, lgt. (HNHM). 1 3: Rovinaja Planina, 7.vii.1931, Dr. J. Fodor, lgt. (HNHM). 1 3: Vareš Han Milosnica, 30.vii.1928, Dr. J. Fodor, lgt. (HNHM). 1 3: Trebevic, Gerger lgt.Vareš Han Milosnica, 30.vii.1928, Dr. J. Fodor, lgt. (HNHM). 1 3: Pazaric, 1931, Fodor lgt. (HNHM). 3 3: Rajtov Han, bei Vareš, 25.ix.1929, Dr. J. Fodor, lgt. (HNHM). 3 3: Jablanica, 9.x.1929, Dr. J. Fodor, lgt. (HNHM). 3 3: Varos, Zvdzda, Fodor. (HNHM). 1 3: Herzegovina, Planina Orjen, Bjela Gora, 1000m, 11.vi.2005, leg. D. Paviċeviċ. (PHKS). 2 3: Herzegovina, Plasa planina, Fodor. (HNHM). 2 3: Bosnia, Zvijezda Planina, 14–18.vii.1930, Dr. J. Fodor lgt. (HNHM, PHKS).

SERBIA: 2 3: eastern Serbia, Zvonačka Banja env, sifting, 28.iv.2002, P. Hlaváč lgt. (PHKS). 1 3: Ovčar- Kablarsk Kl. [Kablarska klisura], Debela gora, 16.4.– 19.5.1996, D. Paviċeviċ lgt. (PHKS). 2 3: Suva Planina, Bojanine Vode, 14.vi.2001, M. Stevanoviċ lgt. (PHKS). 1 3: east Niš, Suva Planina, Jelov Pažar, 4.vii.2006, Stevanoviċ lgt. (PHKS). 1 3, 1 Ƥ: Svrljiške planine, Ribarska Korita, 16.v.2004, Stevanoviċ lgt. (PHKS). 3 3, 3 Ƥ: eastern Serbia, Niš env. Suva Planina, 890m, Bojanine Vode, sifting, 2.vii.2006, P. Hlaváč lgt. (PHKS). 1 3: Kopaonik, Jošanička Banja, 6.vi.2009, M. Stevanoviċ lgt. (PHKS). 4 3, 4 Ƥ: Serbia, 6.6.2012, Majdanpek env, V. Zieris lgt. (VZPC). MACEDONIA: 3 3, 1 Ƥ: Galičnik, Bistra Planina, 7–14. vii. 1937, Dr. J. Fodor lgt. (HNHM). 1 3: Crepolsko, Ketsana Stjena, 26.viii.1936, Dr. J. Fodor lgt. (HNHM). ROMANIA: 2 3: M?.?urean, vf. Nǎiivalea Riu?or, ca. 600m, 11.vi.1998, P. Moravec lgt. (PHKS). 3 3, 1 Ƥ: M?. Vǎlcan, Lupeni env, 800–900 m, 22– 25.vii.2002, P. Moravec lgt. (PHKS). 3 3, 1 Ƥ: M?. Cǎpǎ?inii, cheile Olte?ului-Polovragi, 650–800m, 17.vi.1998, P. Moravec lgt. (PHKS). 1 3: Baie Herculane env., Vf. Demégled, 200–600 m, 27.6.1996, P. Bulirsch lgt. (PHKS). 1 3: Nermed p. Re?i?a, 24.6.1996, P. Bulirsch lgt. (PHKS). 1 3: Nermed p. Re?i?a, 24.6.1996, P. Bulirsch lgt. (PHKS). 4 3, 6 Ƥ: Romania, M?. Trascǎului, Baia de Arieş, Cioara de Sus, 27.5.2010, V. Zieris lgt. (VZPC). 2 3: Romania, M?. Trascǎului, Sub Piatra, 3.6.2010, Huda lui Paparǎ env, sifting, V. Zieris lgt. (VZPC). 2 3, 4 Ƥ: Romania, 25.5.2010, M?. Trascǎului, Rimetea env, vf. Piatra Secului, V. Zieris lgt. (VZPC). 1 3, 3 Ƥ: Romania, 2.6.2010, M?. Trascǎului, Poiana Aiudului env, V. Zieris lgt. (VZPC). 9 3: 0 3.6.2004 SE Romania, SE Crasna, SE Intorsura Bazaului, leg. P. Białooki. (PHKS, PJWP).

Description: Body ( Fig. 1 ) shiny, lighter or darker reddish-brown, body length 2.10–2.20 mm, body width 0.80–0.85 mm; with long, golden pubescence, sparse on elytra, erect on sides of pronotum and posterior part of head. Head slightly wider than long, tempora evenly convergent posteriorly; eyes protuberant; supra-antennal tubercles weakly protuberant. Antennae ( Fig. 5 ) 1.20–1.30 mm long; antennomeres III–VI subequal, antennomere VII transverse, 1.30 time as wide as long, slightly wider than VI; antennomere VIII about 1.70 times as long as wide, about 3 times as long as VII and 1.30 times as long as IX; X subquadrate; terminal antennomere 1.80 time as long as wide and about twice as long as X. Pronotum 1.15–1.25 as long as wide. Elytra 1.50–1.55 times as long as wide, sutural striae reaching basal fourth of elytral length. Aedeagus 0.47 mm in length, as in Fig. 6 .

Distribution: Austria, Bosnia and Herzegovina, Croatia, Czech Republic, France, Germany, Greece, Hungary, Italy, Macedonia, Montenegro, Poland, Romania, Serbia, Slovenia, Ukraine (Davis, 2004).

Differential diagnosis: Euconnus motschulskii differs from other species of the denticornis group by large body, 2.10–2.20 mm long, antennomere VII transverse, 1.30 times as long as wide, antennomere VIII about 1.70 times as long as wide, subquadrate antennomere X. Euconnus motschulskii is similar to E. winneguthi , both are largest species of the subgenus within the studied area. Euconnus motschulskii differs from E. winneguthi in different structure of antennomeres VII–XI and aedeagus.

Note I: The original description of Scydmaenus major by Motschulsky (1851) is too general and simple to allow recognizing this species. He is just comparing it on the general level with E. motschulskii and Stenichnus godarti Latreille, 1806 . Description is most probably based on female because the modification of antennal club is not mentioned, no dimessions are given either. So, the identification of the species is not possible. The finding of this species has never been mentioned since its description although Slovenia is entomologically very well explored country. Species has not been mentioned in further taxonomical studies from the area (Reitter, 1882; Karaman, 1974; Franz, 1971b). All these facts lead us to the conclusion that this species is identical with E. motschulskii .

Note II: Euconnus motschulskii is a common species in mountainous regions of the Balkan Peninsula and eastern part of the Alps. The occurrence in Czech Republic, France, Germany, Hungary and Poland is highly doubtfull and must be confirmed by new records.

Figures

FIGURES 1 – 6. Euconnus motschulskii. 1, male, dorsal habitus (scale: 1 mm); 2, labrum (scale: 0.2 mm); 3, right madible (scale: 0.2 mm); 4, maxilla (scale: 0.2 mm); 5, antennae of male (scale: 0.2 mm); 6, aedeagus, dorsal aspect (scale: 0.2 mm).

FIGURES 15 – 22. 15, Euconnus goerzensis, antennae of male (scale: 0.2 mm); 16, E. goerzensis, aedeagus, dorsal aspect (scale: 0.1 mm); 17, E. ispartaensis, antennae of male (scale: 0.2 mm); 18, E. ispartaensis, aedeagus, dorsal aspect (scale: 0.1 mm); 19, E. meybohmi, antennae of male (scale: 0.2 mm); 20, E. meybohmi, aedeagus, dorsal aspect (scale: 0.1 mm); 21, E. robustus, antennae of male (scale: 0.2 mm); 22, E. robustus, aedeagus, dorsal aspect (scale: 0.1 mm).