Circumphallus, L. Papp, 2011

Papp, L., 2011, Description Of A New Genus And A New Family, Circumphallidae Fam. Nov., Of The Acalyptrate Flies (Diptera), Acta Zoologica Academiae Scientiarum Hungaricae 57 (4), pp. 315-341 : 316-317

Circumphallus gen. n.

( Figs 1–12 , 33–35 )

Type species: Circumphallus significans sp. n.

Gender: masculine.

Description. A small, rather slender fly with moderately thickened legs ( Fig. 33 ).

Head with protruding lower prefrontal edge and clypeus, chaetotaxy as described for family. Proboscis shorter than head with a medium-sized labella, dorso-apically with a medial 0.06 mm long thick sharp seta. Palpus normal, dark, with a medium-long (0.10 mm) apical seta.

Thorax with chaetotaxy as described for family; in addition, postpronotal seta present, 1 presutural and 1 postsutural supra-alar, 2 notopleural, 1 strong prescutellar acrostichal seta, 1 postalar and 2 scutellar setae, all strong. No propleural, anepisternal or katepisternal setae, posterior margin of anepisternum with 1–2 fine setulae, similar setulae on katepisternum. Acrostichal setulae medially unarranged and scattered.

Legs short and thickened, fore femur with 4 long posteroventral setae, mid tibia with one thickened ventroapical seta (“spur”). Pulvilli large, plumose.

Wings ( Fig. 34 ) hyaline, elongate. Costal vein reaching apex of R 4+5 but vanishing before M 1 apex, long setulose over whole its length, setulae longer, or as long as vein wide. Subcostal vein strong in its whole length, more or less parallel to R 1 and reaching costal vein without attenuation. Vein R 4+5 ending anterior of wing apex, cell r 4+5 twice as wide as cell r 2+3. Cell dm short and slightly proximal of R 1 apex. Basal medial cell closed, i.e. basal crossvein present. Cubital cell (cup) short, closed with an arcuate vein. A 1 long, parallel to posterobasal margin in most of its length but reaching wing margin.

Calypters very small, thoracic calypter hardly discernible, alar calypter with sparse dark setulae. Halter normal ochreous.

Abdominal tergites 1–5 shiny in anterior half and microtrichose in posterior half ( Fig. 33 ), with long erect setae on disc, but without marginal setae, usually as long as tergite ( Fig. 35 ). Tergites 1–5 on sides ventrally curved, sternites very narrow: sternite 2 in 2 small parts. Tergites 2–5 bipartite ( Fig. 35 ): they consist of a shiny caudal and a dull basal part each, and tergites with extremely long discal (i.e. not marginal) setae. Spiracles, including 6th pair, in membrane, rather far from tergal edges.

Syntergosternite 6–8 very short, forming single irregular ring and deeply hidden underneath tergite 5.

Male genitalia as described for the family.

to R 1 vein portion of the wing. Scales: 0.2 mm for all part in broadest view (most of the thorns covered). Scales: 0.1 mm for all

Figures

Figs 1–2. Circumphallus significans sp. n., holotype male. 1 = 5th abdominal segment and postabdomen with genitalia(ejaculatory apodeme and distiphallus omitted, surstyli schematic), 2 = humeral

Figs 3–7. Circumphallus significans sp. n., holotype male. 3 = all postabdominal sclerites (syntergosternite) in the view, when dorsal part is at widest, 4 = postabdominal sclerites (syntergosternite), left lateral view, 5 = same, right lateral view, 6–7 = surstylus: 6 = subanterior-sublateral view, 7 = apical

Figs 8–12. Circumphallus significans sp. n., holotype male. 8 = phallus and contours of epandrium, 9 = basal part of phallus and medial part of its second circle in higher magnification, 10 = ejaculatory apodeme, broadest view, 11 = hypandrial plates with basiphallus and contours of epandrium, surstyli and base of phallapodeme ventral view (dashed: insertion surface of phallapodeme and distiphallus/basiphallus), 12 = phallapodeme with edges of surstyli and contours of basiphallus, in its broadest, i.e.

Figs 33–35. Circumphallus significans sp. n., holotype male. 33 = habitus; photo taken before abdomen was removed; 34 = right wing; 35 = abdomen, lateral view