Lepanus ustulatus ( Lansberge, 1874 )

Gunter, Nicole L. & Weir, Thomas A., 2021, Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species, Zootaxa 4923 (1), pp. 1-66: 12-19

publication ID

https://doi.org/10.11646/zootaxa.4923.1.1

publication LSID

lsid:zoobank.org:pub:41D057A7-F855-47C2-A598-8000CFD83A5B

DOI

http://doi.org/10.5281/zenodo.4537921

persistent identifier

http://treatment.plazi.org/id/1A323678-6225-220E-09D9-FE2EFD326BAD

treatment provided by

Plazi

scientific name

Lepanus ustulatus ( Lansberge, 1874 )
status

 

Lepanus ustulatus ( Lansberge, 1874)  

( Figs. 2 View FIGURE 2 A–B, D–E; 3E; 4E; 5E; 6A–B)

Epilissus ustulatus Lansberge, 1874: 7   ; Masters 1886: 10; Gillet 1911: 39; 1925: 5; Paulian 1934: 282.

Panelus ustulatus: Lea 1923: 361   .

Lepanus ustulatus: Matthews 1974: 104   .

Type material digitally examined. Lectotype: “Wide Bay” (handwritten) / “Ex-Musaeo VAN LANSBERGE” (printed) / “MUSÉUM PARIS 1952 COLL. R. OBERTHÜR” (printed) / “ LECTOTYPE ” / “ Lepanus ustulatus (Lansberge)   . LECTOTYPE, Y. CAMBEFORT det. 1981” (hand written) / “ MNHN EC8675” (printed); Paralectotypes: “Ex-Musaeo VAN LANSBERGE” (printed) / “R. PAULIAN Vidit” (printed)/ “MUSÉUM PARIS 1952 coll. R. OBERTHÜR” (printed) / “ PARALECTOTYPE ” / “ PARALECTOTYPE Lepanus ustulatus ( Lansberge 1874)   ” (printed) / “ MNHN EC8676” (printed); “Ex-Musaeo VAN LANSBERGE” (printed) / “R. PAULIAN Vidit” (printed)/ “MUSÉUM PARIS 1952 coll. R. OBERTHÜR” (printed) / “ PARALECTOTYPE ” / “ PARALECTOTYPE Lepanus ustulatus ( Lansberge 1874)   ” (printed) / “ MNHN EC8677” (printed); “Ex-Musaeo VAN LANSBERGE” (printed) / “R. PAULIAN Vidit” (printed)/ “MUSÉUM PARIS 1952 coll. R. OBERTHÜR” (printed) / “ PARALECTOTYPE ” / “ PARALECTOTYPE Lepanus ustulatus ( Lansberge 1874)   ” (printed) / “ MNHN EC8678” (printed); “Brisbane” (handwritten) / “Ex-Musaeo VAN LANSBERGE” (printed) / “R. PAULIAN Vidit” (printed)/ “MUSÉUM PARIS 1952 coll. R. OBERTHÜR” (printed) / “ PARALECTOTYPE ” / “ PARALECTOTYPE Lepanus ustulatus ( Lansberge 1874)   ” (printed) / “ MNHN EC8679” (printed).

Other material examined. 1407 other specimens from over 200 unique localities.

Queensland: Mount Beecher , 0.5 km NW, 23°54’58”S, 151°10’53”E, 27.ix–15.xii.1999 (T105335 1 specimen in QM) GoogleMaps   ; Eurimbula Creek via Round Hill Head , [24°12’00”S 151°46’48”E], 2–5.v.1975, R.I. Storey, (25-058453 1 specimen in QM) GoogleMaps   ; Bulburin Barracks, 600 m, via Builyan [24°31’39”S, 151°28’01”E], 18.ix–2.xii.1989 ( T80281 View Materials T80303 View Materials 23 specimens in QM) GoogleMaps   ; Bulburin Barracks, 600 m, via Builyan [24°31’39”S, 151°28’01”E], 2.xii.1989 – 16.i.1990 ( T80304 View Materials T80319 View Materials 16 specimens in QM) GoogleMaps   ; Bulburin Barracks, 600 m, via Builyan , [24°31’39”S, 151°28’01”E], 7–9.x.1999 ( T89647 View Materials T89648 View Materials 2 specimens in QM) GoogleMaps   ; Bulburin State Forest , [24°32’00”S 151°23’00”E], 8.xii.1972, R.I. Storey, (25-058565 specimens in QDPC) GoogleMaps   ; Bulburin National Park , 24°31’26”S, 151°28’08”E, 570 m, 11–12.iv.2012, G.B. Monteith, (25-053941 2 specimens in ANIC, 25-053942 4 specimens in ANIC) GoogleMaps   ; Bulburin State Forest , [24°35’00”S 151°35’00”E], 12–14.iv.1974, B.P. Moore, (25-058331 1 specimen in ANIC) GoogleMaps   ; Mount Fort William, via Kalpowar [24°39’06”S, 151°20’24”E], 18.i.1990 ( T50268 View Materials T50270,T80268 T80270 View Materials 6 specimens in QM) GoogleMaps   ; Mount Fort William, via Kalpowar [24°39’06”S, 151°20’24”E], 2.xii.1989 – 16.i.1990 ( T50271, T80271 2 specimens in QM) GoogleMaps   ; Hurdle Gully , 10.8 km WSW Monto, 24°54’02”S, 151°01’10”E, 20–21.xii.1997 ( T85382 View Materials 1 specimen in QM) GoogleMaps   ; Forest Station, Bulburin State Forest, via Many Peaks , [24°32’00”S, 151°23’00”E], 200m, 2– 5.iv.1972, S.R. Monteith, (25-058446 13 specimens in QM) GoogleMaps   ; Ventor Tableland, 21km ESE Monto, Site 1, 24°53’35”S, 151°19’59”E, 475 m, 1.x.2013 – 5.xii.2013, G.B. Monteith, (25-057705 15 specimens in ANIC) GoogleMaps   ; One Tree Hill , 1 km SE, 25°17’46”S, 151°54’36”E, 22–26.i.2001 (T105674–T105676 3 specimens in QM) GoogleMaps   ; One Tree Hill , 1.6km ENE, 25°17’05”S, 151°55’10”E, 14.xii.1999 – 19.iii.2000 ( T95331 View Materials 1 specimen in QM) GoogleMaps   ; Yidney Scrub, Fraser Island , [25°19’00”S, 153°11’00”E], 1–4.xii.1975, G. Thompson & J.A. Slater, (25-058450 1 specimen in QM) GoogleMaps   ; Fraser Island, Garawongera Lake , 25°19’54”S, 153°09’14”E, 1–30.xi.1992 ( T80343 View Materials 1 specimen in QM) GoogleMaps   ; Mount Woowoonga , 1.4 km SW, 25°26’28”S, 152°05’55”E, 21–22.i.2001 (T105673 1 specimen in QM) GoogleMaps   ; Mount Woowoonga , 1.4 km SW, 25°26’28”S, 152°05’55”E, 20–23.iii.2001 (T105315–T105321 7 specimens in QM) GoogleMaps   ; Fairlies Knob , 0.5 km SE, 25°30’32”S, 152°17’22”E, 20.xii.2000 – 23.iii.2001 (T105685 1 specimen in QM) GoogleMaps   ; Fairlies Knob , 0.5 km SE, 25°30’32”S, 152°17’22”E, 20.x–20.xii.2000 (T105694–T105696 5 specimens in QM) GoogleMaps   ; Wetheron , 3.5 km SW, 25°34’01”S, 151°41’13”E, 19.xii.1998 – 27.i.1999 ( T85224 View Materials 1 specimen in QM) GoogleMaps   ; Teddington Weir , 25°39’11”S, 152°40’08”E, 30 m, 4–5.i.2012, G.B. Monteith, (25-053880 1 specimen in ANIC) GoogleMaps   ; Tinana Creek , 25°45’07”S, 152°41’38”E, 35 m, 6–7.xii.2011, G.B. Monteith, (25-053885 1 specimen in ANIC) GoogleMaps   ; Tallegalla Weir Junction , 25°45’25”S, 152°41’49”E, 35 m, 6–7.xii.2011, G.B. Monteith, (25-053886 3 specimens in ANIC) GoogleMaps   ; Stockhaven , 3 km SE, 25°48’03”S, 151°59’05”E, 10.x–19.xii.1998 ( T85323 View Materials T85324 View Materials 2 specimens in QM) GoogleMaps   ; Yarraman, Cooyar Range , 26°52’08”S, 151°54’10”E, 7.i–2.iii.1992, ( T78237 View Materials T78242 View Materials 6 specimens in QM) GoogleMaps   ; Yarraman, Cooyar Range , 26°52’08”S, 151°54’10”E, 2.iii–12.iv.1992 ( T78243 View Materials T78244 View Materials 2 specimens in QM) GoogleMaps   ; Yarraman, Cooyar Range , 26°52’08”S, 151°54’10”E, 1.xii.1991 – 7.i.1992 ( T78245 View Materials 1 specimen in QM) GoogleMaps   ; Cooloola, Womalah State Forest , 25°56’58”S, 153°05’31”E, 30.xi.1991 – 8.i.1992 ( T78039 View Materials T78053 View Materials 15 specimens in QM) GoogleMaps   ; Cooloola, Womalah State Forest , 25°56’58”S, 153°05’31”E, 8.i–3.iii.1992 ( T78076 View Materials T78081 View Materials 6 specimens in QM) GoogleMaps   ; Cooloola, Womalah State Forest , 25°56’58”S, 153°05’31”E, 3.iii–21.iv.1992 ( T78082 View Materials T78098 View Materials 17 specimens in QM) GoogleMaps   ; Cooloola, Womalah State Forest , 25°56’58”S, 153°05’31”E, 30.xi.1991 – 8.i.1992 ( T78099 View Materials T78110 View Materials 12 specimens in QM) GoogleMaps   ; Cooloola 3, 25°57’04”S, 153°06’43”E, 3.iii–21.iv.1992 ( T78054 View Materials T78068 View Materials 15 specimens in QM) GoogleMaps   ; Cooloola 3, 25°57’04”S, 153°06’43”E, 30.xi.1991 – 8.i.1992 ( T78069 View Materials T78074 View Materials 6 specimens in QM) GoogleMaps   ; Bymien, Cooloola , 25°57’14”S, 153°06’18”E, 150 m, 3–4.ii.2013, G.B. Monteith, (25-057693 2 specimens, 25-057694, 25-057695 3 specimens, 25-057698 1 specimen in ANIC) GoogleMaps   ; Freshwater Road, Cooloola , [25°58’00”S, 153°09’00”E], 9.xi.1974, G.B. Monteith, (25-058462 1 specimen in QM) GoogleMaps   ; Cooloola, via Gympie , [25°58’00”S, 153°09’00”E], 25–27.iv.1975, R.I. Storey, (25-058448 7 specimens in QM) GoogleMaps   ; Gunalda 3.6 km NE, [25°58’00”S 152°35’00”E], 140 m, 21–26.i.2001, D.J. Cook, (25-060844 2 specimens also as COL582, COL 583 specimens in ANIC) GoogleMaps   ; Gunalda , 3.6 km NE, 25°58’26”S, 152°35’07”E, 18–21.xii.2007 (T154645–T154650 6 specimens in QM) GoogleMaps   ; Gunalda , 3.6 km NE, 25°58’26”S, 152°35’07”E, 15–17.ii.2008 (T154651 1 specimen in QM) GoogleMaps   ; Gunalda , 3.6 km NE, 25°58’26”S, 152°35’07”E, 21– 26.i.2001 (T105677–T105678 2 specimens in QM) GoogleMaps   ; 2 km SE of Camp Milo, Cooloola National Park , [26°00’00”S, 153°05’00”E], 17–18.iv.1982, A. Calder, (25-057974 1 specimen in ANIC) GoogleMaps   ; Mudlo National Park , top, [26°00’40”S 152°13’52”E], 320 m, 20–21.xii.2011, G.B. Monteith, (25-053882 1 specimen in ANIC) GoogleMaps   ; Mudlo Gap , top site, 26°00’48”S, 152°13’46”E, 25.i–7.vi.1999 ( T95330, T85211 T85213 View Materials 4 specimens in QM) GoogleMaps   ; Mudlo Gap , top site, 26°00’48”S, 152°13’46”E, 17.xii.1998 – 25.i.1999 ( T85214 View Materials T85215 View Materials 2 specimens in QM) GoogleMaps   ; Mudlo Gap , top site, 26°00’48”S, 152°13’46”E, 9.x–17.xii.1998 ( T85216 View Materials 1 specimen in QM) GoogleMaps   ; Mudlo Gap , bottom site, 26°01’27”S, 152°13’57”E, 17.xii.1998 – 25.i.1999 ( T85209 View Materials T85210 View Materials 2 specimens in QM) GoogleMaps   ; Mudlo Gap , bottom site, 26°01’27”S, 152°13’57”E, 9.x–17.xii.1998 ( T85217 View Materials T85221 View Materials 5 specimens in QM) GoogleMaps   ; Mudlo Gap , bottom site, 26°01’27”S, 152°13’57”E, 25.i.1999 ( T85222 View Materials 1 specimen in QM) GoogleMaps   ; Mudlo Gap , bottom site, 26°01’27”S, 152°13’57”E, 25.i– 7.vi.1999 ( T85223 View Materials 1 specimen in QM) GoogleMaps   ; Oakview State Forest , barracks, 26°06’47”S, 152°19’36”E, 15–16.i.2002 (T122782 1 specimen in QM) GoogleMaps   ; Nangur State Forest , 1st site, 26°07’45”S, 151°58’15”E, 24.xi.1995 – 3.ii.1996 ( T78290 View Materials T85397 View Materials 8 specimens in QM) GoogleMaps   ; Nangur State Forest , 2nd site, 26°07’55”S, 151°58’43”E, 24.xi.1995 – 3.ii.1996 ( T78296 View Materials T85398 View Materials 3 specimens in QM) GoogleMaps   ; Jack Smith Environmental Park on Boat Mountain , 26°09’17”S, 151°54’34”E, 15.xii.1994 – 26.i.1995 ( T80226 View Materials T80227 View Materials 2 specimens in QM) GoogleMaps   ; Oakview State Forest , summit, 26°09’40”S, 152°19’43”E, 15.i–27.v.2002 (T111420–T111429 10 specimens in QM) GoogleMaps   ; Oakview State Forest , summit, 26°09’40”S, 152°19’43”E, 25.ix.2001 – 15.i.2002 (T116837–T116846 10 specimens in QM) GoogleMaps   ; Oakview State Forest , 26°08’32”S, 152°20’12”E, 12.xii.1997 – 26.iv.1998 ( T85311 View Materials T85315 View Materials 5 specimens in QM) GoogleMaps   ; Gympie , 11 km ESE, 26°14’24”S, 152°45’36”E, 1.i–28.ii.2002 (T111395 1 specimen in QM) GoogleMaps   ; Marys Creek, Gympie , 26°15’44”S, 152°32’40”E, 1974–1975 (T105313 1 specimen in QM) GoogleMaps   ; Cooran Tableland , BS68, 26°15’49”S, 152°49’43”E, 16.i– 27.v.2002 (T111401–T111406 6 specimens in QM) GoogleMaps   ; Cooran Tableland , 26°16’25”S, 152°50’05”E, 30.xi.1991 – 8.i.1992 ( T77869 View Materials T77911 View Materials 3 specimens in QM) GoogleMaps   ; Cooran Tableland , 26°16’25”S, 152°50’05”E, 8.i–2.iii.1992 ( T77912 View Materials T77952 View Materials 41 specimens in QM) GoogleMaps   ; Cooran Tableland , 26°16’25”S, 152°50’05”E, 2.iii–17.iv.1992 ( T77953 View Materials T77969 View Materials 17 specimens in QM) GoogleMaps   ; Cooran Barracks , 0.5 km N, 26°17’01”S, 152°49’57”E, 16.i–27.v.2002 (T111396– T111400 5 specimens in QM) GoogleMaps   ; Cooran Barracks , 0.5 km N, 26°17’01”S, 152°49’57”E, 25.ix.2001 – 16.i.2002 (T116859 1 specimen in QM) GoogleMaps   ; Cooran Plateau, 1300 feet via Gympie , [26°21’00”S, 152°47’00”E], 396 m, 3.x– 3.xi.1973, R.I. Storey, (CMNEN-00030167 specimens in CMNC) GoogleMaps   ; Cooran Plateau via Gympie , [26°21’00”S, 152°47’00”E], 1300 m, 10–11.iii.1973, R.I. Storey, (25-058447 20 specimens in QM, 25-058564 2 specimens in QDPC) GoogleMaps   ; Amamoor Creek , 26°23’55”S, 152°36’00”E, 15.i–27.v.2002 (T111410–T111417 8 specimens in QM) GoogleMaps   ; Amamoor Creek , 26°23’55”S, 152°36’00”E, 24.ix.2001 – 15.i.2002 (T116847–T116851 5 specimens in QM) GoogleMaps   ; Mount Kandanga , 26°27’07”S, 152°34’40”E, 15.i–27.v.2002 (T111418–T111419 2 specimens in QM) GoogleMaps   ; Mount Kandanga , summit, 26°27’07”S, 152°34’40”E, 24.ix.2001 – 15.i.2002 (T116852 specimens in QM) GoogleMaps   ; Dwyer Creek, Imbil , 26°27’53”S, 152°38’09”E, 15.i–27.v.2002 (T111407–T111409 3 specimens in QM) GoogleMaps   ; Dwyer Creek, Imbil , 26°27’53”S, 152°38’09”E, 24.ix.2001 – 15.i.2002 (T116871–T116876 6 specimens in QM) GoogleMaps   ; Perry’s Knob , 4 km NNE Rosewood, 27°36’11”S, 152°36’17”E, 11.xi.1998 – 13.i.1999 ( T85290 View Materials 1 specimen in QM) GoogleMaps   ; Perry’s Knob , 4 km NNE Rosewood, 27°36’11”S, 152°36’17”E, 13.i–16.v.1999 ( T85291 View Materials T85309 View Materials 19 specimens in QM) GoogleMaps   ; 1 mi N Nambour , [26°37’08”S, 152°58’00”E], 27.v–2.vi.1975, R.I. Storey (25-058452 7 specimens in QM) GoogleMaps   ; Little Yabba Creek , 26°37’22”S, 152°41’26”E, 2.iii–12.iv.1992 ( T77845 View Materials T77855 View Materials 11 specimens in QM) GoogleMaps   ; Little Yabba Creek , 26°37’22”S, 152°41’26”E, 7.i.1992 – 2.iii.1992 ( T77856 View Materials T77863 View Materials 8 specimens in QM) GoogleMaps   ; Little Yabba Creek , 26°37’22”S, 152°41’26”E, 30.xi.1991 – 7.i.1992 ( T77864 View Materials T77868 View Materials 5 specimens in QM) GoogleMaps   ; Mapleton Falls National Park , 26°37’49”S, 152°50’21”E, 8.i–03.iii.1992 ( T78298 View Materials T78316 View Materials 17 specimens in QM) GoogleMaps   ; Mapleton Falls National Park , 26°37’49”S, 152°50’21”E, 30.xi.1991 – 8.i.1992 ( T78317 View Materials 1 specimen in QM) GoogleMaps   ; Mapleton Falls National Park , 26°37’49”S, 152°50’21”E, 3.iii–17.iv.1992 ( T80131 View Materials T80147 View Materials 17 specimens in QM) GoogleMaps   ; Jimna , 26°38’37”S, 152°27’12”E, 29.xi.1991 – 7.i.1992 ( T78142 View Materials T78147 View Materials 6 specimens in QM) GoogleMaps   ; Jimna , 26°38’37”S, 152°27’12”E, 7.i– 02.iii.1992 ( T78148 View Materials T78156 View Materials 9 specimens in QM) GoogleMaps   ; Jimna , 26°38’37”S, 152°27’12”E, 02.iii–12.iv.1992 ( T78157 View Materials T78184 View Materials 28 specimens in QM) GoogleMaps   ; 20 km NW Maleny, Booloumba Creek , [26°38’44”S, 152°39’03”E], 150 m, 26– 29.iii.1993, C. Carlton, (CMNEN-00030170 1 specimen in CMNC) GoogleMaps   ; Tungi Creek, Jimna , 26°38’57”S, 152°28’21”E, 18.xii.1996 – 20.i.1997 ( T78185 View Materials T78188 View Materials 4 specimens in QM) GoogleMaps   ; Tungi Creek, Jimna , 26°38’57”S, 152°28’21”E, 20.i–9.iii.1997 ( T78189 View Materials 1 specimen in QM) GoogleMaps   ; Tungi Creek, Jimna , 26°38’57”S, 152°28’21”E, 9.iii–15.v.1997 ( T78190 View Materials T78196 View Materials 7 specimens in QM) GoogleMaps   ; Booloumba Creek , 26°39’09”S, 15802602 °38’32”E, 29.xi.1991 – 8.i.1992 ( T77757 View Materials T77778 View Materials 22 specimens in QM)   ; Booloumba Creek , 26°39’09”S, 152°38’32”E, 8.i–2.iii.1992 ( T77779 View Materials T77806 View Materials 28 specimens in QM) GoogleMaps   ; Booloumba Creek , 26°39’09”S, 152°38’32”E, 2.iii–12.iv.1992 ( T77807 View Materials T77826 View Materials 20 specimens in QM) GoogleMaps   ; Sunday Creek, Conondale Range 26°42’39”S, 152°33’49”E, 7.i–2.iii.1992 ( T80176 View Materials 1 specimen in QM) GoogleMaps   ; Sunday Creek, Conondale Range , 26°42’39”S, 152°33’49”E, 29.xi.1991 – 7.i.1992 ( T80177 View Materials 1 specimen in QM) GoogleMaps   ; Mary Cairncross Park , 26°46’52”S, 152°52’56”E, 29.xi.1991 – 8.i.1992 ( T80251 View Materials T80259 View Materials 9 specimens in QM) GoogleMaps   ; Mary Cairncross Park , 26°46’52”S, 152°52’56”E, 8.i–3.iii.1992 ( T80260 View Materials T80261 View Materials 2 specimens in QM) GoogleMaps   ; Mary Cairncross Park , 26°46’52”S, 152°52’56”E, 3.iii–12.iv.1992 ( T80262 View Materials T80263 View Materials 2 specimens in QM) GoogleMaps   ; Dularcha National Park , 26°47’41”S, 152°58’04”E, 1–3.iv.2004 (T122767 1 specimen in QM) GoogleMaps   ; Landsborough , 2.5 km NE, 26°47’31”S, 152°58’40”E, 25–27.iii.2004 (T122762–T122766 5 specimens in QM) GoogleMaps   ; Yarraman , 2.2 km SE, 26°51’35”S, 151°59’36”E, 14–15.i.1998 ( T78203 View Materials 1 specimen in QM) GoogleMaps   ; Yarraman , 26°51’38”S, 152°00’34”E, 2.iii– 12.iv.1992 ( T78246 View Materials T78248 View Materials 3 specimens in QM) GoogleMaps   ; Yarraman , 26°51’38”S, 152°00’34”E, 7.i–2.iii.1992 ( T78249 View Materials 1 specimen in QM) GoogleMaps   ; Saddle Tree Creek , 26°51’37”S, 151°38’06”E, 1.iii–12.iv.1992 ( T80248 View Materials T80250 View Materials 3 specimens in QM) GoogleMaps   ; Mellum Creek Bridge, 26°51’51”S, 153°01’02”E, 22.ii–14.iii.2008 (T156697 1 specimen in QM) GoogleMaps   ; Saddle Tree Creek, Bunya Mountains , 26°52’09”S, 151°37’26”E, 1.i.1975 – 31.xii.1976 (T105312 1 specimen in QM) GoogleMaps   ; Top of Blackbutt Range , 26°52’38”S, 152°11’32”E, 24.xi.1995 – 3.ii.1996 (T116877, T85400 View Materials 2 specimens in QM) GoogleMaps   ; Blackbutt Range, 6 mi E of Blackbutt, [26°53’00”S, 152°11’00”E], 22–23.xii.1973, R.I. Storey, (25-058451 1 specimen in QM) GoogleMaps   ; Neurum Creek Conservation Park, 26°58’20”S, 152°40’54”E, 25–27.iii.2004 (T122761 1 specimen in QM) GoogleMaps   ; Brisbane, Forest Park, Scrub Creek Road, [27°01’00”S 152°26’00”E], 18.ix–29.x.1998, P. Bouchard, (25-058458 17 specimens in QM) GoogleMaps   ; Mount Glorious , [27°20’00”S, 152°46’00”E], 630 m, 14.xi.1986 – 30.i.1987, T. Hiller, (25-057981 5 specimens in ANIC) GoogleMaps   ; Mount Glorious , [27°20’00”S, 152°46’00”E], 13–20.iv.1975, R.I. Storey, (25-058449 1 specimen in QM) GoogleMaps   ; Mount Glorious , [27°20’00”S, 152°46’00”E], 10.x.1999, P. Bouchard, K. Evans, & J. Skevington (25-058464 4 specimens in QM) GoogleMaps   ; Mount Delaney , 0.9 km SSW, 27°01’14”S, 152°41’58”E, 10–11.ii.2001 (T105679 1 specimen in QM) GoogleMaps   ; Mount Mee , 27°02’53”S, 152°40’49”E, 29.xi.1991 – 8.i.1992 ( T80148 View Materials T80153 View Materials 6 specimens in QM) GoogleMaps   ; Mount Mee , 27°02’53”S, 152°40’49”E, 3.iii–12.iv.1992 ( T80154 View Materials T80164 View Materials 11 specimens in QM) GoogleMaps   ; Mount Mee , 27°02’53”S, 152°40’49”E, 8.i–3.iii.1992 ( T80165 View Materials T80175 View Materials 11 specimens in QM) GoogleMaps   ; Mount Deongwar , 2 km SE, 27°12’53”S, 152°16’45”E, 14.x–31.xii.1998 ( T85322 View Materials 1 specimen in QM) GoogleMaps   ; Mount Deongwar, 3 km S, Site 2, 27°13’49”S, 152°15’25”E, 30.xii.1998 – 26.iii.1999 ( T85316 View Materials T85321 View Materials 5 specimens in QM) GoogleMaps   ; Mount Glorious , barracks, 27°19’54”S, 152°45’12”E, 16–28.iv.2007 (T148364–T148365 2 specimens in QM) GoogleMaps   ; Mount Glorious , 27°20’20”S, 152°45’30”E, 1.i–31.iii.1982 ( T78255 View Materials T78256 View Materials 2 specimens in QM) GoogleMaps   ; Stony Creek, via Samford , 27°20’20”S, 152°47’52”E, 22.x.1994 – 2.ii.1995 ( T78215 View Materials T78232 View Materials 18 specimens in QM) GoogleMaps   ; Stony Creek, via Samford , 27°20’20”S, 152°47’52”E, 2.ii–8.iv.1995 ( T78233 View Materials T78236, T87587 5 specimens in QM) GoogleMaps   ; Cashs Crossing , [27°21’00”S, 152°58’00”E], 16.iii.1959, A. Diatloff, (25-058457 1 specimen in QM) GoogleMaps   ; Ravensbourne National Park , 27°22’09”S, 152°10’44”E, 1.iii–11.iv.1992 ( T77829 View Materials T77835 View Materials 7 specimens in QM) GoogleMaps   ; Ravensbourne National Park , 27°22’09”S, 152°10’44”E, 7.i–1.iii.1992 ( T77836 View Materials T77844 View Materials 9 specimens in QM) GoogleMaps   ; Boombana National Park , site 1, 27°24’08”S, 152°47’22”E, 15–16.ii.2004 (T122060–T122062 3 specimens in QM) GoogleMaps   ; Enoggera Reservoir , site 8, 27°26’37”S, 152°54’14”E, 15.iii–18.v.2000 (T105338 1 specimen in QM) GoogleMaps   ; Enoggera Reservoir , site 2, 27°26’28”S, 152°55’16”E, 27.i–15.iii.2000 (T105339 1 specimen in QM) GoogleMaps   ; Enoggera Reservoir , site 2, 27°26’28”S, 152°55’16”E, 16.x–21.xii.1999 (T105322–T105323 2 specimens in QM) GoogleMaps   ; Enoggera Reservoir , site 3, 27°26’44”S, 152°55’19”E, 16.x–21.xii.1999 (T105324–T105326 3 specimens in QM) GoogleMaps   ; Enoggera Reservoir , site 3, 27°26’44”S, 152°55’19”E, 27.i–15.iii.2000 (T105331–T105332 2 specimens in QM) GoogleMaps   ; Enoggera Reservoir , site 3, 27°26’44”S, 152°55’19”E, 10.xi.1999 (T118119–T118120 2 specimens in QM) GoogleMaps   ; Enoggera Reservoir , site 5, 27°26’44”S, 152°55’19”E, 16.x–21.xii.1999 (T105327–T105328 2 specimens in QM) GoogleMaps   ; Enoggera Reservoir , site 5, 27°26’44”S, 152°55’19”E, 21.xii.1999 – 27.i.2000 (T105329 1 specimen in QM) GoogleMaps   ; Enoggera Reservoir , site 5, 27°26’44”S, 152°55’19”E, 15.iii–18.v.2000 (T105337 1 specimen in QM) GoogleMaps   ; Enoggera Reservoir , site 1, 27°26’31”S, 152°55’27”E, 27.i–15.iii.2000 (T105330 1 specimen in QM) GoogleMaps   ; Enoggera Reservoir , site 1, 27°26’31”S, 152°55’27”E, 15.iii–18.v.2000 (T105336 1 specimen in QM) GoogleMaps   ; Jones Road, Brookfield , 27°28’08”S, 152°54’40”E, 12–13.ii.2011, G.B. Monteith, (25-053874–25-05376 3 specimens also as COL1673, COL1674, COL 1675 specimens in ANIC) GoogleMaps   ; Upper Brookfield Road , 27°28’21”S, 152°50’06”E, 12–13.ii.2011, G.B. Monteith, (25-053873, also as COL71672 View Materials 1 specimen in ANIC) GoogleMaps   ; Slaughter Falls , 27°28’44”S, 152°57’34”E, 6–7.ii.2001 (T105680–T105684 5 specimens in QM) GoogleMaps   ; Upper Brookfield Road, Brisbane , [27°29’00”S, 152°52’00”E], 22.ix–4.x.1975, R.I. Storey, (25-058459 3 specimens in QM) GoogleMaps   ; Upper Brookfield Road, Brisbane , [27°29’00”S, 152°52’00”E], 24.viii–7.ix.1975, R.I. Storey, (25-058460 7 specimens in QM) GoogleMaps   ; Mount Coot–tha , [27°29’00”S, 152°57’00”E], 15–20.iii.1971, G.B. Monteith, (25-058456 1 specimen in QM) GoogleMaps   ; Moggill Creek Brisbane, 27°30’25”S, 152°55’52”E, 22–23.i.2011, G.B. Monteith, (25-060829 1 specimen in ANIC) GoogleMaps   ; The Knobby , 4 km W Glamorgan Vale, 27°30’27”S, 152°35’18”E, 13.i–16.v.1999 ( T85310 View Materials 1 specimen in QM) GoogleMaps   ; Cameron’s Scrub , 27°30’32”S, 152°43’48”E, 75 m, 30.iv–1.v.2012, G.B. Monteith, (25-057576 2 specimens in ANIC) GoogleMaps   ; Cameron’s Scrub , top ridge, 27°30’29”S, 152°43’54”E, 13.i–16.v.1999 ( T85231 View Materials T85234 View Materials 4 specimens in QM) GoogleMaps   ; Cameron’s Scrub , gully, 27°30’35”S, 152°43’54”E, 13.i–16.v.1999 ( T85235 View Materials T85259 View Materials 25 specimens in QM) GoogleMaps   ; Cameron’s Scrub , gully, 27°30’35”S, 152°43’54”E, 11.xi.1998 – 13.i.1999 ( T85260 View Materials T85286 View Materials 27 specimens in QM) GoogleMaps   ; Cameron’s Scrub, Snig track, 27°30’49”S, 152°43’43”E, 13.i–16.v.1999 ( T85225 View Materials T85230 View Materials 6 specimens in QM) GoogleMaps   ; Cameron’s Scrub, Snig track, 27°30’49”S, 152°43’43”E, 11.xi.1998 – 13.i.1999 ( T85287 View Materials T85289 View Materials 3 specimens in QM) GoogleMaps   ; Belmont Hill , top, 27°30’48”S, 153°06’59”E, 12.xii.1997 – 7.v.1998 ( T78197 View Materials T78199, T89630 T89646 View Materials 20 specimens in QM) GoogleMaps   ; Belmont Hills Bushlands , 27°30’47”S, 153°07’05”E, 25–27.ii.2004 (T122063 1 specimen in QM) GoogleMaps   ; Belmont Hill , gully, 27°30’44”S, 153°07’06”E, 12.xii.1997 – 7.v.1998 ( T85325 View Materials - T85338 View Materials 14 specimens in QM) GoogleMaps   ; Pine Mountain (Flinton Hill in QM)   , 27°31’23”S, 152°44’45”E, 1975–1976 ( T78257 View Materials T78258 View Materials 2 specimens in QM) GoogleMaps   ; Redwood Park, Toowoomba , [27°35’00”S 151°59’00”E], 8.xi.1988, E.S. Nielsen & M. Horak, (25-057983 1 specimen in ANIC) GoogleMaps   ; Buhot Creek, Burbank , 27°35’27”S, 153°10’19”E, 6.xi.2003 (T122059 1 specimen in QM) GoogleMaps   ; Mount Cotton , 27°36’58”S, 153°12’21”E, 3.ix–12.xii.1997 ( T78205 View Materials T78209 View Materials 5 specimens in QM) GoogleMaps   ; Mount Cotton, Scott’s Dam , 27°36’58”S, 153°12’21”E, 12.xii.1997 – 7.v.1998 ( T85374 View Materials T85381 View Materials 8 specimens in QM) GoogleMaps   ; Mount Cotton , upper gully, 27°36’58”S, 153°12’32”E, 12.xii.1997 – 7.v.1998 ( T78200 View Materials T78202, T85356 –T785373 21 specimens in QM) GoogleMaps   ; Perry’s Knob , 27°36’00”S, 152°36E, 13.i–16.v.1999 ( T98837 View Materials 1 specimen in QM) GoogleMaps   ; Bahr’s Scrub , 27°45’31”S, 153°10’06”E, 21.i–9.iii.1992 ( T77622 View Materials T77647 View Materials 26 specimens in QM) GoogleMaps   ; Bahr’s Scrub , 27°45’31”S, 153°10’06”E, 10.xii.1991 – 21.i.1992 ( T77648 View Materials T77670 View Materials 23 specimens in QM) GoogleMaps   ; Bahr’s Scrub , 27°45’31”S, 153°10’06”E, 9.iii–30.iv.1992 ( T77671 View Materials T77682 View Materials 12 specimens in QM) GoogleMaps   ; Bahr’s Scrub , 27°45’31”S, 153°10’06”E, 10.xii.1991 – 21.i.1992 ( T77706 View Materials T77726 View Materials 27 specimens in QM) GoogleMaps   ; Bahr’s Scrub , 27°45’31”S, 153°10’06”E, 21.i–9.iii.1992 ( T77727 View Materials T77729 View Materials 3 specimens in QM) GoogleMaps   ; Mount Tamborine National Park , [27°53’11”S, 153°10’51”E], 24.vi–6.vii.1978, S. Peck & J. Peck (CMNEN- 00030168 1 specimen in CMNC) GoogleMaps   ; Tamborine Mountain (general), [27°55’00”S, 153°10’00”E], 7.xii.1967, E. Matthews, (25-057973 1 specimen in ANIC) GoogleMaps   ; Tamborine Mountain (general), [27°55’00”S, 153°10’00”E], 26.xii.1972, R.I. Storey, (25-058455 1 specimen in QM) GoogleMaps   ; Palm Grove, Tamborine [Tamborine Mountain], [27°55’00”S, 153°10’00”E], 28.ix–12.x.1975, R.I. Storey, (25-058461 2 specimens in QM) GoogleMaps   ; North Tamborine , 27°55’13”S, 153°11’50”E, 9.iii–25.iv.1992 (T105311 1 specimen in QM) GoogleMaps   ; North Tamborine , 27°55’13”S, 153°11’50”E, 11.i– 9.iii.1992 ( T78259 View Materials T78277 View Materials 19 specimens in QM) GoogleMaps   ; North Tamborine , 27°55’13”S, 153°11’50”E, 9.iii–25.iv.1992 ( T78278 View Materials T78289 View Materials 12 specimens in QM) GoogleMaps   ; Tamborine, Guanaba shelf, 27°55’19”S, 153°13’14”E, 1.xii.1992 – 31.i.1993 ( T80345 View Materials T80346 View Materials 2 specimens in QM) GoogleMaps   ; Mount French National Park , 28°00’07”S, 152°37’00”E, 2.xii.1991 – 6.i.1992 ( T80237 View Materials T80240 View Materials 4 specimens in QM) GoogleMaps   ; Mount French National Park , 28°00’07”S, 152°37’00”E, 6.i–29.ii.1992 ( T80241 View Materials T80244 View Materials 4 specimens in QM) GoogleMaps   ; Mount French National Park , 28°00’07”S, 152°37’00”E, 29.ii–11.iv.1992 ( T80245 View Materials T80247 View Materials 3 specimens in QM) GoogleMaps   ; Glen Witheren , main scrub, 28°02’48”S, 153°07’12”E, 30.xii.1991 – 24.v.1992 ( T80187 View Materials T80195 View Materials 9 specimens in QM) GoogleMaps   ; Glen Witheren , main scrub, 28°02’48”S, 153°07’12”E, 1.ii– 30.iv.1991 ( T80209 View Materials T80210 View Materials 2 specimens in QM) GoogleMaps   ; Glen Witheren , shelf scrub, 28°02’58”S, 153°06’58”E, 30.x– 30.xii.1991 ( T80196 View Materials T80208 View Materials 13 specimens in QM) GoogleMaps   ; Burleigh Headland National Park , 28°05’45”S, 153°27’25”E, 1.i–4.iii.1992 ( T77683 View Materials T77695, T77741 T77742 View Materials 15 specimens in QM) GoogleMaps   ; Burleigh Headland National Park , 28°05’45”S, 153°27’25”E, 9.xii.1991 – 1.i.1992 ( T77696 View Materials T77705 View Materials 10 specimens in QM) GoogleMaps   ; Burleigh Headland National Park , 28°05’45”S, 153°27’25”E, 4.iii–2.iv.1992 ( T77743 View Materials T77756 View Materials 14 specimens in QM) GoogleMaps   ; Black Rock Scrub , 28°07’32”S, 152°39’48”E, 2.xii.2000 – 13.v.2001 (T105663–T105672, T106763–T106764 12 specimens in QM) GoogleMaps   ; Black Rock Scrub , 28°07’32”S, 152°39’48”E, 6.x–3.xii.2000 (T105686–T105693 8 specimens in QM) GoogleMaps   ; Black Rock Scrub , 28°07’32”S, 152°39’48”E, 2.xii.2000 – 13.v.2001 (T105341–T105342 2 specimens in QM) GoogleMaps   ; Black Rock Scrub , 28°07’32”S, 152°39’48”E, 6.x–3.xii.2000 (T105333–T105334 2 specimens in QM) GoogleMaps   ; Environment   Centre, Numinbah Valley , 28°07’24”S, 153°14’10”E, 1978–1979 (T105314 1 specimen in QM) GoogleMaps   ; Bonogin Road , 0.5 km W Smith Road, 28°07’50”S, 153°21’11”E, 3–7.iii.2008 (T156691–T156692 2 specimens in QM) GoogleMaps   ; Smith Road , gully rainforest, 28°08’47”S, 153°21’36”E, 3–7.iii.2008 (T156693–T156696 4 specimens in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–300–D, 28°08’31.2”S, 153°07’58.8”E, 13–23.i.2007 (T145225 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–300–D, 28°08’31.2”S, 153°07’58.8”E, 16–26.x.2006 (T145229, T145234 2 specimens in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–300–D, 28°08’31.2”S, 153°07’58.8”E, 21–26.x.2006 (T145238 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–300–D, 28°08’31.2”S, 153°07’58.8”E, 18–23.i.2007 (T145243 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–300–A, 28°08’52.8”S, 153°08’13.2”E, 13–23.i.2007 (T145222 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–300–A, 28°08’52.8”S, 153°08’13.2”E, 16–26.x.2006 (T145232 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–300–C, 28°09’3.6”S, 153°08’16.8”E, 13–23.i.2007 (T145224 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–300–C, 28°09’3.6”S, 153°08’16.8”E, 16–26.x.2006 (T145231 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–300–C, 28°09’3.6”S, 153°08’16.8”E, 13–18.i.2007 (T145240 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–300–C, 28°09’3.6”S, 153°08’16.8”E, 18–23.i.2007 (T145242 1 specimen in QM) GoogleMaps   ; Beechmont Road , 28°09’22”S, 153°11’42”E, 580 m, 29.ii.2012 –iii, 2012, G.B. Monteith, (25-053910 1 specimen in ANIC) GoogleMaps   ; Springbrook , north end, 28°09’47”S, 153°15’42”E, 30.viii–31.x.1997 ( T78204 View Materials 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–300–B, 28°09’18”S, 153°08’20.4”E 13–23.i.2007 (T145223 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–300–B, 28°09’18”S, 153°08’20.4”E 16–26.x.2006 (T145230–T145235 6 specimens in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–300–B, 28°09’18”S, 153°08’20.4”E 21–26.x.2006 (T145237 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–300–B, 28°09’18”S, 153°08’20.4”E 13–18.i.2007 (T145239 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–300–B, 28°09’18”S, 153°08’20.4”E, 18–23.i.2007 (T145241 1 specimen in QM) GoogleMaps   ; Gwinganna , 6 km SW by S of Tallebudgera, 28°11’00”S, 153°23’00”E, 18–23.iv.1994, D. Rentz, W. Lee & M. Upton, (25-057982 1 specimen in ANIC) GoogleMaps   ; Nichol’s Scrub , 28°11’26”S, 153°25’33”E, 1.i–4.iii.1992 ( T78111 View Materials T78123 View Materials 13 specimens in QM) GoogleMaps   ; Nichol’s Scrub , 28°11’26”S, 153°25’33”E, 4.iii–29.iv.1992 ( T78124 View Materials T78134 View Materials 11 specimens in QM) GoogleMaps   ; Nichol’s Scrub , 28°11’26”S, 153°25’33”E, 9.xii.1991 – 1.i.1992 ( T78135 View Materials T78141 View Materials 7 specimens in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–500–C, 28°12’36”S, 153°08’20.4”E, 22.iii–1.iv.2007 (T108394 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–500–C, 28°12’36”S, 153°08’20.4”E, 15–25.i.2007 (T145228 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–700–D, 28°12’14.4”S, 153°07’44.4”E, 14–24.i.2007 (T145226 1 specimen in QM) GoogleMaps   ; Lower Coomera , 28°12’20”S, 153°10’46”E, 9.i.1995 – 6.iv.1995 (T126027–T126029, T80219 View Materials T80223 View Materials 5 specimens in QM) GoogleMaps   ; Lower Coomera , 28°12’20”S, 153°10’46”E, 3.xii.1994 – 9.i.1995 ( T80215 View Materials T80218, T80344 5 specimens in QM) GoogleMaps   ; Tullawallal, Binna Burra , 28°12’36”S, 153°11’22”E, 3.xii.1994 – 9.i.1995 ( T80224 View Materials T80225 View Materials 2 specimens in QM) GoogleMaps   ; Mount Gannon , 1.5 km SE, 28°12’39”S, 153°19’27”E, 9–12.ii.2006 (T135162–T135166 5 specimens in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–500–B, 28°12’43.2”S, 153°08’27.6”E, 15–25.i.2007 (T145227 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–500–B, 28°12’43.2”S, 153°08’27.6”E, 18–28.x.2006 (T145233 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–500–B, 28°12’43.2”S, 153°08’27.6”E, 18–23.x.2006 (T145236 1 specimen in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–500–B, 28°12’43.2”S, 153°08’27.6”E, 20–25.i.2007 (T145244 1 specimen in QM) GoogleMaps   ; Mount Tallebudgera , 0.4 km NE, 28°12’55”S, 153°21’35”E, 31.i–3.ii.1997 ( T85388 View Materials T85395 View Materials 8 specimens in QM) GoogleMaps   ; Orams Dump , 28°13’24”S, 153°18’59”E, 26.i–2.ii.2006 (T133766–T133767 2 specimens in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–900–D, 28°13’37.2”S, 153°07’51.6”E, 20.i.2008 (T155890 1 specimen in QM) GoogleMaps   ; Mount Cougal , W Peak, 1 km NNW, 28°13’42”S, 153°18’34”E, 26.i–2.ii.2006 (T133768 1 specimen in QM) GoogleMaps   ; Tallebudgera Valley , falls area, 28°13’40”S, 153°18’46”E, 19.x.2003 (T124780 1 specimen in QM) GoogleMaps   ; Tallebudgera Valley , falls area, 28°13’40”S, 153°18’46”E, 1.v.2005 (T132352 1 specimen in QM) GoogleMaps   ; Tallebudgera Valley , falls area, 28°13’40”S, 153°18’46”E, 29.x.2004 (T132353 1 specimen in QM) GoogleMaps   ; Tallebudgera Valley , falls area, 28°13’40”S, 153°18’46”E, 26.ix.1998 – 21.v.2000 (T116858 1 specimen in QM) GoogleMaps   ; Tallebudgera Valley , falls area, 28°13’40”S, 153°18’46”E, 11.xi.1997 – 28.i.1998 ( T85383 View Materials T85387 View Materials 4 specimens in QM) GoogleMaps   ; Bilbrough Lookout , 1.3 km ENE, 28°13’47”S, 153°17’59”E, 4– 11.ii.2006 (T135160–T135161 2 specimens in QM) GoogleMaps   ; Springbrook Track , 300 m, 28°13’36”S, 153°18’30”E, 25–26.xi.2001 (T116836 1 specimen in QM) GoogleMaps   ; Tallebudgera Valley , 420 m, 28°13’57”S, 153°18’13”E, 4–28.iv.1997 ( T78075 View Materials 1 specimen in QM) GoogleMaps   ; Numinbah Arch , 28°13’51”S, 153°14’37”E, 1.i–29.ii.1992 ( T77598 View Materials T77621, T77737 T77740 View Materials 28 specimens in QM) GoogleMaps   ; Numinbah Arch , 28°13’51”S, 153°14’37”E, 29.ii–11.iv.1992 ( T77730 View Materials T77735, T77827 T77828 View Materials 8 specimens in QM) GoogleMaps   ; Upper Tallebudgera 330 m, 28°14’00”S, 153°18’32”E, 2.i–17.ii.1995 ( T78250 View Materials T78254 View Materials 5 specimens in QM) GoogleMaps   ; Tallebudgera Valley , 28°14’07”S, 153°17’59”E, 28.xii.1991 – 6.iii.1992 ( T65288 View Materials T65291 View Materials 4 specimens in QM) GoogleMaps   ; Tallebudgera Valley , 28°14’07”S, 153°17’59”E, 6.iii–28.iv.1992 ( T65292 View Materials T65296 View Materials 5 specimens in QM) GoogleMaps   ; Tallebudgera Valley , 28°14’07”S, 153°17’59”E, 18.xi–28.xii.1991 ( T65297 View Materials T65298 View Materials 2 specimens in QM) GoogleMaps   ; Lamington National Park, IBISCA Site IQ–900–A, 28°14’2.4”S, 153°08’27.6”E, 21–26.i.2007 (T145245 1 specimen in QM) GoogleMaps   ; Numinbah Valley Natural Arch National Park ., [28°14’19”S, 153°14’17”E], 300 m, 7.vii.1978, S. Peck & J. Peck (CMNEN-00030169 1 specimen in CMNC) GoogleMaps   ; Levers Plateau , 28°18’57”S, 152°50’53”E, 6.i–29.ii.1992 ( T65299 View Materials T65312 View Materials 14 specimens in QM) GoogleMaps   ; Levers Plateau , 28°18’57”S, 152°50’53”E, 2.xii.1991 – 6.i.1992 ( T65313 View Materials T65316 View Materials 4 specimens in QM) GoogleMaps   ; Levers Plateau , 28°18’57”S, 152°50’53”E, 29.ii–11.iv.1992 ( T65317 View Materials T65323 View Materials 7 specimens in QM) GoogleMaps   ; Levers Plateau via Rathdowney , [28°20’00”S, 152°52’00”E], 640 m, 1.xi.1975, I.D. Naumann, (25-058465 1 specimen in QM) GoogleMaps   ; Boyds Butte , 2 km ESE,, 28°14’21”S, 153°21’17”E 13.vii.2005 (T132354–T132355 2 specimens in QM) GoogleMaps   ; New South Wales: Wiangaree State Forest Isaksson Ridge, [28°23’00”S, 153°06’00”E], 1050 m, 29.ii–3.iii.1980, A. Newton & M. Thayer, (25-057978 2 specimens in ANIC) GoogleMaps   ; Wiangaree State Forest, Sheepstation Creek , [28°23’00”S, 153°06’00”E], 600 m, 29.ii–3.iii.1980, A. Newton & M. Thayer, (25-057975–7 3 specimens in ANIC) GoogleMaps   ; Unumgar State Forest near Woodenbong, Pole Bridge road 788, [28°24’00”S, 152°40’00”E], 430 m, 2–11.i.1987, A. Newton & M. Thayer, (25-058330 8 specimens in ANIC) GoogleMaps   ; Byrrill Creek , campground, 28°26’55”S, 153°11’35”E, 30.i–1.ii.2008 (T154644 1 specimen in QM) GoogleMaps   ; Byrrill Creek , lower creek, 28°26’51”S, 153°11’39”E, 30.i–1.ii.2008 (T154642 1 specimen in QM) GoogleMaps   ; Beaury National Forest, Koreelah Range 1.8 km from Tooloom Lookout along Tucker box Road , 28°28’23”S, 152°24’04”E, 23.iii–9.v.1999, S. Lassar & C. Lemann, (25-060773 also as COL 753 specimens in AM) GoogleMaps   ; Tooloom Plateau , 14 km W Urbenville, [28°29’00”S, 152°24’00”E], 15–21.ii.1984, I.D. Naumann, (25-057980 1 specimen in ANIC) GoogleMaps   ; Whian Whian State Forest , gate, 28°33’29”S, 153°20’09”E, 29–31.i.2008 (T154636 1 specimen in QM) GoogleMaps   ; The Channon , 8 km NNE, 28°36’08”S, 153°17’48”E, 30.i–2.ii.2008 (T154643 1 specimen in QM) GoogleMaps   ; Minyon Falls , 3.5 km S, 28°38’27”S, 153°21’39”E, 29–31.i.2008 (T154637 1 specimen in QM) GoogleMaps   ; Boat harbour, site 2, 28°46’42”S, 153°19’45”E, 29.i– 31.i.2008 (T154638 1 specimen in QM) GoogleMaps   ; Lion’s Park, via Boatharbour , [28°47’00”S, 153°20’00”E], 19–20.i.1974, R.I. Storey, (25-058454 1 specimen in QM) GoogleMaps   ; Rotary Park, Lismore , site 2, 28°48’41”S, 153°17’46”E, 29–31.i.2008 (T154641 1 specimen in QM) GoogleMaps   ; Wilson Nature Reserve Lismore, [28°49’00”S, 153°16’00”E], 17.xii.1998, G. Williams & B. Williams, (25-058111 1 specimen in GWC) GoogleMaps   ; Wilson Park, Lismore , 28°50’02”S, 153°17’25”E, 29– 31.i.2008 (T154639 1 specimen in QM) GoogleMaps   ; Davis Scrub , site 2, 28°52’02”S, 153°24’20”E, 29–31.i.2008 (T154640 1 specimen in QM) GoogleMaps   ; Victoria Park, via Alstonville , [28°54’00”S, 153°24’00”E], 16.xi.1974, G.B. Monteith, (25- 058463 1 specimen in QM) GoogleMaps   ; Victoria Park near Alstonville [28°54’00”S, 153°24’00”E], 6.ii.1980, (1 specimen in AM) GoogleMaps   ; Huonbrook [28°32’00”S, 152°21’00”E], 2.iii.1964, (2 specimens in AM) GoogleMaps   ; Yabbra Scrub, Yabbra State Forest [28°31’00”S, 152°34’00”E], 14.xii.1988 (1 specimen in AM) GoogleMaps   ; Lorien, Lansdowne , [31°52’00”S, 152°29’00”E], 7.ii.1981, G. Williams & T. Williams, (25-058112 1 specimen in GWC) GoogleMaps   ; Lorien Wildlife Refuge 3km N Lansdowne, 31°45’00”S, 152°32’00”E, 16.ix.2010, G. Williams & T. Williams, (25-060880 also as COL1145 1 specimen in ANIC) GoogleMaps   ; 4 km W. Lansdowne, Lorien , [31°45’00”S, 152°32’00”E], 4.i.1987, H. Howden & A. Howden (CMNEN- 00030171 1 specimen in CMNC) GoogleMaps   ; Newbys Lane, Lansdowne , [31°47’00”S, 152°32’00”E], 14.i.1981, G. Williams & T. Williams, (25-057979 3 specimens in ANIC) GoogleMaps   ; Swans Crossing, Kerewong State Forest , 12 km WNW of Kendall, 31°36”S, 152°34”E, 24.x.1993 (1 specimen in GWC)   .

Description. Lectotype predominately black, humeri and apical edges of elytra reddish brown. Most other specimens also usually have red elytral humeri. Apical antennomere (3 rd) of antennal clubs white, basal 2 antennomeres (1 st, 2 nd) grey.

Total length: 5.0– 6.5 mm. Lectotype measurements (taken from photograph): total length 6.5 mm, elytral width 4.35 mm.

Male. Head: Width to length ratio 45: 32. Surface very finely reticulate with fine punctation. Dorsal part of eyes wide, separated by an interocular space approximately 8 times eye width (65: 8); eye canthus not dividing eye.

Prothorax: Pronotum with posterolateral angles distinct, lateral margins bordered. Surface finely punctate and finely reticulate in anterior half, finely punctate and nitid in posterior half. Pronotum width to length ratio 78: 47. Hypomeral surface finely reticulate, hypomeral stria absent.

Elytra: Surface reticulate. All striae on disc equal width, impunctate. Striae 6 and 7 not extending to the elytral base. 8th stria sometimes with a few punctures where it curves inwards posteriorly. Ratio of length of the elytra along suture to maximum elytral width 85: 100.

Legs: Protibia with ventral ridge not crenulate. Protibia with 3 teeth on the outer edge. Front edge truncate, apical digit broad and blunt apically, and apical spur absent. Mesotibia with a brush of setae apically on inner edge. Metatibia slightly bent with variously crenulated inner edge, and no round expansion apically. Metafemur slightly angulate basally on inner edge. Basal metatarsomere lobed on inner edge.

Abdomen: Pygidium with a transverse sinuate fold across base that is curved and impressed medially. Pygidial disc with a large flat, reticulate central area that occupies more than half the disc not extending under the transverse sinuate fold and separated from it by an impressed line. Ventrite 6 very finely punctate.

Pterothorax: Mesometaventral suture arched. Medial lobe of metaventrite very finely reticulate and virtually impunctate, broadly margined between mesocoxae. Lateral lobe of metaventrite very finely reticulate, finely punctate and setose. Mesoventrite very finely reticulate and notched anteriorly. Mesepimeron very finely reticulate. Metanepisternum reticulate.

Female. Protibia with front edge truncate and apical spur arising directly from front edge, apical digit absent ( Fig. 2B View FIGURE 2 ). Inner edge of metatibia not crenulate and inner edge of metafemur not angulate ( Fig. 2E View FIGURE 2 ). Mesotibia without a brush of setae apically.

Distribution. Known from rainforests, wet and dry sclerophyll forest from south of Gladstone, Queensland to Lansdowne, New South Wales ( Fig. 6A View FIGURE 6 ). It has been recorded from the following Interim Biogeographic Regionalisations for Australia (IBRA) bioregions: South Eastern Queensland (SEQ), the most northerly extents of New South Wales North Coast (NNC) and New England Tablelands (NET), Sydney Basin (SYB), the most easterly extents of Brigalow Belt South (BBS). Notably, there is a large gap in the distribution that extends across most of the NNC bioregion, separating records north of 29°S from those south of 31°36”S. BIOCLIM modelling suggests that most of the NNC bioregion, except for a small gap at ~ 30°S, has suitable climatic conditions for the species ( Fig. 6B View FIGURE 6 ). The region has been well sampled and the gap in the distribution likely reflects true absence.

Comments. Lepanus ustulatus   is a generalist species, collected in traps with a variety of baits including macropod or human dung, mushroom, and carrion (chicken or squid). Also collected at light, in berlesed leaf litter, flight intercept and window traps. In a food preference study that tested attraction to 10 different baits including a variety of dung, carrion, fermenting fruits and mushroom lures, seven specimens of L. ustulatus   were collected including six at domestic fowl dung and one specimen at vertebrate carrion ( Ebert et al. 2019).

In southern Queensland and northern New South Wales, L. ustulatus   appears to be part of a mimicry ring involving several species of scarabs and hydrophilids that are attracted to dung and other decaying organic resources ( Fikacek et al. 2014). The species listed in the potential mimicry complex also includes Onthophagus arrilla Matthews 1974   ( Coleoptera   : Scarabaeidae   : Scarabaeinae   : Onthophagini), O. sydneyensis ( Lansberge, 1874)   , Liparochrus nanus Paulian, 1980   ( Hybosoridae   ), and Austrotypus nothofagi Fikacek, Minoshima & Newton, 2014   ( Hydrophilidae   ) and are all superficially similar, most notably with distinct reddish-brown elytral humeri on the otherwise black body. Other recently described Lepanus   that occur in the region, L. burnetti Gunter & Weir, 2019   and L. meierae   , may also be part of this mimcry ring. Beyond the region, a number of other Lepanus   species also have this colour pattern including most species in the L. ustulatus   species group, although the reddish-brown humeri are less defined than in L. ustulatus   .

Lepanus ustulatus   is one of only seven species in the genus that has been assessed for the IUCN redlist and was classified as a species of least concern due to its wide distribution and expected lack of significant threats to its populations ( Weir 2013). This broad distribution of L. ustulatus   from Gladstone to Landsdowne, especially the absence of records in much of the New South Wales North Coast Bioregion, is interesting to consider. There are no consistent morphological differences that support splitting the northern and southern populations of this species and sequence data from specimens from SYB and SEQ IBRA bioregions is congruent with expected levels of intraspecific variation. The disjunction within the distribution could represent an extinction of populations in the New South Wales North Coast bioregion.

Matthews (1974) assigned a lectotype for L. ustulatus   from the series of MNHN specimens, formerly of the van Lansberge collection, and selected a male labelled “Wide Bay”. Lepanus ustulatus   is the only species within this species group that occurs in southern Queensland including the region around Wide Bay. Specimens listed in Matthews (1974) from Paluma (Wet Tropics), Eungella and near Mackay (central Queensland) represent the morphologically similar and cryptic species, Lepanus lemannae   described below. The two specimens of L. ustulatus   reported by Aristophanous (2014) from the Atherton region were not examined but based on geographic range of these superficially similar species they are probably Lepanus lemannae   .

Dorsal, lateral, ventral views, a close up of the pygidium and all labels of the lectotype of L. ustulatus   , and the dorsal habitus, pygidia, and label images of the four additional paralectotypes are published on the Muséum national d’Histoire naturelle, Paris ( France) online database.

http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec8675;

http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec8676;

http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec8677;

http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec8678;

http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec8679;

MNHN

Museum National d'Histoire Naturelle

QM

Queensland Museum

QDPC

Queensland Primary Industries Insect Collection

ANIC

Australian National Insect Collection

AM

Australian Museum

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Lepanus

Loc

Lepanus ustulatus ( Lansberge, 1874 )

Gunter, Nicole L. & Weir, Thomas A. 2021
2021
Loc

Lepanus ustulatus:

Matthews, E. G. 1974: 104
1974
Loc

Panelus ustulatus:

Lea, A. M. 1923: 361
1923
Loc

Epilissus ustulatus

Paulian, R. 1934: 282
Gillet, J. J. E. 1925: 5
Gillet, J. J. E. 1911: 39
Masters, G. 1886: 10
Lansberge, G. van 1874: 7
1874