Euselasia satyroides Lathy, 1926

Santos, Fabio Luis Dos, Dias, Fernando Maia Silva, Leite, Luis Anderson Ribeiro, Dolibaina, Diego Rodrigo, Casagrande, Mirna Martins & Mielke, Olaf Hermann Hendrik, 2014, Taxonomic notes on some species of Euselasia Hübner, [1819] from the " Uriiformes " group, with the description of the immature stages of Euselasia satyroides Lathy, 1926, stat. rev. (Lepidoptera: Riodinidae: Euselasiinae), Zootaxa 3869 (5), pp. 501-522 : 509-510

Euselasia satyroides Lathy, 1926 stat. rev.

( Figs 9–12 , 21–22 )

Euselasia satyroides Lathy, 1926 . p. 144.

Euselasia modesta [misidentification]; Hayward, 1973. p. 168.— Callaghan & Lamas, 2004. p. 143.—Marchiori & Romanowski, 2006. p. 1034—1035.— Núñez-Bustos et al., 2011. p. 40.

Euselasia eugeon [misidentification]; Jørgensen, 1924. p. 87.— Hayward, 1969. p.112—Biezanko et al. 1979. p. 9.— Canals, 2003. p. 218 (male dorsal, male ventral).

Euselasia eugeon brevicauda [misidentification]; Hayward, 1951. p. 167.

Euselasia eusepus brevicauda [misidentification]; Hayward, 1973. p. 168.

Euselasia geon [misidentification]; Núñez-Bustos, 2009. p. 74.— Núñez-Bustos et al., 2011. p. 40.

Euselasia sp.; Siewert, Iserhard, Romanowski, Callaghan & Moser, 2014. (additional file 1).

Diagnosis. E. satyroides is distinguished from other species of the “Uriiformes” group by the brown color of the wings upper side, dull towards the apex and without purple or ocher tinges; and the shape of the hind wing, evenly convex and pointed at the tornus, not forming a distinct projection at 2A. This species is most likely to be confused with E. modesta and E. brevicauda , but can be distinguished from the former by the presence of a ring of white scales basal to the club in the antennae, frons with whitish scaling around the eyes, labial palpus and legs yellow; forewing apex angular; hind wing pointed at the tornus; submarginal and post discal bands and spots in the wings underside conspicuous. It can be distinguished from E. brevicauda by the above cited characteristics and the general appearance of the post discal bands on the wings underside, slightly oblique in the forewing; in the hind wing, the post discal band runs closer to the posterior submarginal band and ends at the inner margin closer to the tornus than in E. brevicauda .

Taxonomic comments. Lathy (1926) described E. satyroides based on four specimens, two males and two females from San Ignacio, Misiones, Argentina (wrongly spelled "San Inguacio" in the original description). This species closest ally probably is E. brevicauda , considering the general appearance of the post discal bands on the underside of the wings and structure of male and female genitalia. Euselasia satyroides hardly ever was recognized as a valid species since the catalog of Stichel (1930). Despite having access to photographs of the type series, Stichel (1927) considered the habitus E. satyroides identical to the description of E. modesta provided by Bates (1868), which he did not have any illustration of the types. However, as above cited under E. modesta , there are a number of differences between these two species. Euselasia satyroides is frequently misidentified as E. eugeon and E. modesta , probably due to the misleading informations provided by Seitz (1913, 1916) and Stichel (1928, 1930) perpetrated in the literature. D’Abrera (1994) completely ignores this species; Jørgensen (1924), Hayward (1969), Biezanko et al. (1979) and Canals (2003: 218) misidentified it as E. eugeon ; and Núñez-Bustos (2009) and Núñez- Bustos et al. (2011) misidentified it as E. modesta and/or E. geon . Biezanko et al. (1979) regarded the species as "very rare" in the southeastern Rio Grande do Sul, Brazil. Jørgensen (1924) and Hayward (1969) recorded Chrysophyllum gonocarpum (Sapotaceae) as the host plant of E. satyroides in Paraguay and Argentina, respectively. The immature stages of E. satyroides are detailed described herein.

Distribution. Southern Atlantic forests, in the states of Paraná and Rio Grande do Sul, Brazil; Misiones, Argentina; and Paraguay ( Jørgensen 1924).

Type material. The lectotype of Euselasia satyroides Lathy, 1926 , here designated to avoid future uncertainty about the identity of the species ( ICZN 1999; Art. 74.7.), has the following labels: / Type HT/ SYN-TYPE/ LECTOTYPUS /REPUBLIQUE ARG ne [ Argentina] Haut-Parana [Upper Paraná] San-Ignacio–Missions [Misiones]/Janvier [January]/Collection WAGNER /B. M. TYPE No. Rh 1191/ Euselasia ♂ satyroides Lathy Spec. Typicum / SYNTYPE Euselasia ♂ satyroides Lathy det. P. Ackery, 1990/ LECTOTYPE Euselasia satyroides Lathy, 1926 Dias, Dolibaina, Santos, Leite, Mielke & Casagrande, 2013 / ( BMNH). The characteristic lectotype label will be sent to the curator of the collection.

Examined material. BRAZIL— Paraná: Tuneiras do Oeste — REBIO das Perobas, 8- X-2012, 1 male, LABLEP [LELN-UFPR] leg., DZ 26.788 ( DZUP). Rio Grande do Sul: Catuípe, 28- IV-2004, 1 male, F. L. Santos leg., DZ 23.099 ( DZUP), 23- III-2009, 1 male, F. L. Santos leg., FLS 12.604 (FLS), 4- V-2009, 1 male, F. L. Santos, leg., FLS 12.723 (FLS), Ponte Santo Antônio, RS-342, Km 95, -28.248º -54.013º, 327 m, IV-2009, 1 female, F. L. Santos leg., FLS 12.724 ( FSL), 30-III-16- IV-2009, 9 males and 1 female, F. L. Santos leg., DZ 22.939*, DZ 23.159, DZ 23.089, DZ 23.199, DZ 23.109, DZ23.079, DZ 23.149, DZ 23.169, DZ 23.179, DZ 22.949*, DZ 23.119, DZ 23.1298*, DZ 23.189, DZ 23.209, DZ 23.139 ( DZUP) and 5 females (FLS), (all reared in Pouteria salicifolia ( Sapotaceae )); Pelotas, 26- V-1966, 1 female, Biezanko leg., DZ 22.969 ( DZUP).

Figures

FIGURES 1 – 16. Habitus of species of Euselasia Hübner, [1819] belonging to the " Uriiformes " group. 1 – 4. E. brevicauda. 1 – 2. Male dorsal and ventral (Brazil, Acre, Xapuri). 3 – 4. Female dorsal and ventral (Brazil, Rondônia, Ariquemes). 5 – 8. E. eugeon. 5 – 6. Male dorsal and ventral (Brazil, Amazonas, Presidente Figueiredo). 7 – 8 Female dorsal and ventral (Brazil, Pará, Óbidos). 9 – 12. E. satyroides, stat. rev. 9 – 10. Male dorsal and ventral (Brazil, Rio Grande do Sul, Catuípe). 11 – 12 Female dorsal and ventral (Brazil, Rio Grande do Sul, Catuípe). 13 – 14. E. modesta, male dorsal and ventral (French Guiana, Saint- Laurent-du-Maroni, Saül). 15 – 16. E. uria, male dorsal and ventral (Brazil, Amazonas, Manaus). Arrowheads indicate characters cited in the taxonomic key. Scale bar = 1 cm.

FIGURES 17 – 22. Genitalia of species of Euselasia Hübner, [1819] belonging to the " Uriiformes " group. 17 – 18. E. brevicauda. 17. Male genitalia, lateral. 18. Female genitalia, ventral. 19 – 20. E. eugeon. 19. Male genitalia, lateral. 20. Female genitalia, ventral. 21 – 22. E. satyroides, stat. rev. 21. Male genitalia, lateral. 22. Female genitalia, ventral.