Subancistrocerus clypeatus Tan, 2023

Tan, Jiang-Li, Wang, Meng, Xu, Hongli, Tang, Yan & Liu, Ying, 2023, Key to the species of the genus Subancistrocerus de Saussure, 1855 (Hymenoptera, Vespidae, Eumeninae) from China with description of a new species, Journal of Hymenoptera Research 95, pp. 113-127 : 113

Subancistrocerus clypeatus Tan sp. nov.

Figs 4 , 5 , 6

Material examined.

Holotype, 1♀, Shunxi, Qingliangfeng (30.13°N, 119.04°E), Zhejiang, China, 24.viii.2018, coll. Jiangli Tan. paratype, 1♂, same data as holotype.

Diagnosis.

The species is similar to two Chinese species S. kankauensis and S. compressus and to a species S. indochinensis from Laos mainly basis of the deep concavity of A11-A12 and A13 flat, reaching the basis of A11 and mid- basitarsomere curved, about 5.0 × as long as wide, but can be separated from S. kankauensis and S. compressus by the characters indicated in the key. It also differs from S. indochinensis by the following characters: 1) in S. clypeatus sp. nov., clypeus of male slightly longer than wide (width: length = 2.0: 2.2) with apical emarginate (width: depth = 1.0: 0.2), but in S. indochinensis , clypeus as wide as long (width: length = 2.0: 2.0) with apical margin slightly emarginate (width: depth = 1.4: 0.2). 2) in S. clypeatus sp. nov., clypeus of female as wide as long (width: length = 2.0: 2.0) with apical slightly margin emarginate (width: depth = 1.4: 0.2), black laterally, yellow medially with a black transverse spot, while in S. indochinensis , the clypeus of female slightly wider than long (2.5: 2.3) with apical margin slightly emarginate (width: depth = 1.4: 0.2), black with a yellow transverse band basally and two yellow spots above the apical margin. 3) apical yellow band of T3 and S3 present in S. clypeatus sp. nov., but absent in S. indochinensis .

Description.

Holotype. ♀, Length of body (up to apex of T2) 6.3 mm with forewing 5.4 mm long.

Body black with yellow to brownish yellow maculation. Basal 2/5 of the mandible yellow except a small black triangle, and its distal 3/5 dark brown. Clypeus black laterally, yellow medially with a black transverse spot, and with its apical margin brown translucent. Other yellow or brownish yellow parts normally smooth or sparsely punctate as follows (Fig. 4A-D ): an inter-antennal spot, scape ventrally, spot on ventral side of each flagellum, a small spot on the apical ocular sinus, a spot behind the eye on upper surface of temple, two small separated irregular spots on pronotum dorsally, tegula sub -core periphery, parategula, separated metanotal spots, propodeal valvula and apical lamella of submarginal carina of propodeum, dorso-apical fore femora, dorsal fore tibiae, wide apical bands of terga 1-2 and sternum 2, narrow band of T3, very small trace of S3 medially. Wings hyaline with veins and pterostigma dark brown, and with a brown cloudy patch along the costal margin; body with white to yellowish brown pubescence (Fig. 5A ).

Head (Fig. 5B, C, E ). Frons deeply punctate and reticulate; temple relatively small and sparsely punctate. Clypeus moderately convex, as wide as long with medial depression in shield shaped that bounded laterally by two indistinct sinuate carinae; apical margin emarginate (width: depth = 1.4: 0.2) between distinct projecting teeth, 0.3 × of maximum width of the clypeus, laterally angulate; clypeus densely punctate laterally, weakly punctate to smooth at the medial depression portion, but densely punctate at black part. Antenna short and clavate, length of the first flagellar segment F1: width = 3.3: 2.5, F2 as wide as long, length of F3: width = 6: 7.

Mesosoma (Fig. 5D, F ). Mesosoma 1.4 × as long as wide and 1.6 × as its high. Humeral corner of pronotum rounded; pronotum, mesoscutum, mesoscutellum densely punctate, punctures obviously larger and deeper than those on head. Mesepisternum densely reticulate-rugose, meshes on the upper part larger than on lower part. Propodeum densely reticulate-rugose dorsally and laterally; its posterior surface concaved, more or less shiny with fine, microscopic, diagonally striped (Fig. 4B ); metapleuron impunctate and somewhat coriaceous with transverse wrinkles and densely white pubescence.

Metasoma (Fig. 5G-I ). T1 about 1.8 × wider than long. Surface between the two transverse carinae of T1 smooth and shiny; horizontal surface of T1 behind second carina to apical margin 2.5 × as wide as long. T1 coarsely punctate behind second transverse carina, with backwards open punctures, and with very narrow transparent lamellate apical margin; diameter of punctures less than distance between punctures; apical yellow band of T1 evenly wide; S1 coriaceous without punctures. T2 punctate with punctures sparser medially than basally and subapically; S2 sparsely punctate with punctures relatively smaller than T2, diameter of punctures more than distance between punctures; T3 and S3 densely punctate similar to T1, lateral arched carina indistinct; T4 and S4 smooth basally and sparsely punctate distally, punctures gradually larger and deeper towards apex. Fifth to the remaining metasomal segments smooth basally and very finely punctate towards apex.

Male (Fig. 6A ): Similar to female except as follows: a large spot on ocular sinus; clypeus and mandible both entirely yellow with long pubescence; clypeus slightly longer than wide (length: width = 2.2: 2.0) without carinae, smooth except basal half finely and sparsely punctate; medially convex with ventral 1/3 portion strongly depressed; apical margin emarginate (width: depth = 1.0: 0.2), 0.36 × of maximum width of clypeus (Fig. 6F ); dorsal apex of femora, tibiae except a dark brown patch on its ventral surface and basitarsomere of all legs yellow. Antennae clavate with an apical cavity on the article 11-12 (A11-A12); terminal article A13 flat, shaped as a corresponding cover reaching the basis of A11, 1.8 × as long as wide, rounded apically; A3-A9 laterally each with an indistinct (A3) or a distinct tyloid (a carina-like prominence) without concavities ventrally (Fig. 6B-E ). A3, A4 and A5 is 1.7 ×, 1.2 × and 1.0 × as long as wide. Basal half of fore femur ventrally compressed (Figs 6G-H ); basitarsomere of mid leg curved, 5.0 × as long as wide, 1.2 × longer than the combine length of tarsomeres 2-5 and pretarsus (Fig. 6I ).

Distribution.

China (Zhejiang)

Etymology.

The new species is named from Latin "clypeus, shield" referring to its shield shaped patch on the middle of female clypeus for the depression bordered by two longitudinal sinuate carinae.

Figures

Figure 4. Subancistrocerus clypeatus Tan, sp. nov., holotype, ♀ A wing B propodeum posterior surface C fore tarsi D mid- and hind tibiae and pretarsus.

Figure 5. Subancistrocerus clypeatus Tan, sp. nov., female, holotype A forewing B head, frontal aspect C ibid, dorsal aspect D mesosoma, dorsal aspect E head, lateral aspect F mesosoma lateral aspect G metasoma, dorsal aspect H ibid, ventral aspect I ibid, lateral aspect.

Figure 6. Subancistrocerus clypeatus Tan, sp. nov., male, paratype A habitus, lateral aspect B, E right antenna, ventral aspect, show the tyloids and yellow patches and deep concavity of A 11 - A 12 C ibid, dorsal aspect D ibid, A 4 - A 13 enlarged head, frontal aspect F head, frontal aspect G left fore-femora and tibia, frontal aspect H foreleg, caudal aspect I mid-leg without pretarsus, frontal aspect.