Phanaeus nimrod Harold, 1863

Moctezuma, Victor, Halffter, Gonzalo & Lizardo, Viridiana, 2021, The Phanaeus tridens species group (Coleoptera: Scarabaeoidea): a dung beetle group with genital morphological stasis but a changing ecological niche, Acta Entomologica Musei Nationalis Pragae 61 (2), pp. 447-482 : 464

publication ID

https://doi.org/ 10.37520/aemnp.2021.025

publication LSID

lsid:zoobank.org:pub:85434EFF-F859-4BBF-8AB5-F50B9BA08771

DOI

https://doi.org/10.5281/zenodo.6303366

persistent identifier

https://treatment.plazi.org/id/1B2B878A-8A3B-FFAC-FC68-FEEFEB1FF7A0

treatment provided by

Plazi

scientific name

Phanaeus nimrod Harold, 1863
status

 

Phanaeus nimrod Harold, 1863 View in CoL

( Figs 24–26 View Figs 24–26 , 40 View Figs 32–42 , 52 View Figs 43–54 , 60 View Figs 55–62 )

Phanaeus nimrod Harold, 1863: 167 View in CoL . Type locality: Mexico.

Phanaeus babori Balthasar, 1939:246 View in CoL .Synonymy established by ARNAUD (1982b: 125) and confirmed here.

Type material examined. MEXICO: OAXACA: HOLOTYPE of P. babori (designated by monotypy by BALTHASAR 1939, examined from photographs; Fig. 26 View Figs 24–26 ): ♂, Oaxaca ( NMPC: Mus. Nat. Pragae Inv. 26352). W I THOUT SPEC I F I C LOCAL I TY: L ECTOTYPE of P. nimrod (designated by ARNAUD 1982a, examined from photographs; Fig.25 View Figs 24–26 ):♂, without specific locality ( MNHN: MNHN EC10555). P ARALECTOTYPES of P. nimrod : 1♂ 1 ♀ (examined from photographs), without specific locality ( MNHN: MNHN EC10556, MNHN EC10557).

Non-type material examined. MEXICO: OAXACA: 1 ♂ 1 ♀, Ocotlán ( GHVM); 6 ♂♂ 6 ♀♀, Santa Cruz Papalutla ( JLSHVM: 3 ♂♂ 3 ♀♀; VMPM: 3 ♂♂ 3 ♀♀); 1 ♀, 2 km S Sola de Vega ( IEXA).

Diagnosis. Metallic turquoise ( Fig. 25 View Figs 24–26 ), green-blue ( Figs 26 View Figs 24–26 , 60 View Figs 55–62 ), yellow-green, green-red ( Fig. 24 View Figs 24–26 ) or dark blue. Sides of pronotal disc finely granulate to finely granulorugose ( Figs 24–26 View Figs 24–26 , 40 View Figs 32–42 , 52 View Figs 43–54 , 60 View Figs 55–62 ). Pronotal disc smooth to superficially punctate anteriorly, sparsely roughened posteriorly ( Figs 24–26 View Figs 24–26 , 52 View Figs 43–54 ). Posteromedial process of pronotum produced into denticle, strongly widened laterally, not bifurcated apically ( Figs 40 View Figs 32–42 , 52 View Figs 43–54 ). Anteromedial portion of pronotal disc with two denticles, distinctly separate ( Fig. 52 View Figs 43–54 ).Anterolateral margins of pronotal disc with distinctly developed line of tubercles ( Fig. 52 View Figs 43–54 ). Posterolateral angles of pronotum much shorter than posteromedial process of pronotum ( Figs 40 View Figs 32–42 , 52 View Figs 43–54 ). Elytral striae scabriculous, distinctly impressed, superficially punctate ( Figs 24–26 View Figs 24–26 ). Elytral interstriae scabriculous, superficially punctate, convex ( Figs 24–26 View Figs 24–26 ).

Variability. Minor male. Similar to major male, except for reduction of secondary sexual characters (i.e. cephalic horn, pronotal processes and posterolateral angles). Female. Similar to male, except for head showing trituberculate carina; pronotal sculpture granulate, becoming smooth to superficially punctate posteriorly; pronotum with anteromedial black macula, and anteromedial carina followed by posterior concavity ( Fig. 60 View Figs 55–62 ). This species is highly variable in colour ( Figs 24–26 View Figs 24–26 , 52 View Figs 43–54 , 60 View Figs 55–62 ).

Comments. Apparently, this species is well defined and is not involved in taxonomical controversies. Phanaeus nimrod is typical of the Central Valleys of Oaxaca ( Fig. 67 View Fig ), but DELOYA et al. (2014) reported it from an isolated locality in Guerrero (El Ocotito). The distribution of this species needs to be confirmed since doubtful records from the Pacific Coast of Oaxaca have been provided ( EDMONDS 1994, 2003; DELOYA & MORÓN 1998). None of the specimens of the P. tridens species group revised in the present work from the Mexican Pacific coast belong to P. nimrod . The major male figured by DELOYA & COVARRUBIAS-MELGAR (2014) seems to be a green specimen of P. nimrod .

Distribution. Mexico: Central Oaxaca to central Guerrero ( Fig. 67 View Fig ). Records from Guerrero and the Pacific coast of Oaxaca were omitted for the distribution model because they need confirmation.

NMPC

National Museum Prague

MNHN

Museum National d'Histoire Naturelle

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Phanaeus

Loc

Phanaeus nimrod Harold, 1863

Moctezuma, Victor, Halffter, Gonzalo & Lizardo, Viridiana 2021
2021
Loc

Phanaeus babori

BALTHASAR V. 1939: 246
1939
Loc

Phanaeus nimrod

HAROLD E. 1863: 167
1863
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