Cambalida deorsa, Murthappa, Prashanthakumara S., Prajapati, Dhruv A., Sankaran, Pradeep M. & Sebastian, Pothalil A., 2016

Murthappa, Prashanthakumara S., Prajapati, Dhruv A., Sankaran, Pradeep M. & Sebastian, Pothalil A., 2016, First records of the genus Cambalida Simon, 1909 (Araneae: Corinnidae, Castianeirinae) from Asia, with the description of two new species from India and one new combination, Zootaxa 4103 (6), pp. 526-536 : 527-531

publication ID	https://doi.org/ 10.11646/zootaxa.4103.6.3
publication LSID	lsid:zoobank.org:pub:D035CE30-3C63-4397-BE81-ABD3EF07A2A8
DOI	https://doi.org/10.5281/zenodo.6089746
persistent identifier	https://treatment.plazi.org/id/1B4D6A15-0E0A-FFA3-FF2E-FBAEFC9D95D3
treatment provided by	Plazi
scientific name	Cambalida deorsa
status	sp. nov.

Treatment

Cambalida deorsa View in CoL sp. n.

( Figs 1A–K View FIGURE 1 A – K , 2A–E View FIGURE 2 A – E , 3A–E View FIGURE 3 A – E )

Type material. Holotype: ♂ with left leg I missing ( ADSH 18102A), INDIA: Karnataka: Shimoga: Shankaraghatta: Jnana Sahyadri campus of Kuvempu University, 13°44'00.92"N, 75°37'44.22"E, 680 m a.s.l., S.M. Prashanthakumara leg., 28 May 2015, by hand from the ground; Paratypes: 2 ♀ ( ADSH 18102B), same data as holotype except 14 September 2015; 1 ♀ ( ADSH 18102C), INDIA: Gujarat: Gandhinagar, The Serenity Library and Botanical Garden in Koteshwar village, 23°06'42.27''N, 72°37'23.95''E, 63 m a.s.l., D.A. Prajapati leg., 26 June 2015, by hand from the ground.

Etymology. The specific epithet is an adjective and is derived from the downwardly directed median turn of the embolus. Latin deorsum = downward.

Diagnosis. Cambalida deorsa sp. n. is most similar to C. compressa Haddad, 2012 from West Africa but can be distinguished by the following combination of characters: embolus nearly uniform in width along the entire length ( Figs 2A–C View FIGURE 2 A – E , 3A–C View FIGURE 3 A – E ) (embolus in C. compressa broad proximally but narrow distally, see Haddad 2012: fig. 59); median turn of embolus oblique to the longitudinal axis and directed downward ( Figs 2C View FIGURE 2 A – E , 3C View FIGURE 3 A – E ) (in C. compressa and all other described Cambalida spp. with either transverse or oblique and distally directed median embolar turn, see Haddad 2012: figs 50–56); embolic tip directed at 11 o’ clock in ventral view ( Figs 2C View FIGURE 2 A – E , 3C View FIGURE 3 A – E ) (embolic tip in C. compressa directed at 12 o’ clock in ventral view, see Haddad 2012: fig. 50); epigyne with circular plate-like ridges ( Figs 2D View FIGURE 2 A – E , 3D View FIGURE 3 A – E ) ( C. compressa with 6-shaped epigynal ridges, see Haddad 2012: fig. 57); copulatory ducts with sharp median curve ( Fig. 3E View FIGURE 3 A – E ) (copulatory ducts of C. compressa with less prominent median curve, see Haddad 2012: fig. 58); fertilization ducts short ( Fig. 3E View FIGURE 3 A – E ) ( C. compressa with long fertilization ducts, see Haddad 2012: fig. 58); and posterior border of epigyne W-shaped ( Figs 2D, 2E View FIGURE 2 A – E , 3D, 3E View FIGURE 3 A – E ) (posterior border of epigyne in C. compressa convex, see Haddad 2012: figs 57, 58).

Description. Male in alcohol (holotype, Figs 1A–E View FIGURE 1 A – K ): Prosoma, chelicerae, fangs, sternum reddish-brown. Prosoma pear-shaped, with tiny tubercles; fovea short, distinct. Eyefield black. Clypeus, labium, maxillae brownish. Clypeus provided marginally with long thick bristles. Maxillae with scopulae, labium not. Chelicerae with apical fang shield mound bearing long, unmodified setae; cheliceral promargin with 3 teeth, median largest; retromargin with 2 teeth. Coxae yellowish-brown; femora I & II brownish, III & IV yellowish; tarsi I & II brownish, III & IV yellowish. Opisthosoma rectangular, with dorsal, ventral, epigastric and tracheal (inframamillary) scuta; tracheal scute broad, nearly circular in shape ( Fig. 1E View FIGURE 1 A – K ); post-epigastric sclerites well developed ( Fig. 1E View FIGURE 1 A – K ); dorsum blackish-brown, with large whitish patch posteriorly just above the spinnerets; venter reddish-brown. Prosoma, opisthosoma, legs covered with feathery hairs; femora III and IV with distal constrictions ( Fig. 1K View FIGURE 1 A – K , arrow). Spinnerets grey with whitish tip. Body length 3.7. Prosoma length 1.88, width 1.36, height 0.58. Opisthosoma length 1.82, width 1.16, height 1.04. Eye diameters: ALE 0.09, AME 0.08, PLE 0.11, PME 0.10. Eye interdistances: AME–AME 0.06, ALE–ALE 0.26, ALE–PME 0.16, PLE–PLE 0.36, PME–PME 0.12, PME–PLE 0.02, AME–ALE 0.01, AME–PME 0.12, ALE–PLE 0.07. Clypeus height at ALE 0.11, at AME 0.15. Chelicerae length 0.58. Measurements of palp and legs. Palp (right) 1.75 [0.57, 0.23, 0.24, 0.71], I (right) 4.97 [1.37, 0.45, 1.20, 1.10, 0.84], II 4.29 [1.22, 0.41, 1.01, 0.94, 0.71], III 4.23 [1.15, 0.42, 0.91, 1.15, 0.60], IV 6.66 [1.73, 0.53, 1.58, 1.93, 0.89]. Leg formula: 4123. Spination. Palp: femur do 2, patella do 1, tibia pl 1, cymbium/tarsus pl 3; legs: femora: I (right)–II pl 1 do 3, III–IV pl 2 do 3 rl 1; patellae: I–IV spineless; tibiae: I (right) plv 2 rlv 1, II rlv 1, III pl 2 do 1 rl 2 plv 2 rl 1 rlv 1, IV pl 2 do 1 rl 2 plv 3 rlv 2; metatarsi: I (right) plv 2 rlv 2, II plv 1 rlv 2, III pl 2 rl 2 plv 3 rlv 2 vt 1, IV pl 3 rl 3 plv 3 rlv 2 vt 1; tarsi: I–IV spineless. Pedipalp ( Figs 2A–C View FIGURE 2 A – E , 3A–C View FIGURE 3 A – E ). Palpal segments dark brown; cymbium/tarsus apically with six stout setae on dorsal surface, arranged in three rows (2–2–2); apical cymbium shows an unusual modification forming a ‘thick cone’ ( Figs 2A View FIGURE 2 A – E , 3A View FIGURE 3 A – E ). Bulb straw coloured; sperm duct broad along the entire length; embolus with nearly uniform thickness, with 1¼ coils, with pointed tip directed at 11 o’ clock in ventral view ( Figs 2B–C View FIGURE 2 A – E , 3B–C View FIGURE 3 A – E ).

Female in alcohol (paratype, Figs 1F–K View FIGURE 1 A – K ): In all details like male except the following: Prosoma with a median longitudinal dark band between the fovea and the posterior eye row, with dorso-lateral black striations. Chelicerae with apical fang shield mound bearing long, unmodified setae ( Fig. 1J View FIGURE 1 A – K ). Opisthosoma ovate, greyish; dorsum with small anterior scutum, with two pairs of sigilla; posteriorly with a pair of short, parallel, rectangular white patches; venter with four longitudinal discontinuous lines, without ventral scutum; post-epigastric sclerite poorly defined. Leg segments of III and IV brownish. Palpal segments yellowish-brown. Body length 4.94. Prosoma length 2.24, width 1.68, height 0.62. Opisthosoma length 2.70, width 1.76, height 1.74. Eye diameters: ALE 0.12, AME 0.11, PLE 0.14, PME 0.135. Eye interdistances: AME–AME 0.06, ALE–ALE 0.31, ALE–PME 0.15, PLE–PLE 0.49, PME–PME 0.13, PME–PLE 0.07, AME–ALE 0.01, AME–PME 0.12, ALE–PLE 0.07. Clypeus height at ALE 0.10, at AME 0.10. Chelicerae length 0.89. Measurements of palp and legs. Palp 2.17 [0.65, 0.31, 0.43, 0.78], I 6.01 [1.68, 0.57, 1.47, 1.31, 0.98], II 5.45 [1.58, 0.53, 1.26, 1.17, 0.91], III 5.17 [1.51, 0.40, 1.07, 1.40, 0.79], IV 7.94 [2.03, 0.60, 1.96, 2.24, 1.11]. Leg formula: 4123. Spination. Palp: femur do 2, patella pl 1 do 2, tibia pl 2 rl 2, tarsus pl 2 rl 1 plv 1 rlv 1; legs: femora: I pl 1 do 3, II pl 1 do 3 rl 1, III–IV pl 2 do 3 rl 1; patellae: I–IV spineless; tibiae: I plv 2 rlv 2, II pl 1 rlv 2, III–IV pl 2 do 1 rl 2 plv 3 rlv 2; metatarsi: I–II plv 2 rlv 2, III–IV pl 4 do 1 rl 4 plv 2 rlv 2 vt 1; tarsi: I–IV spineless. Epigynum ( Figs 2D–E View FIGURE 2 A – E , 3D–E View FIGURE 3 A – E ): Weakly sclerotised with circular epigynal ridges, with medium sized lateral copulatory openings, with W-shaped posterior margin ( Figs 3D–E View FIGURE 3 A – E ). Spermathecae dumb-bell shaped with spherical anterior part (spermathecae II) and reniform posterior part (spermathecae I). Copulatory ducts long, dark, with sharp median inward curving ( Figs 2E View FIGURE 2 A – E , 3E View FIGURE 3 A – E ). Fertilization ducts tiny, medially placed on spermathecae I ( Fig. 3E View FIGURE 3 A – E ).