Gudeodiscus (Gudeodiscus?) anceyi (Gude, 1901)

Pall-Gergely, Barna, Hunyadi, Andras, Ablett, Jonathan, Lương, Hao Văn, Fred Naggs, & Asami, Takahiro, 2015, Systematics of the family Plectopylidae in Vietnam with additional information on Chinese taxa (Gastropoda, Pulmonata, Stylommatophora), ZooKeys 473, pp. 1-118 : 10

Taxon classification Animalia Stylommatophora Plectopylidae

Gudeodiscus (Gudeodiscus?) anceyi (Gude, 1901) Figures 2B, 9G, 11 C–F

Plectopylis Anceyi Gude 1901a, Journal de Conchyliologie, 49: 208-209., Figs 6 a–e, Plate 6, Figs 6 a–c. ["Bac-Kan (le type); secteur de Nac-Ri; entre Cho-Moi et That-Khé”]

Gudeodiscus anceyi , - Páll-Gergely & Hunyadi 2013, Archiv für Molluskenkunde, 142 (1): 8.

Types examined.

Tonkin, Bac-Kan, leg. Messager, MNHN 24600 (syntype, Figure 2B); Tonkin, Bac-Khan, NHMW 50858 (2 syntypes).

Museum material examined.

Tonkin, coll. Jetschin ex Berlier 1908, SMF 118124/2; Tonkin, Bac-Khan, coll. Jaeckel, S. H. ex Rolle, SMF 207668/1; Tonkin, Bac-Khan, coll. Dosch ex Rolle, SMF 172078/4; Tonkin, Than-Moi, probably ex Messager, SMF 150135/1; Central-Tonkin, Chiam-Hoa, coll. Möllendorff ex Fruhstorfer SMF 150134/1; Tonkin, Bac-Kan, leg. Messager, 22.11.1898, RBINS/5; Secteur de Nac-Ri, RBINS/1; Secteur de Nac-Ri, leg. Messager (n. 33), RBINS/1; Tonkin, entre Cho-Moi, et That-Khé, leg. Messager (n. 33), RBINS/11; Tonkin, Bac-Kan, RBINS/4; Muong-Kong, leg. Messager, MNHN-IM-2012-2139/1; Secteur de Nac-Ri, Bac-Kan, leg. Messager, MNHN-IM-2012-2250/343; Bac-Kan, leg. Messager, MNHN-IM-2012-2252/60; Bac-Kan, leg. Messager, MNHN-IM-2012-2258/38; That-Khé, coll. Mansuy, MNHN-IM-2012-2259/12; Cho-Moi, leg. Messager, MNHN-IM-2012-2263/48; Bac-Kan, leg. Messager, MNHN-IM-2012-2265/30; Long-Phai, leg. Messager, MNHN-IM-2012-2270/36; Cho-Moi, leg. Messager, MNHN-IM-2012-2275/30; Long-Phai, leg. Messager, MNHN-IM-2012-2277/26; Cho-Moi, leg. Messager, MNHN-IM-2012-2283/95; Cao-Bang, leg. Messager, MNHN-IM-2012-2468/1; Na-Ri, leg. Messager, MNHN-IM-2012-2285/40; Long- Phai, leg. Messager, MNHN-IM-2012-2286/36; Bac-Kan, coll. Letellier, 1949, MNHN-IM-2012-2287/1; Cho-Moi, leg. Messager, MNHN-IM-2012-2300/25; Bac-Kan, coll. Lavezzari, 1929, MNHN-IM-2012-2301/15; Bac-Kan, leg. Messager, MNHN-IM-2012-2305/62; Long-Phai, leg. Messager, MNHN-IM-2012-2312/30; Bac-Kan, coll. Staadt, 1969, MNHN-IM-2012-2313/4; Na-Ri, leg. Messager, MNHN-IM-2012-2376/34; Pakhé, leg. Messager, MNHN-IM-2012-2453/1; Tonkin, Bac-Khan, coll. Rolle, 4/11/08, NHMUK 20130585/3; Tonkin, Bac-Khan, coll. Rolle, 4/11/08, NHMUK 20130586/3; Tonkin, Bac-Kan, 13/6/01, NHMUK 20130587/3; Tonkin, Bac-Kan, coll. Rolle, 4/11/08, NHMUK 20130588/3; Tonkin, 4/11/8, NHMUK 20130589/2; Tonkin, Bac-Kan, coll. Salisbury ex Beddome, NHMUK 20130590/2; Tonkin, coll. Lucas, NHMUK 20130591/2; Tonkin, Bac-Khan, NHMUK 1916.03.16.1-2/2; Tonkin, NHMUK 1901.08.01.22/1; Tonkin, NHMUK 1901.7.11.89-90/2; Tonkin, Bac-Kan, coll. Rušnov ex Rolle ex Messager, NHMW 92556/6; Tonkin, Bac-Kan, coll. Wagner ex Messager, NHMW 92557/2; Tonkin, Cho-Moi, coll. Oberwimmer ex. Rosen, NHMW 71640/O/9480/1; Tonkin, Ngam-Son, coll. Wagner ex Messager, NHMW 82558/2; Tonkin, Cho-Moi, coll. Rosen, NHMW 71640/O/9479/2; Tonkin, Bac-Khan, coll. Rolle ex Messager, NHMW 50858/2; Tonkin, That-Khé, entre Cho-Moi, coll. Steenberg, ZMUC-GAS-1809/2.

New material examined.

Vn10-33B Bắc Kạn Province, Ba Bể Nat. Park, surroundings of Na Phoong cave, GPS not recorded, leg. Hemmen, Ch. & J., 10.10.2010., PGB/1; GS21 Bắc Kạn Prov, Na Rì District, left side of road from Kim Hỷ to Bắc Kạn, 2 km after Kim Hỷ, in leaf litter bellow high limestone walls above road, 583 m, 22°16.861'N, 106°2.169'E, leg. Grego, J. & Śteffek, J., 06.04.2012., JG/1; GS22 Bắc Kạn Prov, Na Rì District, 2 km S of Bản Dền (=Dền Village), limestone rocks at side of the valley near gold quarry, in small cavern in dense rain forest, ca 590 m, 22°14.547'N, 106°0.527'E, leg. Grego, J. & Śteffek, J., 06.04.2012., JG/1, PGB/1; GS24 Bắc Kạn Prov, Na Rì District, 2 km S of Bản Dền, W slopes of a deep sinkhole covered with forest, leaf litter under high limestone wall, ca 640 m, 22°14.506'N, 106°0.521'E, leg. Grego, J. & Śteffek, J., 06.04.2012., JG/1; 2011/82 Lạng Sơn Province, Lũng Phầy Pass, Thất Khê N 13 km, 475 m, 22°20.363'N, 106°27.098'E, leg. Hunyadi, A., 15.11.2011., HA/4; 2011/91 Bắc Kạn Province, Ba Bể Nat. Park, 500 m on the path starting from the bungalows, 240 m, 22°25.072'N, 105°37.941'E, leg. Hunyadi, A., 17.11.2011., HA/3; 2011/93 Bắc Kạn Province, Ba Bể Nat. Park, Đầu Đằng Waterfall, above the waterfall, 175 m, 22°27.159'N, 105°34.193'E, leg. Hunyadi, A., 18.11.2011., HA/1; 2011/94 Bắc Kạn Province, Ba Bể Nat. Park, Ao Tiên, near the lake, 155 m, 22°26.831'N, 105°37.023'E, leg. Hunyadi, A., 18.11.2011., HA/3+1jb; 2011/96 Bắc Kạn Province, Ba Bể Nat. Park, Thẳm Kịt Cave 2 km, look-out tower, 335 m, 22°24.686'N, 105°37.710', leg. Hunyadi, A., 19.11.2011., HA/1; 2011/100 Bắc Kạn Province, Ba Bể Nat. Park, Bố Lù, 600 m from the harbour towards Pắc Ngòi, right side of the road, 175 m, 22°23.989'N, 105°37.523'E, leg. Hunyadi, A., 19.11.2011., HA/3; 2011/101 Bắc Kạn Province, Ba Bể Nat. Park, Na Phoong Cave, south of Bố Lù, 215 m, 22°23.341'N, 105°36.812'E, leg. Hunyadi, A., 19.11.2011., HA/3; 2012/45 Bắc Kạn Province, Na Rì Distr., Kim Hỷ SSE, 1.5 km on a by-road from the road nr. 279, 420 m, 22°16.988'N, 106°02.990'E, leg. Hunyadi, A., 29.05.2012., HA/3; Vn10-68 Cao Bằng Province, right off old rd., ca. 33 km from Cao Bằng to Đông Khê, 22°27.547'N, 106°22.331'E, leg. Hemmen, Ch. & J., 26.03.2010., HE/1; Vn11-159 Lạng Sơn Province, at km 74.8 on road 1B, Đồng Đăng to Thái Nguyên (8 km S Bắc Sơn), 21°54.543'N, 106°17.298'E, leg. Hemmen, Ch. & J., 02.04.2011., HE/7; Vn11-31C Bắc Kạn Province, Ba Bể Nat. Park, near Puổng Cave, 22°27.835'N, 105°38.997'E, leg. Hemmen, Ch. & J., 17.03.2011., HE/1; same data, leg. Hemmen, Ch. & J., 19.10.2009., PGB/2.

Diagnosis.

Shell very small, finely ribbed, whole shell with easily-visible spiral lines, spire elevated, umbilicus deep; aperture with well-developed, long apertural fold (Figure 9G). Parietal wall with two lamellae, the anterior is fused with the lower plica, upper plica missing (or short and fused to the anterior lamella); palatal plicae oblique, short, sometimes connected with a ridge (Figures 11 C–F).

Measurements

(in mm): D = 7.4-7.9, D: 3.5-4 (shells from different localities, n=3); D = 9.2-9.8, H = 4.5-4.6 (Vn11-31C).

Differential diagnosis.

Gudeodiscus messageri is larger than Gudeodiscus anceyi and lacks the apertural fold and spiral lines on the ventral surface of the shell. Gudeodiscus anceyi is smaller than typical Gudeodiscus phlyarius , has stronger spiral lines, and has no horizontal plica under the lamellae, which are present in most populations assigned to Gudeodiscus phlyarius . The Gudeodiscus phlyarius populations living near the Chinese border (typical anterides , gouldingi , fallax , verecunda ) are usually larger than Gudeodiscus anceyi and they often lack the apertural fold and the spiral lines on the ventral side of the shell. For differences with Gudeodiscus hemmeni sp. n. and Sicradiscus mansuyi , see under those species.

Intraspecific diversity.

Relatively low; shell characters, namely the size and general shell and aperture shape are rather stable. The morphology of the palatal plicae shows some diversity. The species is easily recognisable and can be separated from other plectopylid species without major problems.

Distribution

(see Figure 40): We have newly-collected material only from Bắc Kạn Province. The species was previously recorded from That Khé (Lạng Sơn Province) and Nac Ri ( Hà Giang Province) ( Gude 1901a, see also Figure 39).