Paramblynotus yangambicolus, Noort & Buffington, 2013
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publication ID |
https://dx.doi.org/10.3897/jhr.31.4072 |
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publication LSID |
lsid:zoobank.org:pub:DFD1344D-FCA6-42CD-BD68-4FDF2E73F9AC |
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persistent identifier |
https://treatment.plazi.org/id/1C26338B-EFD9-3487-548B-701D101D7E63 |
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treatment provided by |
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scientific name |
Paramblynotus yangambicolus |
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species-group |
Paramblynotus yangambicolus species-group
Remarks.
Historically this taxon has been recognized variously at generic ( Decellea Benoit, 1956) or subgeneric level ( Liu et al. 2007). Decellea was synonymized with Paramblynotus by Weld (1962); followed by re-establishment of generic status by Ronquist (1995); and subsequent re-synonymisation based on phylogenetic analyses, which showed this species-group to be deeply nested within the Paramblynotus clade ( Liu et al. 2007).
Previously the species-group was only known from the African mainland with three described species ( Paramblynotus mixtus Liu et al., Paramblynotus alveolatus Liu et al. and Paramblynotus yangambicolus Benoit). We describe two further species from Madagascar (Paramblynotus behara sp. n. and Paramblynotus zohy sp. n.).
Diagnosis.
This species-group is characterized in females by excavations (spiracular peritremata) on the terminal portion of T8 associated with the spiracle ( Figs 7B View Figure 7 ; 9 D View Figure 9 ; 12A-B View Figure 12 ), not referred to by Liu et al. (2007). A distinct pronotal crest is present, medially forming a conspicuous, slightly backward pointing, ridge-like tooth ( Figs 4C View Figure 4 ; 6C View Figure 6 ; 8C-D View Figure 8 ; 10C View Figure 10 ). The mesoscutum has rough discontinuous transverse costae produced into irregularly raised and slightly backward pointing teeth ( Figs 4C-D View Figure 4 ; 6C-D View Figure 6 ; 8E-F View Figure 8 ; 10C-D View Figure 10 ). The speculum is longitudinally costate ( Fig. 11D View Figure 11 , and the median propodeal area is not delimited by lateral propodeal carinae.
Key to species of the Paramblynotus yangambicolus species-group (modified from Liu et al., 2007)
| 1 | Forewing with RS+M vein arising at or very close to base of basal vein ( Figs 5B View Figure 5 ; 12C View Figure 12 ) (Madagascar) | 2 |
| - | Forewing with RS+M vein arising at basal third or mid-way up basal vein ( Figs 9B; 9E View Figure 9 ) (Africa) | 3 |
| 2 | Head and mesosoma black ( Figs 4C-D View Figure 4 ). Abdominal tergite 2 (petiole) dorsally sparsely variolate with confused, weak, longitudinal carinulation ( Fig. 4F View Figure 4 ); laterally with strong, widely spaced longitudinal carinae ( Fig. 4E View Figure 4 ). RS+M, basal vein and M+Cu1 nebulous ( Fig. 5B View Figure 5 ). RS+M arising from base of basal vein, this juncture represented by a pigmented spot ( Fig. 5B View Figure 5 ). (Female unknown) | Paramblynotus behara sp. n. |
| - | Head pronotum, mesoscutum and scutellum reddish-brown ( Figs 10 C-D View Figure 10 ). Abdominal tergite 2 (petiole) densely longitudinally carinate ( Fig. 11E View Figure 11 ). RS+M, basal vein and M+Cu1 distinct ( Fig. 12C View Figure 12 ). RS+M arising slightly above base of basal vein ( Fig. 12C View Figure 12 ). (Female T8 with two excavations, a smaller circular one surrounding the spiracle and a second larger oval one adjacent to the first and extending to posterior margin) ( Figs 12A-B View Figure 12 ) | Paramblynotus zohy sp. n. |
| 3 | Body length about 4 mm. Body colour entirely dark ( Fig. 6A View Figure 6 ). Eyes prominent, protruding distinctly beyond genae ( Fig. 6E View Figure 6 ). Median frontal carina almost reaching clypeus. Antennal scrobes rugose. Speculum distinctly longitudinally carinate ( Fig. 6C View Figure 6 ). Upper mesopleuron foveate-reticulate. Mesoscutellum sloped posteriorly ( Fig. 6C View Figure 6 ). Wings entirely transparent ( Fig. 6A View Figure 6 ). Metasoma somewhat, but not strongly, compressed laterally. T6 almost about the same size as the three preceding tergites ( Fig. 6F View Figure 6 ). Sterna 4-6 entirely covered by sternum 3 | Paramblynotus mixtus |
| - | Body length about 6-10 mm. Head and mesosoma dark, metasoma yellow to brown ( Fig. 8A View Figure 8 ). Eyes not protruding distinctly beyond genae ( Fig. 9A View Figure 8 ). Median frontal carina absent in lower face. Antennal scrobes entirely longitudinally carinate. Speculum very finely and superficially carinate ( Fig. 8E View Figure 8 ). Upper mesopleuron entirely longitudinally costate. Mesoscutellum raised posteriorly, forming a flat dorsal surface ( Fig. 8C View Figure 8 ). Forewing evenly ferruginous with darker marginal cell and a dark narrow strip along anterior-interior margin of the first submarginal cell ( Fig. 9B View Figure 9 ). Metasoma strongly compressed laterally ( Fig. 9B View Figure 9 ). T6 distinctly larger than any of the 3 preceding tergites ( Fig. 9C View Figure 9 ). Sterna 4-6 exposed, not covered by sternum 3 | 4 |
| 4 | Face evenly curved in lateral view. Genae ventrally strongly expanded posteriorly. Median mesopleural impression distinct. Lower mesopleuron densely punctate with pubescence. Apical teeth of metatibia rounded apically; first metatarsomere 2.0 × length of remaining 4 metatarsomeres combined | Paramblynotus alveolatus |
| - | Face distinctly raised medially and curved inward ventrally in lateral view ( Fig. 8D View Figure 8 ). Median mesopleural impression usually obscured by extension of longitudinal carinae in upper mesopleuron ( Fig. 8C View Figure 8 ). Lower mesopleuron glabrate and sparsely punctate with sparse pubscence. Apical teeth of metatibia pointed apically ( Fig. 9C View Figure 9 ); first metatarsomere 1.5 × length of remaining 4 metatarsomeres combined | Paramblynotus yangambicolus |
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