Ametadoria leticiamartinezae Fleming & Wood

Fleming, AJ, Wood, D. Monty, Smith, M. Alex, Hallwachs, Winnie & Janzen, Daniel, 2015, Three new species of Ametadoria Townsend (Diptera: Tachinidae) from Area de Conservacion Guanacaste, Costa Rica, Biodiversity Data Journal 3, pp. 5039-5039 : 5039

Ametadoria leticiamartinezae Fleming & Wood sp. n.

Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018970 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018970; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 02-SRNP-11285; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Santa Elena ; verbatimLocality: Vado Quebrada Calera; verbatimElevation: 305; verbatimLatitude: 10.867; verbatimLongitude: -85.647; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.867; decimalLongitude: -85.647; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 02-Sep-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018981 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018981; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 96-SRNP-296; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Murcielago ; verbatimLocality: Pozo Del General; verbatimElevation: 40; verbatimLatitude: 10.898; verbatimLongitude: -85.731; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.898; decimalLongitude: -85.731; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 01-Jun-1996; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018973 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018973; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 02-SRNP-11281; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Santa Elena ; verbatimLocality: Vado Quebrada Calera; verbatimElevation: 305; verbatimLatitude: 10.867; verbatimLongitude: -85.647; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.867; decimalLongitude: -85.647; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 16-Aug-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018978 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018978; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 02-SRNP-11286; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Santa Elena ; verbatimLocality: Vado Quebrada Calera; verbatimElevation: 305; verbatimLatitude: 10.867; verbatimLongitude: -85.647; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.867; decimalLongitude: -85.647; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 24-Aug-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018979 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018979; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 96-SRNP-322; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Murcielago ; verbatimLocality: Pozo Del General; verbatimElevation: 40; verbatimLatitude: 10.898; verbatimLongitude: -85.731; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.898; decimalLongitude: -85.731; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 15-Jun-1996; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018980 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018980; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 96-SRNP-300; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Murcielago ; verbatimLocality: Pozo Del General; verbatimElevation: 40; verbatimLatitude: 10.898; verbatimLongitude: -85.731; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.898; decimalLongitude: -85.731; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 05-Jun-1996; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018971 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018971; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 02-SRNP-10747; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Santa Elena ; verbatimLocality: Vado Quebrada Calera; verbatimElevation: 305; verbatimLatitude: 10.867; verbatimLongitude: -85.647; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.867; decimalLongitude: -85.647; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 14-Oct-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018974 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018974; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 02-SRNP-11311; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Santa Elena ; verbatimLocality: Vado Quebrada Calera; verbatimElevation: 305; verbatimLatitude: 10.867; verbatimLongitude: -85.647; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.867; decimalLongitude: -85.647; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 18-Aug-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018982 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018982; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 96-SRNP-164; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Murcielago ; verbatimLocality: Pozo Del General; verbatimElevation: 40; verbatimLatitude: 10.898; verbatimLongitude: -85.731; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.898; decimalLongitude: -85.731; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 24-Feb-1996; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018977 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018977; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 02-SRNP-11297; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Santa Elena ; verbatimLocality: Vado Quebrada Calera; verbatimElevation: 305; verbatimLatitude: 10.867; verbatimLongitude: -85.647; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.867; decimalLongitude: -85.647; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 12-Aug-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018983 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018983; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 96-SRNP-136; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Murcielago ; verbatimLocality: Pozo Del General; verbatimElevation: 40; verbatimLatitude: 10.898; verbatimLongitude: -85.731; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.898; decimalLongitude: -85.731; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 11-Mar-1996; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018975 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018975; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 02-SRNP-11306; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Santa Elena ; verbatimLocality: Vado Quebrada Calera; verbatimElevation: 305; verbatimLatitude: 10.867; verbatimLongitude: -85.647; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.867; decimalLongitude: -85.647; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 26-Jul-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018976 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018976; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 02-SRNP-11312; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Santa Elena ; verbatimLocality: Vado Quebrada Calera; verbatimElevation: 305; verbatimLatitude: 10.867; verbatimLongitude: -85.647; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.867; decimalLongitude: -85.647; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 04-Aug-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018985 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018985; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 96-SRNP-317; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Murcielago ; verbatimLocality: Pozo Del General; verbatimElevation: 40; verbatimLatitude: 10.898; verbatimLongitude: -85.731; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.898; decimalLongitude: -85.731; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 31-May-1996; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018984 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018984; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 96-SRNP-158; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Murcielago ; verbatimLocality: Pozo Del General; verbatimElevation: 40; verbatimLatitude: 10.898; verbatimLongitude: -85.731; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.898; decimalLongitude: -85.731; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 10-Mar-1996; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018972 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros; individualID: DHJPAR0018972; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 02-SRNP-11328; Taxon: scientificName: Ametadoria leticiamartinezae; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Ametadoria; specificEpithet: leticiamartinezae; scientificNameAuthorship: Fleming & Wood, 2015; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservación Guanacaste; locality: Sector Santa Elena ; verbatimLocality: Vado Quebrada Calera; verbatimElevation: 305; verbatimLatitude: 10.867; verbatimLongitude: -85.647; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.867; decimalLongitude: -85.647; Identification: identifiedBy: Fleming & Wood; dateIdentified: 2014; Event: samplingProtocol: reared from caterpillar of Lactura subfervensDHJ01 (Lacturidae); verbatimEventDate: 13-Aug-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Description

Male (Fig. 3a, b, c). Head (Fig. 3c): frontal vitta dark black, narrowed apically to less than the width of the ocellar triangle, fronto-orbital plate as wide as frontal vitta; ocellar bristles well developed, proclinate, half as long as arista; frontal bristles extending below level of arista by width of one bristle; small black setulae intermingled with frontal bristles all along the parafrontal, not reaching below upper margin of pedicel; antenna black; arista black, minutely pubescent at its base, and bare at its tip; proclinate orbital bristles absent; parafrontal entirely silver; parafacial silver; palpus yellow and haired at its tip; gena 1/7 height of head. Thorax (Fig. 3a, b): gray when viewed dorsally with four longitudinal gray vittae, these only slightly visible post-suturally, appearing broken at thoracic suture; four post-sutural dorsocentral bristles; scutellum bearing silver tomentosity over its entirety (this tomentosity extending to underside of scutellum). Legs black; anteroventral surface of hind femur bearing 5-6 long erect bristles of irregular length and irregularly spaced; all surfaces of hind tibia with tightly adpressed, short, regularly spaced hairs, 1 bristle on each of the anterodorsal, anteroventral, posterodorsal and posteroventral surfaces of the tibia, 4X as long as the tibia is wide. Wing (Fig. 3a): pale smoky yellowish in color, calypters pale white translucent, slightly yellow and haired along their margins. Abdomen (Fig. 3a): ground color of abdominal tergites brown dorsally overall, dark velvety black medially with a bright gray band covering 3/4 or more of tergal surface. Tergites 3, 4 and 5 possessing median marginal bristles, but syntergite 1+2 without such bristles. Ventrolateral surface of syntergite 1+2, tg 3, and tg 4 orange. Sex patches present on ventral surface of tg 4, and tg 5. Terminalia (Figs 5b, 6b, 7b): sternite 5 with a deep and wide V-shaped median cleft, forming two rounded outer lobes, these covered in dense tomentum; lobes of sternite 5 bearing 5 long bristles marginally, the longest approximately 2X as long as other bristles present on sternite 5. Cerci almost flat when viewed laterally, narrow and finger-like in posterior view, not fused medially, maintaining an almost equidistant separation along their entire length; cerci covered in long hair dorsally, tapering at around 4/5 the length of the appendage, leaving only the apex bare. Surstylus short and truncate, approximately 2/3 as long as cerci, rounded and sub-triangular with a slight hook to the tip when viewed laterally; lateral surface of surstylus with several stout bristles confined to anterior apex, these being much stouter and thicker than any other bristles present on the genitalia. Epandrium, when viewed laterally, not extending backwards, 2/3 the length of cerci. Phallus short and stout, approximately same length as cerci. Postgonite 1/3 as long as phalloapodeme with no forward curve, and with a slightly lobed tip.

Female (Fig. 3d, e, f). Head (Fig. 3f): frontal vitta dark black, narrowed apically to less than the width of the ocellar triangle, fronto-orbital plate as wide as frontal vitta; frontal bristles not reaching below level of pedicel; 2 proclinate orbital bristles present; parafrontal entirely silver with minute setulae over its entire surface; parafacial silver; palpi orange and haired. Thorax (Fig. 3d, e): gray when viewed dorsally with four longitudinal gray vittae, these only slightly visible post-suturally, appearing broken at thoracic suture; scutellum bearing silver tomentosity over its entirety; legs and wings as in males. Abdomen (Fig. 3d): abdominal tergites dark velvety black medially, with a bright gray band covering 3/4 or more of tergal surface. Tergites 3, 4 and 5 possessing median marginal bristles, but syntergite 1+2 lacking such bristles.

Diagnosis

Fronto-orbital plate without golden tomentum, or golden tomentum present only in trace amounts and directly adjacent to the ocellar triangle; parafacial silver; calypters pale white translucent with yellow margins; hind tibia lacking a regularly spaced comb of hairs; instead, hairs on tibia are tightly adpressed with a few sparse long spines; abdominal tergites with black ground color bearing bright gray tomentose bands covering 3/4 or more of tergal surface; syntergite 1+2 lacking median marginal bristles. Epandrium when viewed laterally not extending backwards, 2/3 length of cerci. Phallus short and stout, approximately same length as cerci. Postgonite 1/3 as long as phalloapodeme with no forward curve, resembling a slightly lobed tip. Ametadoria leticiamartinezae can be differentiated from all other species of Ametadoria by the following combination of traits: the lack of golden tomentum on the fronto-orbital plate, the presence of orange ground color on the ventrolateral surface of the abdomen, and the gray tomentosity of the abdomen.

Etymology

Ametadoria leticiamartinezae is named in recognition of Ana Leticia Martínez Eras for her contributions to the accounting team for Area de Conservación Guanacaste, the forest this fly lives in.

Distribution

Costa Rica, ACG, Prov. Guanacaste, dry forest, 40-305 m elevation.

Ecology

Host: Ametadoria leticiamartinezae was reared from caterpillars of Lactura subfervens ( Lacturidae ) feeding on Sideroxylon spp. ( Sapotaceae ).