Sophiodela japonica ( Thunberg, 1781 ) Fukuda & Ogawa & Hori, 2019

Fukuda, Yuki, Ogawa, Ryo & Hori, Michio, 2019, The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia, Zootaxa 4661 (2), pp. 271-308 : 284

Sophiodela japonica ( Thunberg, 1781) , comb. nov.

( Figs 7B , 12B )

Cicindela japonica Thunberg, 1781: 25 .

Cicindela chinensis japonica: Fleutiaux, 1892:116 .

Cicindela chinensis japonica konoi Goto & Iga, 1950:14 [synonymy by Nakane 1976: 4].

Cicindela chinensis yakushimana Nakane, 1963:3 [synonymy by Nakane 1976: 4].

Cicindela (Sophiodela) japonica: Nakane 1955: 26 .

Cicindela (Sophiodela) chinensis japonica: Cassola & Nidek 1984:11 .

Cicindela (Sophiodela) japonica: Fukuda et al. 2015:271 .

Specimens examined. (N = 4) [ Japan] 2♂♂ ( EUMJ), Fruit Tree Research Center, Shimoidai, Matsuyama-shi , Ehime Pref., 2. X. 2018, R. Ogawa leg ; 2♂♂ ( EUMJ), ditto, 25. X. 2018, R. Ogawa leg.

Description. Internal sac everted dorsally in lateral view. DA largest in the bladders: DLL the second largest: VA as large as B. VLR, BLR and VB not exist.

VA short triangular everted dorso-apically. DA everted basally. B oval swollen ventrally, with well sclerotized spines, which are forming a narrow band. DLL swollen dorsally, covered numerously with well sclerotized spines, which are a forming band. Flagellum well short, curved and formed S shape. Medial tooth short, orthogonally curved right ventro-laterally.

Remarks. This species shares the characteristics of the position of each bladder with Sophiodela okinawana , S. ferriei , and S. c. chinensis , but is distinguished from the three species by the following characteristics: B oval swollen ventrally; VA triangular everted dorso-apically; medial tooth orthogonally curved.

Figures

FIGURE 7. Habitus of Sophiodela and Calochroa. A, Sophiodela chinensis chinensis (DeGeer) stat. nov.; B, Sophiodela japonica (Thunberg) stat. nov.; C, Sophiodela okinawana (Nakane) stat. nov.; D, Sophiodela ferriei (Fleutiaux) stat. nov.; E, Sophiodela chinensis flammifera (Horn) stat. nov.; F, Calochroa octonotata (Wiedemann); G, Calochroa carissima (Fleutiaux).

FIGURE 12. Everted internal sacs of Sophiodela. A, Sophiodela ferriei (Fleutiaux) stat. nov.; B, S. japonica (Thunberg) stat. nov.; C, S. okinawana (Nakane) stat. nov.; D, S. chinensis chinensis (DeGeer) stat. nov.; E, S. chinensis flammifera (Horn) stat. nov. Scales: 1.0 mm.