Philonthus costatus, Baudi, 1848

Taphylininae, Iochini, Aorothiini, Thiini, Latyprosopini, Taphylinini, Mblyopinina, Nisolinina, Yptiomina, Hilonthina & Herman, Lee H., 2001, Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina), Bulletin of the American Museum of Natural History 2441 (265), pp. 2435-3021 : 2792-2796

publication ID	https://doi.org/ 10.1206/0003-0090.265.1.5
persistent identifier	https://treatment.plazi.org/id/1D1E87B6-FEB7-FEED-FF00-4F39FD374990
treatment provided by	Marcus
scientific name	Philonthus costatus
status

Treatment

costatus Baudi, 1848 View in CoL , see: varians Paykull, 1789 .

couleensis Hatch, 1957: 185 ( Philonthus ; Type locality: Washington: Grand Coulee, Dry Falls). — Smetana, 1965: 25 ( Philonthus ; subgenus Philonthus ; characters; Miquelon; Nova Scotia; Newfoundland).

— Smetana, 1995e: 327 ( Philonthus ; characters; notes; Alberta; British Columbia; Manitoba;

New Brunswick; Newfoundland; Northwest Territories; Nova Scotia; Ontario; Quebec; Saskatchewan; Idaho; Illinois; Indiana; Massachusetts; Michigan; New Jersey; New York; Washington; Wisconsin).

- DISTRIBUTION: Canada, USA.

couloni Drugmand, 1987: 272 ( Philonthus ; subgenus Philonthus ; Type locality: Belgique: province de Brabant, Forest).

- DISTRIBUTION: Belgium.

coxalus Hochhuth 1872 , see: Gabrius .

coxatus Curtis, 1836 , see: laminatus Creutzer, 1799 .

coxatus Bernhauer, 1914 , see: Gabrius .

crassicornis Fauvel, 1895: 264 ( Philonthus ; Type locality: Bírmanie, Carin Cheba , 900–

1100 m; Tenasserim, Malewoon; Tonkin; Singapore; Java; Ceylan).

— Fauvel, 1904a: 59 ( Philonthus ; Nilghiris; Ceylan; Bírmanie; Tenasserim; Tonkin; Singapore; Sumatra; Java; Lombock; Maurice; Réunion).

— Fauvel, 1905e: 83 ( Philonthus ; Java).

— Bernhauer and Schubert, 1914: 333 ( Philonthus ; catalog).

— Cameron, 1921a: 377, 405 ( Philonthus ; characters; Singapore).

— Bernhauer, 1928: 16 ( Philonthus ; Sumatra).

— Cameron, 1932: 114 ( Philonthus ; characters; Ceylon; India).

— Scheerpeltz, 1933: 1338 ( Philonthus ; catalog).

— Cameron, 1933d: 387 ( Philonthus ; Sumatra).

— Jarrige, 1957: 109 ( Philonthus ; Réunion; also listed from Madagascar and Mauritius).

— Hayashi, 1996: 110 ( Philonthus ; Taiwan).

— Lecoq, 1990: 199 ( Philonthus ; Mascarene Islands).

- DISTRIBUTION: Madagascar, Mascarene Islands, Sri Lanka, India, Myanmar, Vietnam, Singapore, Indonesia, Taiwan.

crassulus Horn, 1884 , see: Gabrius .

crassulus Sharp, 1885 , see: alius Bernhauer and Schubert, 1914 .

crebrior Bernhauer, 1931 , see: donckieri Bernhauer, 1915 .

crestonensis Hatch, 1957 , see: Bisnius .

cribellatus Solsky, 1868 , see: Styngetus .

cribratus Erichson, 1840 , see: Orthidus .

cribrellus Fauvel, 1891: 112 ( Philonthus ; Type locality: Caracas ; Nouvelle Grenade).

— Bernhauer and Schubert, 1914: 334 ( Philonthus ; catalog).

- DISTRIBUTION: Venezuela.

cribricephalicus Tottenham, 1949b: 323 ( Philonthus ; Type locality: Mashonaland: Salisbury. Also cited from Natal ).

— Tottenham, 1962: 187 ( Philonthus ; characters).

- DISTRIBUTION: Zimbabwe, South Africa.

cribriceps Bernhauer, 1921c: 105 ( Philonthus ; Type locality: Brasilien: Petropolis ).

— Scheerpeltz, 1933: 1338 ( Philonthus ; catalog).

- DISTRIBUTION: Brazil.

cribricollis Erichson, 1840 , see: Paederomimus .

cribripennis Germain, 1903 , see: Loncovilius .

cribripennis Sharp, 1885: 403 ( Philonthus ; Type locality: Guatemala; Panama, Volcan de Chiriqui 2000 to 3000 feet).

- DISTRIBUTION: Guatemala, Panama.

cribriventris Bernhauer, 1912d: 43 ( Philonthus ; Type locality: Argentinien: Prov. Tucuman; Chaunar Region [Note: In a discussion of Neobisnius parcepunctatus, Frank, 1981: 12 , 13 thought this locality probably referred to Chañar Region, La Rioja]).

— Bernhauer and Schubert, 1914: 334 ( Philonthus ; catalog).

- DISTRIBUTION: Argentina.

crinitus Schubert, 1909: 293 ( Philonthus ; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d’Orizaba).

— Bernhauer and Schubert, 1914: 334 ( Philonthus ; catalog).

- DISTRIBUTION: Mexico.

crotchi Horn, 1884: 203 ( Philonthus ; Type locality: Vancouver; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Vancouver to Arizona; near Lake Tahoe ).

— Bernhauer and Schubert, 1914: 334 ( Philonthus ; catalog).

— Hatch, 1957: 186 ( Philonthus ; subspecies of lomatus ; characters; British Columbia; Washington; Oregon).

— Smetana, 1963a: 5 ( Philonthus ; aedeagus).

— Smetana, 1995e: 416 ( Philonthus ; lectotype designation: Van [ Vancouver ]; characters; notes; Alberta; British Columbia; Alaska; California; Montana; Nevada; Oregon; Washington) .

- DISTRIBUTION: Canada, USA.

nigritoides Smetana, 1964: 35 ( Philonthus ; subgenus Philonthus ; Type locality: Alaska: Kodiak Island).

— Smetana, 1965a: 7 ( Philonthus ; characters; Alaska).

— Smetana, 1995e: 416 ( Philonthus ; synonym of crotchi ).

cruentatus Gmelin, 1790: 2035 ( Staphylinus ; Type locality: Europa).

— Zschach, 1788: 44 ( Staphylinus ; species number 975; characters stated but not named and referred to as cruentatus by Gmelin).

— Panzer, 1795: 355 ( Staphylinus ; [Misidentification: Bernhauer and Schubert, 1914: 334 cited bipustulatus Panzer as a synonym of cruentatus Gmelin but Panzer did not describe bipustulatus as new, he attributed it to Fabricius who attributed it to Linné]; characters; Germany).

— Panzer, 1796c: 10 ( Staphylinus ; [Note: Misidentification: Bernhauer and Schubert, 1914: 334 cited bipustulatus Panzer as a synonym of cruentatus ; Panzer did not describe bipustulatus as new, he attributed it to Linné]; illustration; characters).

— Turton, 1802: 519 ( Staphylinus ; characters; Europe).

— Latreille, 1804: 321 ( Staphylinus ; cited as bipustulatus Panzer ; characters; Germany; France).

— Panzer, 1808: 8 ( Staphylinus ; [Note: Misidentification: Bernhauer and Schubert, 1914: 334 cited bipustulatus Panzer, 1795: 10 as a synonym of cruentatus Gmelin but Panzer did not describe bipustulatus as new, he attributed it to Fabricius who attributed it to Linné]; characters; illustration).

— Kraatz, 1857d: 601 ( Philonthus ; [Note: Cited as bipustulatus Panzer ]; characters; Germany).

— Thomson, 1860: 164 ( Philonthus ; [Note: Cited as bipustulatus Panzer ]; characters; Scandinavia).

— Fauvel, 1874: 253 [= 1874a: 479] ( Philonthus ; characters; France; Caucasus; Transcaucasia; Buchara; Lake Baikal).

— Seidlitz, 1875: 277 ( Philonthus ; cited as bipustulatus Panzer ; characters; Baltic region).

— Mulsant and Rey, 1876: 468 [= 1877: 325] ( Philonthus ; characters; France).

— J. Sahlberg, 1876: 20 ( Philonthus ; Finland).

— Fowler, 1888: 272 ( Philonthus ; characters; Britain).

— Eppelsheim, 1892: 334 ( Philonthus ; Turkestan).

— Ganglbauer, 1895: 454 ( Philonthus ; characters; Europe; Transcaucasian region; Siberia).

— Sainte-Claire Deville, 1906: 99 ( Philonthus ; Corsica).

— Reitter, 1909: 130 ( Philonthus ; characters; Germany).

— Bernhauer and Schubert, 1914: 334 ( Philonthus ; catalog).

— Johansen, 1914: 408 ( Philonthus ; characters; Denmark).

— Gridelli, 1920a: 18, 20 ( Philonthus ; characters; Italy).

— Portevin, 1929: 363 ( Philonthus ; characters; France).

— Cameron, 1932: 130 ( Philonthus ; characters; India).

— Scheerpeltz, 1933: 1338 ( Philonthus ; catalog).

— Wüsthoff, 1934: 61 ( Philonthus ; illustration of aedeagus).

— Voris, 1936 ( Philonthus ; rapid spread in North America ; habitat; New York; Michigan; Indiana; Kentucky; Tennessee; Illinois; Wisconsin; Minnesota; North Dakota; South Dakota; Nebraska; Missouri; Kansas; Colorado; Wyoming; Utah; Nevada; California; Idaho; Oregon; Washington; Ontario; Manitoba) .

— Uyttenboogaart, 1937: 82 ( Philonthus ; notes; Canary Islands).

— C. Koch, 1941: 63 ( Philonthus ; Morocco).

— Hatch, 1957: 189 ( Philonthus ; characters; British Columbia; Idaho; Washington; Oregon).

— Smetana, 1958: 194 ( Philonthus ; subgenus Philonthus ; characters; ecology; Czechoslovakia).

— Smetana, 1959: 157, 158 ( Philonthus ; characters).

— Scheerpeltz, 1963: 13 ( Philonthus ; Afghanistan).

— Palm, 1963b: 88 ( Philonthus ; characters; Sweden; Denmark; Norway; Finland).

— Lohse, 1964: 183 ( Philonthus ; subgenus Philonthus ; characters; central Europe).

— Focarile, 1964: 59 ( Philonthus ; Italy).

— Smetana, 1965: 16 ( Philonthus ; notes; Miquelon; St. Pierre; Newfoundland).

— Horion, 1965: 158 ( Philonthus ; Ireland; Britain; Holland; Belgium; France to Portugal and Spain; Corsica; Italy; Istria; Dalmatia; Bosnia; Serbia; Macedonia; Greece; France to Poland and Russia; Germany; Austria; Denmark; Sweden; Norway; Finland).

— Coiffait, 1967a: 419 ( Philonthus ; characters; Europe; Siberia).

— Raianu, 1968a: 466 ( Philonthus ; Romania).

— Szujecki, 1968a: 730 ( Philonthus ; Poland).

— Coiffait, 1970d: 143 ( Philonthus ; Uzbekistan).

— Coiffait, 1974: 241 ( Philonthus ; characters; Europe; Britain; Scandinavia; Siberia;

North Africa; Lebanon; Caucasus; India).

— Shilov, 1975a: 375 [= 1975b: 98] ( Philonthus ; Komi region of Russia).

— Pace and Zanetti, 1977: 295 ( Philonthus ; Italy).

— Pope, 1977: 29 ( Philonthus ; Britain).

— Hinton, 1981: 661 ( Philonthus ; characters of egg).

— Frank, 1982: 39 ( Philonthus ; list of parasites).

— Coiffait, 1983e: 477 ( Philonthus ; Afghanistan).

— Bordoni, 1984: 342 ( Philonthus ; Lebanon).

— Tóth, 1984: 70 ( Philonthus ; characters; Hungary).

— Lucht, 1987: 105 ( Philonthus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Nowosad, 1990: 105 ( Philonthus ; collected from mole nest; notes; Poland).

— Pilon and Zanetti, 1991: 59 ( Philonthus ; Italy).

— Hunter, Kincher, Bay, Beerwinkle, 1991: 168 ( Philonthus ; seasonal distribution and diel flight activity; Texas).

— Smetana, 1993b: 48 ( Philonthus ; subgenus Philonthus ; Czech Republic; Slovakia).

— Smetana, 1995e: 232 ( Philonthus ; characters; notes; Alberta; British Columbia; Manitoba;

New Brunswick; Newfoundland; Nova Scotia; Ontario; Prince Edward Island; Quebec; Saskatchewan; Alabama; Arizona; Arkansas; California; Colorado; Connecticut; Idaho; Illinois; Indiana; Iowa; Kansas; Kentucky; Maryland; Massachusetts; Michigan; Missouri; Montana; Nebraska; Nevada; New Hampshire; New Jersey; New Mexico; New York; North Carolina; North Dakota; Ohio; Oklahoma; Oregon; Pennsylvania; South Carolina; South Dakota; Tennessee; Texas; Vermont; Virginia; Washington; Wisconsin; Wyoming; Mexico).

— Ciceroni and Zanetti, 1995: 28 ( Philonthus ; Italy).

— Downie and Arnett, 1996: 378 ( Philonthus ; characters; USA).

— Schmidt, 1999: 108 ( Philonthus ; food for adults and larvae used for rearing species; rearing methods).

cunicularius Gistel, 1857: 80 ( Philonthus ; [Note: Nomen dubium]; Type locality: Bavaria). - DISTRIBUTION: Germany.

cupreipennis Cameron, 1926 , see: tractatus Eppelsheim, 1895 .

cupreonitens Eppelsheim, 1895a: 125 ( Philonthus ; Type locality: Gabon).

— Fauvel, 1903e: 241 ( Philonthus ; Cameroon; Gabon; Togo).

— Bernhauer and Schubert, 1914: 334 ( Philonthus ; catalog).

— Tottenham, 1949b: 332 ( Philonthus ; characters).

— Cameron, 1950g: 36 ( Philonthus ; Belgian Congo).

— Scheerpeltz, 1971c: 170 ( Philonthus ; Cameroon).

- DISTRIBUTION: Togo, Cameroon, Gabon, Congo.