Tolmerinus auronotatus, Fauvel, 1895

Taphylininae, Iochini, Aorothiini, Thiini, Latyprosopini, Taphylinini, Mblyopinina, Nisolinina, Yptiomina, Hilonthina & Herman, Lee H., 2001, Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina), Bulletin of the American Museum of Natural History 2441 (265), pp. 2435-3021 : 2508-2521

auronotatus Fauvel, 1895 , see: Tolmerinus .

bicolor Bernhauer, 1906 , see: Diatrechus .

birmanus Schillhammer, 1993b: 93 ( Anisolinus ; replacement name for malaisei Scheerpeltz, 1956: 275 ).

- DISTRIBUTION: Myanmar.

malaisei Scheerpeltz, 1965: 275 ( Amaurochlamys ; [preoccupied nec Scheerpeltz, 1965: 267]; Type locality: N.E. Burma, Kambaiti , 7000 ft.).

— Schillhammer, 1999b: 93 ( Anisolinus ; synonym of birmanus ).

caeruleipennis Cameron, 1932: 250 ( Craspedomerus ; Type locality: Sikkim: Rungbong Valley). — Scheerpeltz, 1933: 1410 ( Craspedomerus ; catalog).

— Scheerpeltz, 1976a: 148 ( Craspedomerus ; characters).

— Coiffait, 1984e: 374 ( Craspedomerus ; cited as coeruleipennis ; Nepal).

— Schillhammer, 1999b: 94 ( Anisolinus ; notes).

- DISTRIBUTION: India, Nepal.

elegans Sharp, 1889: 114 ( Anisolinus ; Type locality: Miyanoshita).

— Bernhauer and Schubert, 1914: 365 ( Anisolinus ; catalog).

— Nakane, 1963a: 92 ( Anisolinus ; characters; habitus photograph; Japan).

— Naomi, 1981: 110, 111 ( Anisolinus ; characters; Japan).

— Naomi, 1991: 2 ( Anisolinus ; characters; Japan).

— Hayashi, 1993b: 284 ( Anisolinus ; Japan).

- DISTRIBUTION: Japan.

fasciiventris Fauvel, 1895 , see: Tolmerinus .

femoratus Schubert, 1911 , see: Diatrechus .

hayashii Sawada, 1961: 4 ( Anisolinus ; Type locality: Japan: Honshu: Mt. Koya, Wakayama). — Naomi, 1981: 110, 111 ( Anisolinus ; characters; Japan).

— Naomi, 1991: 5 ( Anisolinus ; characters; Japan).

— Hayashi, 1993b: 284 ( Anisolinus ; Japan).

- DISTRIBUTION: Japan.

humeralis Fauvel, 1907 , see: Diatrechus .

kawanoi Sawada, 1963 , see: tsurugiensis Sawada, 1961 .

malaisei Scheerpeltz, 1965: 267 ( Hesperodes ; Type locality: N.E. Burma, Kambaiti, 1800 m). — Lundgren, 1987: 14 ( Blackwelderella ).

— Schillhammer, 1999b: 93 ( Anisolinus ).

- DISTRIBUTION: Myanmar.

malaisei Scheerpeltz, 1965: 275 , see: birmanus Schillhammer, 1999 .

nitidus Scheerpeltz, 1965: 269 ( Hesperodes ; Type locality: N.E. Burma, Kambaiti, 2000 m). — Lundgren, 1987: 14 ( Blackwelderella ).

— Schillhammer, 1999b: 93 ( Anisolinus ).

- DISTRIBUTION: Myanmar.

picticornis Sharp, 1889: 113 ( Anisolinus ; Type locality: Miyanoshita).

— Bernhauer and Schubert, 1914: 365 ( Anisolinus ; catalog).

— Nakane, 1963a: 92 ( Anisolinus ; characters; habitus photograph; Japan).

— Naomi, 1981: 110, 112 ( Anisolinus ; characters; Japan).

— Naomi, 1991: 6 ( Anisolinus ; characters; Japan).

— Hayashi, 1993b: 284 ( Anisolinus ; Japan).

— Watanabe, 1996a: 13 ( Anisolinus ; cited as pilicornis ; cave collection; Japan).

- DISTRIBUTION: Japan.

raffrayi Fauvel, 1905 , see: Diatrechus .

rougemonti Naomi, 1993: 185 ( Anisolinus ; Type locality: Nepal: Sawa , 2500 m).

- DISTRIBUTION: Nepal.

rufipennis Fauvel, 1895 , see: Tolmerinus .

taoi Naomi, 1981: 113 ( Anisolinus ; Type locality: Japan: Yamanashi Pref.: Daibosatsu-pass). — Naomi, 1991: 5 ( Anisolinus ; characters; Japan).

— Hayashi, 1993b: 284 ( Anisolinus ; Japan).

- DISTRIBUTION: Japan.

tsurugiensis Sawada, 1961: 5 ( Anisolinus ; Type locality: Japan: Shikoku : Tokushima,

Mt. Tsurugi).

— Nakane, 1963a: 92 ( Anisolinus ; [Note: Misidentified as kawanoi Sawada according to Shibata, 1984: 109]; characters; habitus photograph; Japan).

— Naomi, 1981: 110, 112 ( Anisolinus ; characters; Japan).

— Naomi, 1991: 5 ( Anisolinus ; characters; Japan).

- DISTRIBUTION: Japan.

kawanoi Sawada, 1963: 92 ( Anisolinus ; Type locality: Shikoku).

— Shibata, 1984: 109 ( Anisolinus ; synonym of tsurugiensis but stated to be misidentification). wittmeri Coiffait, 1982a: 269 ( Philonthopsis ; [preoccupied]; Type locality: Nepal: Thodung via Those, 3100 m).

— Schillhammer, 1994: 74 ( Philonthopsis ; species to be placed in another genus, perhaps Anisolinus , later).

— Schillhammer, 1999b: 93 ( Anisolinus ; notes).

- DISTRIBUTION: Nepal.

Hesperosoma

[3 species; Palaearctic and Oriental Regions]

Hesperosoma Scheerpeltz, 1965: 270 (species included: malaisei ). TYPE SPECIES: Hesperosoma malaisei Scheerpeltz , fixed by original designation and monotypy.

— Hayashi, 1993b: 290 (characters).

— Hayashi, 2000: 24 (notes on empodial setae).

Paramichrotus Naomi, 1982a: 37 , 38 (subgenus of Amichrotus ; species included: miwai ). TYPE SPECIES: Amichrotus miwai Bernhauer , fixed by original designation and monotypy.

— Hayashi, 1993b: 290 (synonym of Hesperosoma ).

malaisei Scheerpeltz, 1965: 271 ( Hesperosoma ; Type locality: N.E. Burma, Kambaiti , 7000 ft.).

- DISTRIBUTION: Myanmar.

miwai Bernhauer, 1943a: 177 ( Amichrotus ; Type locality: Formosa: Arisan ).

— Shibata, 1976a: 10 ( Amichrotus ; characters; Taiwan).

— Naomi, 1982a: 38 ( Amichrotus ; subgenus Paramichrotus ; characters; Taiwan).

— Hayashi, 1993b: 290 ( Hesperosoma ; characters; Taiwan).

- DISTRIBUTION: Taiwan.

nanshanchiana Hayashi, 1993a: 123 ( Hesperosoma ; subspecies of miwai ; Type locality: Taiwan: Nanshanchi, Nantou Hsien).

nanshanchiana Hayashi, 1993 , see: miwai Bernhauer, 1943 .

sakoi Hayashi, 1993a: 124 ( Hesperosoma ; Type locality: Taiwan: Fungchiifo , Chiai Hsien).

- DISTRIBUTION: Taiwan.

Misantlius

[7 species; Neotropical Region]

Misantlius Sharp, 1885: 393 (species included: optatus , torquatus , carinulatus , discedens ). TYPE SPECIES: Misantlius carinulatus Sharp , fixed by subsequent designation by R. Lucas, 1920: 420.

— Bernhauer and Schubert, 1914: 367 (world catalog; 6 species).

— Blackwelder, 1944: 136 (checklist of species from Mexico, Central and South America, and the West Indies).

— Blackwelder, 1952: 249 (type species).

carinulatus Sharp, 1885: 394 ( Misantlius ; Type locality: Mexico; Toxpam ).

— Bernhauer and Schubert, 1914: 367 ( Misantlius ; catalog).

- DISTRIBUTION: Mexico.

discedens Sharp, 1885: 395 ( Misantlius ; Type locality: Mexico, Yolotepec ).

— Bernhauer and Schubert, 1914: 367 ( Misantlius ; catalog).

- DISTRIBUTION: Mexico.

gebieni Bernhauer, 1942: 26 ( Misantlius ; Type locality: Kostarika: Vara Blanca, 1700 m).

- DISTRIBUTION: Costa Rica.

jalappae Bernhauer, 1910b: 378 ( Misantlius ; Type locality: Mexiko: Jalapa ).

— Bernhauer and Schubert, 1914: 367 ( Misantlius ; catalog).

- DISTRIBUTION: Mexico.

optatus Sharp, 1885: 394 ( Misantlius ; Type locality: Mexico; Misantla ).

— Bernhauer and Schubert, 1914: 367 ( Misantlius ; catalog).

- DISTRIBUTION: Mexico.

rufipennis Sharp, 1887: 786 ( Misantlius ; Type locality: Mexico, Yolotepec , Juquila).

— Bernhauer and Schubert, 1914: 367 ( Misantlius ; catalog).

- DISTRIBUTION: Mexico.

torquatus Sharp, 1885: 394 ( Misantlius ; Type locality: Guatemala, Calderas ).

— Bernhauer and Schubert, 1914: 367 ( Misantlius ; catalog).

- DISTRIBUTION: Guatemala.

Montguillonius

[1 species; Palaearctic Region]

Montguillonius Coiffait, 1987: 497 (replacement name for Hesperopsis Coiffait ). TYPE SPECIES: Hesperopsis sikkimensis Coiffait , fixed by objective synonymy with Hesperopsis for which a type had been fixed.

Hesperopsis Coiffait, 1982a: 268 (species included: sikkimensis ; preoccupied by Dyar, 1905). TYPE SPECIES: Hesperopsis sikkimensis Coiffait , fixed by original designation and monotypy.

— Coiffait, 1987: 497 (synonym of Montguillonius ).

sikkimensis Coiffait, 1982a: 268 ( Hesperopsis ; Type locality: Sikkim: Dzongri , 3000 m).

- DISTRIBUTION: India.

Philomyceta

[2 species; Palaearctic Region]

Philomyceta Cameron, 1944: 12 (species included: caeruleipennis , affinis ). TYPE SPECIES: Philomyceta caeruleipennis Cameron , by original designation.

— Blackwelder, 1952: 301 (type species: caeruleipennis ).

affinis Cameron, 1944: 13 ( Philomyceta ; Type locality: Darjeeling: Ghum district).

- DISTRIBUTION: India.

caeruleipennis Cameron, 1944: 12 ( Philomyceta ; Type locality: Darjeeling: Ghum district).

- DISTRIBUTION: India.

Subtribe Hyptiomina

Hyptiomina Casey, 1906 .

— Newton and Thayer, 1992: 65 (subtribe of Staphylinini ).

— Smetana, 1995e: 48 (characters).

— Newton, Thayer, Ashe, and Chandler, 2000: 397 (characters; notes).

Hyptiomae Casey, 1906: 359, 361 (genera included: Hyptioma ). TYPE GENUS: Hyptioma

(see Newton and Thayer, 1992: 65).

atratulus Sharp, 1885: 464 ( Holisus ; Type locality: Guatemala, El Tumbador 2500 feet, Capetillo).

— Bernhauer and Schubert, 1914: 321 ( Holisus ; catalog).

- DISTRIBUTION: Guatemala.

cingulatus Cameron, 1918 , see: Holisomimus .

collaris Baudi, 1848: 136 ( Lithocharis ; Type locality: Venezuela).

— Kraatz, 1857b: 176 ( Holisus ; transferred from Lithocharis ).

— Fauvel, 1891: 119 ( Holisus ; Venezuela).

— Bernhauer and Schubert, 1914: 321 ( Holisus ; catalog).

- DISTRIBUTION: Venezuela.

condei Wendeler, 1956a: 229 ( Holisus ; Type locality: Brasilien: Espirito Santo, Leopoldina Col., Rio Bonito ).

- DISTRIBUTION: Brazil.

confinis Sharp, 1885: 462 ( Holisus ; Type locality: Panama, Bugaba ).

— Bernhauer and Schubert, 1914: 321 ( Holisus ; catalog).

- DISTRIBUTION: Panama.

cubensis Casey, 1906 , see: debilis Erichson, 1839 .

debilis Erichson, 1839b: 299 ( Holisus ; Type locality: Americae meridionalis ins. St. Johannis). — Bernhauer and Schubert, 1914: 321 ( Holisus ; catalog).

— Blackwelder, 1943: 461 ( Holisus ; characters; Bahamas; Cuba; Haiti; Puerto Rico; St John;

St. Lucia; St. Vincent).

- DISTRIBUTION: West Indies.

cubensis Casey, 1906: 362 ( Hyptioma ; Type locality: Cuba, Havana).

— Bernhauer and Schubert, 1914: 320 ( Hyptioma ; catalog).

— Scheerpeltz, 1933: 1326 ( Holisus ; catalog).

— Blackwelder, 1943: 461 ( Holisus ; synonym of debilis ).

rufoniger Cameron, 1922: 117 ( Holisus ; Type locality: Haiti; St. Vincent).

— Scheerpeltz, 1933: 1326 ( Holisus ; catalog).

— Blackwelder, 1943: 461 ( Holisus ; synonym of debilis ).

depressus Sharp, 1876: 180 ( Holisus ; Type locality: Ega).

— Bernhauer and Schubert, 1914: 321 ( Holisus ; catalog).

- DISTRIBUTION: Brazil.

discedens Sharp, 1876: 183 ( Holisus ; Type locality: Not cited).

— Bernhauer and Schubert, 1914: 321 ( Holisus ; catalog).

- DISTRIBUTION: Brazil.

elatus Sharp, 1885: 465 ( Holisus ; Type locality: Guatemala, San Geronimo; Panama, Bugaba, Volcan de Chiriqui).

— Bernhauer and Schubert, 1914: 322 ( Holisus ; catalog).

- DISTRIBUTION: Guatemala, Panama.

eleusoides Sharp, 1885: 466 ( Holisus ; Type locality: Guatemala, Las Mercedes 3000 feet, Cerro Zunil 4000–5000 feet).

— Bernhauer and Schubert, 1914: 322 ( Holisus ; catalog).

- DISTRIBUTION: Guatemala.

excavatus Sharp, 1876: 182 ( Holisus ; Type locality: Ega).

— Bernhauer and Schubert, 1914: 322 ( Holisus ; catalog).

- DISTRIBUTION: Brazil.

filum Sharp, 1885: 464 ( Holisus ; Type locality: Guatemala, Teleman in the Polochic Valley).

— Bernhauer and Schubert, 1914: 322 ( Holisus ; catalog).

- DISTRIBUTION: Guatemala.

guildingii Erichson, 1839b: 300 ( Holisus ; Type locality: Americae meridionalis ins.

St. Vincentii).

solskyi Sharp, 1885: 463 ( Holisus ; Type locality: Mexico, Cordova ).

— Solsky, 1868a: 141 (Holius; [Note: Misidentification: Bernhauer and Schubert, 1914: 322, cited ater Solsky as a synonym of solskyi ; Solsky did not describe the species as new, he attributed it to Motschulsky]; Mexico).

— Bernhauer and Schubert, 1914: 322 ( Holisus ; catalog).

- DISTRIBUTION: Mexico.

sparsiceps Bernhauer, 1942: 24 ( Holisus ; Type locality: Kostarika).

- DISTRIBUTION: Costa Rica.

umbra Sharp, 1876: 182 ( Holisus ; Type locality: Ega).

— Bernhauer and Schubert, 1914: 322 ( Holisus ; catalog).

- DISTRIBUTION: Brazil.

Subtribe Philonthina

Philonthina Kirby, 1837 .

— Lohse, 1964: 170 (characters; key to central European species).

— Dettner, 1983 (comparative anatomy of defensive glands and defensive secretions).

— Dettner, 1987: 30 (defensive glands).

— Newton and Thayer, 1992: 65 (subtribe of Staphylinini ).

— Dettner, 1993a (summary and discussion of defensive secretions and exocrine glands and their bearing on phylogeny).

— Smetana, 1995e (characters; revision of and key to species of USA and Canada; key to holarctic genera; revision of genera of USA and Canada; phylogeny; historical discussion; natural history; habitat; life cycle; immature stages; fossils).

— Klimaszewski, Newton, and Thayer, 1996: 151 (list of New Zealand genera; number of

New Zealand species).

— Smetana and Davies, 2000: 12, 13 (characters).

— Newton, Thayer, Ashe, and Chandler, 2000: 332, 395 (key to Nearctic genera; characters; notes).

— Hayashi, 2000: 23 (notes on empodial setae).

Philonthidae Kirby, 1837: 91 (genera included: Philonthus ). TYPE GENUS: Philonthus

(see Newton and Thayer, 1992: 65).

Rémates Mulsant and Rey, 1877: 452 (genera included: Remus ). TYPE GENUS: Remus

(see Newton and Thayer, 1992: 65).

— Newton and Thayer, 1992: 65 (synonym of Philonthina ).

Craspedomerina Bernhauer, 1911.

— Newton and Thayer, 1992: 65 (subtribe of Staphylinini ).

— Smetana, 1995e: 48 (characters).

— Smetana and Davies, 2000: 12 (synonym of Philonthina ).

Craspedomeri Bernhauer, 1911: 88 (genera included: Craspedomerus ). TYPE GENUS: Craspedomerus (see Newton and Thayer, 1992: 65).

— Cameron, 1932: 54, 248 (characters).

— Scheerpeltz, 1933: 1410 (world catalog supplement).

— Scheerpeltz, 1940: 47 (key to Palaearctic genera).

— Moore, 1960: 100 (key to genera).

Philonthini Coiffait, 1956e: 183 (genera included: none cited). TYPE GENUS: Philonthus (see Newton and Thayer, 1992: 65).

— Coiffait, 1963: 6 (characters; key to genera).

— Pototskaia, 1966: 152 (larval characters; key to some genera using larval characters).

— Pototskaia, 1967: 52 (larval characters).

— Coiffait and Saiz, 1968: 347 (key to Chilean genera).

— Scheerpeltz, 1968a: 64 (catalog of Austrian species).

— Coiffait, 1974: 2 (characters; key to genera and species of western Palaearctic region).

manikaensis Levasseur, 1967b: 1628 ( Afrorabigus ; Type locality: région de Kolwezi, Katanga, Manika).

- DISTRIBUTION: Congo.

muscicola Levasseur, 1965: 60 ( Afrorabigus ; Type locality: Katanga: Kolwezi ).

- DISTRIBUTION: Congo.

robusticornis Bernhauer, 1915d: 137 ( Philonthus ; Type locality: Deutsch-Ostafrika: Arusha-Ju; Moshi ).

— Scheerpeltz, 1933: 1358 ( Philonthus ; catalog).

— Bernhauer and Chapman, 1939: 68 ( Philonthus ; Kenya).

— Tottenham, 1949b: 328 ( Philonthus ; characters).

— Tottenham, 1962: 137 ( Philonthus ; Tanganyika).

— Levasseur, 1965: 62 ( Afrorabigus ; characters; Zaire; Kenya; Sudan; Kilimanjaro).

- DISTRIBUTION: Sudan, Kenya, Tanzania, Congo.

toumodiensis Levasseur, 1967b: 1629 ( Afrorabigus ; Type locality: Côte d’Ivoire, Lamto , Toumodi).

- DISTRIBUTION: Ivory Coast.

Agacerus

[1 species; Oriental Region]

Agacerus Fauvel, 1895: 245 (replacement name for Eurycerus Fauvel ). TYPE SPECIES: Agacerus pectinatus Fauvel , fixed by objective synonymy with Eurycerus .

— Bernhauer and Schubert, 1914: 396 (world catalog; 1 species).

— Cameron, 1925: 71 (catalog of Indian species).

— Cameron, 1932: 255 (characters).

— Scheerpeltz, 1933: 1411 (world catalog supplement).

— Blackwelder, 1952: 40 (type species: pectinatus ).

— Scheerpeltz, 1965: 278 (characters; type species: pectinatus ).

Eurycerus Fauvel, 1895: 244 (species included: pectinatus ; preoccupied by Illiger, 1807, Dejean, 1833, and Kaup, 1844; [Note: Eurycerus was validated on page 244 and replaced on page 245. It might be argued that Eurycerus was originally published as

a junior synonym and is therefore an unavailable name (Article 11.6). However, since it was published on one page and synonymized on the next, it can be argued that the name was not originally published as a synonym]). TYPE SPECIES: Agacerus pectinatus Fauvel , fixed by monotypy.

— Bernhauer and Schubert, 1914: 396 (synonym of Agacerus ).

— Cameron, 1932: 255 (synonym of Agacerus ).

— Blackwelder, 1952: 158 (type species).

pectinatus Fauvel, 1895: 245 ( Agacerus ; Type locality: Bírmanie, Carin Cheba , 900–1100 m).

— Bernhauer and Schubert, 1914: 396 ( Agacerus ; catalog).

— Cameron, 1932: 256 ( Agacerus ; characters; Burma).

— Scheerpeltz, 1933: 1411 ( Agacerus ; catalog).

- DISTRIBUTION: Myanmar.

Atopocentrum

[1 species; Neotropical]

Atopocentrum Bernhauer, 1906c: 327 (species included: mirabile ). TYPE SPECIES: Atopocentrum mirabile Bernhauer , fixed by monotypy.

— Bernhauer and Schubert, 1914: 365 (world catalog; 1 species).

— Blackwelder, 1944: 138 (in checklist of species from Mexico, Central and South America,

and the West Indies).

— Blackwelder, 1952: 69 (type species: mirabile ).

mirabile Bernhauer, 1906c: 327 ( Atopocentrum ; Type locality: Peru: Cushi , 1800 m).

— Bernhauer and Schubert, 1914: 365 ( Atopocentrum ; catalog).

- DISTRIBUTION: Peru.

Although Motschulsky used the name Trapezidera in Helotidae in 1860, the names are not homonyms]). TYPE SPECIES: Trapeziderus bicolor Motschulsky , fixed by objective synonymy with Trapeziderus .

— Fauvel, 1895: 266 (synonym of Belonuchus ).

— Blackwelder, 1952: 392 (type species).

Musicoderus Sharp, 1885: 455 (species included: cephalotes , gracilis ). TYPE SPECIES: Musicoderus cephalotes Sharp , fixed by subsequent designation by Blackwelder, 1943: 420 [Note: Blackwelder, 1952: 251, stated that R. Lucas, 1920: 425, designated the type species but the designation was ambiguous because two species were cited as “ cephalotes et gracilis ”].

— Bernhauer and Schubert, 1914: 369 (world catalog; 3 species).

— Scheerpeltz, 1933: 1377 (subgenus of Belonuchus ).

— Blackwelder, 1943: 420 (synonym of Belonuchus ; type species: cephalotes ).

— Blackwelder, 1952: 251 (type species).

— Smetana, 1995e: 722 (synonym of Belonuchus ).

abdominalis Cameron, 1937: 20 ( Belonuchus ; Type locality: Java: G. Tangkoeban Prahoe,

alt. 4000–5000 ft).

- DISTRIBUTION: Indonesia.

abyssinus Fauvel, 1881 , see: Philonthus .

aeneipennis Fauvel, 1895: 268 ( Belonuchus ; Type locality: Nord-ouest de Sumatra , Tebingtinggi ; Tandjong Morawa , Serdang; Borneo, Sarawak; Malacca; Singapore).

— Fauvel, 1904a: 60 ( Belonuchus ; Nilghiris; Bírmanie; Malacca; Singapore; Borneo; Sumatra).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

— Cameron, 1921a: 378, 405 ( Belonuchus ; characters; Singapore).

— Scheerpeltz, 1933: 1377 ( Belonuchus ; catalog).

- DISTRIBUTION: Indonesia, Singapore, Borneo, Malaysia, Myanmar, India.

aeneiventris Fauvel, 1878f: 270 ( Belonuchus ; Type locality: Nouvelle-Guinée, Andai,

Ile Yule; Ternate, Acquiconora).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: New Guinea, Indonesia.

aequalis Sharp, 1876: 161 ( Belonuchus ; Type locality: Para, Ega, St. Paulo; Conceicao ,

Rio Mauhes).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: Brazil.

agilis Erichson, 1840: 423 ( Belonuchus ; Type locality: Cuba).

— Guérin-Méneville, 1857: 37 ( Belonuchus ; characters).

— Chevrolat and Fauvel, 1863: 433 ( Belonuchus ; characters; Cuba).

— Fauvel, 1901: 87 ( Belonuchus ; Jamaica; Haiti; Cuba; Colombia).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

— Blackwelder, 1943: 427 ( Belonuchus ; characters; Cuba; Jamaica).

- DISTRIBUTION: West Indies, Colombia.

chevrolati Fauvel, 1863: 432 ( Belonuchus ; Type locality: La Havane, Cuba).

— Fauvel, 1869: 489 ( Belonuchus ; synonym of agilis ).

— Fauvel, 1901: 87 ( Belonuchus ; synonym of agilis ).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; synonym of agilis ).

— Blackwelder, 1943: 427 ( Belonuchus ; synonym of agilis ).

albovariegatus Bernhauer, 1916 g: 276 ( Belonuchus ; Type locality: West-Columbien: Umgebung Cali am Rio Cauca: St. Antonio , 2000 m; Alto de las cruces, 2200–2400 m).

— Scheerpeltz, 1933: 1377 ( Belonuchus ; catalog).

- DISTRIBUTION: Colombia.

alternatus Sharp, 1885: 430 ( Belonuchus ; Type locality: Guatemala, near the City ,

San Geronimo; Nicaragua, Chontales; Panama, Bugaba).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: Panama, Nicaragua, Guatemala.

altissimus Bernhauer, 1916 g: 275 ( Belonuchus ; Type locality: West-Columbien: Am Monte Socorro in den West-Cordilleren, 3800 m).

— Scheerpeltz, 1933: 1377 ( Belonuchus ; catalog).

- DISTRIBUTION: Colombia.

altivagans Bernhauer, 1916 g: 272 ( Belonuchus ; Type locality: West-Columbien: Pacho in den Ost-Cordilleren, 2000 m).

— Scheerpeltz, 1933: 1377 ( Belonuchus ; catalog).

- DISTRIBUTION: Colombia.

amplipennis Fauvel, 1891: 117 ( Belonuchus ; Type locality: Colonie Tovar; San Esteban).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: Venezuela.

amplus Blackwelder, 1943: 425 ( Belonuchus ; Type locality: St. Lucia: one-half mile northwest of St. Remy on road to Soufriere, 2.5 miles southeast of Soufriere).

- DISTRIBUTION: West Indies.

analis Schubert, 1911: 29 ( Belonuchus ; Type locality: Lombok).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: Indonesia.

andinus Bernhauer, 1916 g: 278 ( Belonuchus ; Type locality: Bolivien: Yuracares ).

— Scheerpeltz, 1933: 1377 ( Belonuchus ; catalog).

- DISTRIBUTION: Bolivia.

angusticeps Scheerpeltz, 1960c: 117 ( Belonuchus ; subgenus Musicoderus ; Type locality: Bolivien: Cochabamba, Yungas de Palmar, 2000 m).

- DISTRIBUTION: Bolivia.

angusticollis Bernhauer, 1906c: 330 ( Belonuchus ; Type locality: Peru: Chanchamayo ).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: Peru.

anthracinus Schubert, 1911: 30 ( Belonuchus ; Type locality: Transvaal, Zoutpansberg, 800 m). — Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: South Africa.

antiguae Blackwelder, 1943: 434 ( Belonuchus ; Type locality: Antigua: near summit of

Boggy Peak, parish of St. Mary, elevation 1300 feet).

- DISTRIBUTION: West Indies.

aphaobius Smetana, 1995e: 740 ( Belonuchus ; Type locality: Texas: Austin ).

- DISTRIBUTION: USA.

apicicornis Bernhauer, 1916 g: 270 ( Belonuchus ; subgenus Musicoderus ; Type locality: West-Columbien: Cali am Rio Cauca: Rio Aquatal, 1800 m; S. Antonio, 2000 m;

Rio Vitaco, 2200 m).

— Scheerpeltz, 1933: 1377 ( Belonuchus ; catalog).

- DISTRIBUTION: Colombia.

apiciventris Sharp, 1885: 419 ( Philonthus ; Type locality: Mexico, Juquila ).

— Bernhauer and Schubert, 1914: 329 ( Philonthus ; catalog).

— Navarrete-Heredia and Márquez-Luna, 1993 ( Philonthus ; cave collections; Mexico).

— Navarrete-Heredia, 1995: 84, 89, 100 ( Philonthus ; cited as apaciventris; characters; habits; natural history notes; host mushrooms; Mexico).

— Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 61 ( Belonuchus ; Mexico).

- DISTRIBUTION: Mexico.

argentinus Bernhauer, 1916 g: 278 ( Belonuchus ; Type locality: Argentinien: Misiones ; Brasilien: Espirito Santo, Rio de Janeiro).

— Scheerpeltz, 1933: 1377 ( Belonuchus ; catalog).

- DISTRIBUTION: Argentina, Brazil.

arizonicus Casey, 1915: 448 ( Belonuchus ; Type locality: Arizona: Benson and Sta. Rita Mts.; [Note: See lectotype designation by Smetana, 1995e]).

— Smetana, 1995e: 743 ( Belonuchus ; lectotype designation: Arizona: Benson and Sta. Rita Mts.; characters; notes; Arizona; California; New Mexico; Texas) .

- DISTRIBUTION: USA.

texanus Casey, 1915: 447 ( Belonuchus ; Type locality: Texas: Brownsville; [Note: See lectotype designation by Smetana, 1995e]).

— Scheerpeltz, 1933: 1379 ( Belonuchus ; catalog).

— Smetana, 1995e: 743 ( Belonuchus ; lectotype designation; synonym of arizonicus ).

armatus Sharp, 1876: 163 ( Belonuchus ; Type locality: Abacaxis).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: Brazil.

arrogans Bernhauer, 1916 g: 274 ( Belonuchus ; Type locality: West-Columbien: Ost- Cordilleren, Pacho, 2000 m).

— Scheerpeltz, 1933: 1377 ( Belonuchus ; catalog).

- DISTRIBUTION: Colombia.

asperatus Bernhauer, 1906c: 336 ( Belonuchus ; Type locality: Peru: Pachitea ).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

— Scheerpeltz, 1960c: 120 ( Belonuchus ; subgenus Musicoderus ; Bolivia).

- DISTRIBUTION: Peru, Bolivia.

assamensis Cameron, 1932: 176 ( Belonuchus ; Type locality: Assam: Naga Hills ).

— Scheerpeltz, 1933: 1377 ( Belonuchus ; catalog).

- DISTRIBUTION: India.

associatus Cameron, 1938b: 150 ( Belonuchus ; Type locality: Not cited, presumably Java).

- DISTRIBUTION: Indonesia.

morosus Cameron, 1938b: 150 ( Belonuchus ; variety of associatus ; Type locality: E. Java: Res. Kediri, Toempoek).

avidus Erichson, 1840: 423 ( Belonuchus ; Type locality: Novae Valenciae in Columbia).

— Fauvel, 1891: 117 ( Belonuchus ; Venezuela).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: Colombia, Venezuela.

bafutensis Levasseur, 1968: 1381 ( Belonuchus ; Type locality: Cameroun: forêt de Bafut, Nguemba).

- DISTRIBUTION: Cameroon.

bakeri Bernhauer, 1915a: 25 ( Belonuchus ; Type locality: Philippinen: Mts. Banahao ).

— Scheerpeltz, 1933: 1377 ( Belonuchus ; catalog).

— Jarrige, 1951: 134 ( Belonuchus ; Tonkin).

- DISTRIBUTION: Philippines, Vietnam.

barbicornis Bernhauer, 1916 g: 271 ( Belonuchus ; subgenus Musicoderus ; Type locality: West-Columbien: Alto de las Cruces, 2200 m; S. Antonio, 2200 m).

— Scheerpeltz, 1933: 1377 ( Belonuchus ; catalog).

- DISTRIBUTION: Colombia.

basiventris Sharp, 1885: 418 ( Philonthus ; Type locality: Mexico, Cordova, Puebla Yolos, Oaxaca, Jalapa ).

— Bernhauer and Schubert, 1914: 330 ( Philonthus ; catalog).

— Marquez-Luna and Navarrete Heredia, 1995: 35 ( Philonthus ; in detritus of Atta mexicana ; Mexico).

— Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 61 ( Belonuchus ; necrophilous species; distribution in Mexico).

- DISTRIBUTION: Mexico.

batesi Sharp, 1876: 157 ( Belonuchus ; Type locality: Ega).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: Brazil.

bicolor Motschulsky, 1860 , see: grandiceps Kraatz, 1859 .

bicoloratus Schubert, 1911: 30 ( Belonuchus ; Type locality: Lombok).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: Indonesia.

bicoloripennis Bernhauer, 1928 g: 121 ( Belonuchus ; Type locality: Belg.- Kongo: Haute Uele : Moto).

— Scheerpeltz, 1933: 1377 ( Belonuchus ; catalog).

- DISTRIBUTION: Congo.

bidens Sharp, 1885: 430 ( Belonuchus ; Type locality: Mexico, Jalapa, Oaxaca, Mirador, Tuxtla ; Nicaragua, Chontales).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: Nicaragua, Mexico.

biformis Cameron, 1951b: 127 ( Belonuchus ; Type locality: Nigeria: Jos , Plateau Bauchi).

- DISTRIBUTION: Nigeria.

rufopygus Cameron, 1951b: 128 ( Belonuchus ; variety of biformis ; Type locality: not stated but possibly Nigeria: Jos, Plateau Bauchi).

bipunctatus Bernhauer, 1935c: 90 ( Belonuchus ; Type locality: Kolumbien [ohne nähere Fundortangabe ]; Cali, Villa Elvira).

- DISTRIBUTION: Colombia.

birmanus Cameron, 1932: 174 ( Belonuchus ; Type locality: Burma: Ruby Mines ).

— Scheerpeltz, 1933: 1377 ( Belonuchus ; catalog).

- DISTRIBUTION: Myanmar.

boops Sharp, 1885: 435 ( Belonuchus ; Type locality: Panama, Bugaba ).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: Panama.

borneensis Cameron, 1933b: 346 ( Belonuchus ; Type locality: British North Borneo: Kenokok, 3300 feet).

- DISTRIBUTION: Borneo.

boxi Bernhauer, 1934b: 212 ( Belonuchus ; Type locality: Argentinien: Tucuman).

- DISTRIBUTION: Argentina.

brasilianus Bernhauer, 1917c: 19 ( Belonuchus ; Type locality: Brasilien: S. Catharina).

— Scheerpeltz, 1933: 1377 ( Belonuchus ; catalog).

- DISTRIBUTION: Brazil.

breviceps Fauvel, 1891: 117 ( Belonuchus ; Type locality: Colonie Tovar).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: Venezuela.

brevicollis Fauvel, 1877: 267 ( Belonuchus ; Type locality: Australie orientale, Queensland, Wide Bay ).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

— Lea, 1925: 228 ( Belonuchus ; characters; Australia).

- DISTRIBUTION: Australia.

bruchi Bernhauer, 1912e: 175 ( Belonuchus ; Type locality: Argentinien: Prov. Buenos Aires). — Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: Argentina.

bruchi Bernhauer, 1934 , see: duplicatus Blackwelder, 1944 .

bruchianus Bernhauer, 1911a: 413 ( Belonuchus ; Type locality: Argentinien, Gob. Misiones). — Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

- DISTRIBUTION: Argentina.

bugnioni Fauvel, 1901: 86 ( Belonuchus ; Type locality: Martinique).

— Bernhauer and Schubert, 1914: 369 ( Belonuchus ; catalog).

— Blackwelder, 1943: 428 ( Belonuchus ; characters; Guadeloupe; Martinique).

- DISTRIBUTION: West Indies.