Philonthus nigriceps, Mannerheim, 1830: 34

Taphylininae, Iochini, Aorothiini, Thiini, Latyprosopini, Taphylinini, Mblyopinina, Nisolinina, Yptiomina, Hilonthina & Herman, Lee H., 2001, Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina), Bulletin of the American Museum of Natural History 2441 (265), pp. 2435-3021 : 2456-2474

publication ID

https://doi.org/ 10.1206/0003-0090.265.1.5

persistent identifier

https://treatment.plazi.org/id/1D1E87B6-FFC7-FFA3-FF60-4EDFFD9B486A

treatment provided by

Marcus

scientific name

Philonthus nigriceps
status

 

nigriceps Mannerheim, 1830: 34 View in CoL ( Gyrohypnus ; Type locality: Suecia: Kinnekulle ).

— Mannerheim, 1831: 448 ( Gyrohypnus ; Sweden).

— Nordmann, 1837: 120 ( Gyrohypnus ; Sweden).

— Erichson, 1839b: 297 ( Othius ; synonym of pilicornis ).

— Kraatz, 1857d: 660 ( Baptolinus ; synonym of alternans ).

— Mulsant and Rey, 1877a: 244 ( Baptolinus ; synonym of pilicornis ).

— Ganglbauer, 1895: 474 ( Baptolinus ; synonym of affinis ).

— Reitter, 1908: 108 ( Baptolinus ; synonym of affinis ).

— Bernhauer and Schubert, 1914: 316 ( Baptolinus ; synonym of affinis ).

— Smetana, 1958: 121 ( Atrecus ; synonym of affinis ).

— Coiffait, 1972: 363 ( Baptolinus ; synonym of affinis ).

— Bordoni, 1982: 404 ( Baptolinus ; synonym of affinis ).

glabricornis Stephens, 1833: 254 ( Othius ; Type locality: near London; Crwmlyn sandhills).

— Stephens, 1839: 402 ( Othius ; characters; England).

— Hardy, 1851: 37 ( Othius ; synonym of pilicornis ).

— G. Waterhouse, 1858: 26 ( Othius ; synonym of alternans ).

— Ganglbauer, 1895: 474 ( Baptolinus ; synonym of affinis ).

— Reitter, 1908: 108 ( Baptolinus ; synonym of affinis ).

— Bernhauer and Schubert, 1914: 316 ( Baptolinus ; synonym of affinis ).

— Smetana, 1958: 121 ( Atrecus ; synonym of affinis ).

— Coiffait, 1972: 363 ( Baptolinus ; synonym of affinis ).

— Bordoni, 1982: 404 ( Baptolinus ; synonym of affinis ).

frigidus Dufour, 1843: 35 ( Xantholinus ; Type locality: vallée d’Ossau).

— Fauvel, 1869: 490 ( Baptolinus ; synonym of pilicornis ).

— Fauvel, 1873b: 87 [= 1873a: 373] ( Baptolinus ; synonym of affinis ).

— Ganglbauer, 1895: 474 ( Baptolinus ; synonym of affinis ).

— Reitter, 1908: 108 ( Baptolinus ; synonym of affinis ).

— Bernhauer and Schubert, 1914: 316 ( Baptolinus ; synonym of affinis ).

— Smetana, 1958: 121 ( Atrecus ; synonym of affinis ).

— Coiffait, 1972: 363 ( Baptolinus ; synonym of affinis ).

— Bordoni, 1982: 404 ( Baptolinus ; synonym of affinis ).

dimidiatus Motschulsky, 1860: 565 ( Othius ; Type locality: Sibérie orientale).

— Hochhuth, 1862: 69 ( Othius ; characters; Siberia).

— Fauvel, 1873b: 87 [= 1873a: 373] ( Baptolinus ; synonym of affinis ).

— Bernhauer and Schubert, 1914: 316 ( Baptolinus ; synonym of affinis ).

— Smetana, 1958: 121 ( Atrecus ; synonym of affinis ).

— Coiffait, 1972: 363 ( Baptolinus ; synonym of affinis ).

— Bordoni, 1982: 404 ( Baptolinus ; synonym of affinis ).

caucasicus Roubal, 1933: 77 ( Baptolinus ; subspecies of affinis ; Type locality: Ca. occ.: Krasnaja Poljiana; Teberda).

— Coiffait, 1972: 362, 363 ( Baptolinus ; subspecies of affinis ; characters; Caucasus).

— Coiffait, 1978d: 163 ( Baptolinus ; subspecies of affinis ; Turkey).

— Assing, 1998c: 345 ( Atrecus ; discussion; synonym of affinis ).

alternans Gravenhorst, 1802 , see: affinis Paykull, 1789 .

americanus Casey, 1906: 426 ( Baptolinus ; Type locality: Lake Superior; [Note: See lectotype designation by Smetana, 1982]).

— Bernhauer and Schubert, 1914: 316 ( Baptolinus ; catalog).

— Notman, 1920: 17 ( Baptolinus ; New York).

Smetana, 1982: 44 ( Atrecus ; lectotype designation; characters; Nova Scotia; Ontario; Quebec; Maine; Massachusetts; Michigan; New Hampshire; New York; North Carolina; Pennsylvania; Vermont; West Virginia) .

— Smetana, 1990c: 83 ( Atrecus ; Arkansas; Tennessee).

— Downie and Arnett, 1996: 395 ( Atrecus ; characters; USA).

- DISTRIBUTION: Canada, USA.

ardeanus Ciceroni, 1990: 55 ( Atrecus ; Type locality: Italia: Massiccio del Pollino , Piana del Pollino, 1700 m).

— Ciceroni and Zanetti, 1995: 26 ( Atrecus ; Italy).

— Assing, 1998c: 350 ( Atrecus ; characters).

- DISTRIBUTION: Italy.

brevicornis Smetana, 1967d: 199 ( Atrecus ; Type locality: Mongolei: Central aimak, 12 km SO v. Ulan-Baator, Nucht in Bogdo ul, 1500 m).

— Coiffait, 1972: 364 ( Baptolinus ; characters).

— Assing, 1998c: 350 ( Atrecus ; characters).

- DISTRIBUTION: Mongolia.

casalei Bordoni, 1987: 568 ( Baptolinus ; Type locality: Grecia orientale: M.te Pangaeon, Akrovunion, Kavala, 1300 m).

— Assing, 1998c: 349 ( Atrecus ; characters).

— Assing and Wunderle, 1999a: 481 ( Atrecus ; characters; notes; Greece).

- DISTRIBUTION: Greece.

caucasicus Roubal, 1933 , see: affinis Paykull, 1789 .

dimidiatus Motschulsky, 1860 , see: affinis Paykull, 1789 .

fraternus Casey, 1906 , see: punctiventris Fall, 1901 .

frigidus Dufour, 1843 , see: affinis Paykull, 1789 .

fulvicollis Fairmaire and Germain, 1861 , see: Cheilocolpus .

glabricornis Stephens, 1833 , see: affinis Paykull, 1789 .

longiceps Fauvel, 1872: 374 [= 1873b: 88] ( Baptolinus ; Type locality: Nancy; Savoie, Rhonnes; Grande Chartreuse).

— Mulsant and Rey, 1877a: 247 ( Baptolinus ; characters; France).

— Ganglbauer, 1895: 474 ( Baptolinus ; characters; France; Switzerland; Germany; Austria; Tuscany; North America; [Note: According to Smetana, 1982: 37, 47, longiceps does not occur

in North America, most specimens so identified are americanus ]).

— Reitter, 1908: 107 ( Baptolinus ; characters; central Europe).

— Reitter, 1909: 133 ( Baptolinus ; characters; Germany).

— Bernhauer and Schubert, 1914: 316 ( Baptolinus ; catalog).

— Portevin, 1929: 368 ( Baptolinus ; characters; France).

— Scheerpeltz, 1933: 1321 ( Baptolinus ; catalog).

— V. Hansen, 1952: 22 ( Baptolinus ; characters; Denmark).

— Coiffait, 1956a: 43 ( Baptolinus ; characters in key).

— Smetana, 1958: 120 ( Atrecus ; characters; ecology; Czechoslovakia).

— Palm, 1963b: 54 ( Baptolinus ; characters; Sweden; Denmark; Norway; Finland).

— Szujecki, 1963a: 33 ( Atrecus ; Poland).

— Lohse, 1964: 167 ( Baptolinus ; characters; central Europe).

— Horion, 1965: 104 ( Baptolinus ; Finland; Sweden; Norway; Denmark; France; Switzerland; Italy; Slovenia; Dalmatia; Bosnia; Slovakia; Czech Republic; Poland; Austria; Germany).

— Smetana, 1966: 330 ( Atrecus ; notes; Czechoslovakia).

— Coiffait, 1972: 364 ( Baptolinus ; characters; references; Europe; Scandinavia; Italy; Siberia; Britain).

— Legner and Moore, 1977: 176 ( Atrecus ; found under bark).

— Bordoni, 1982: 405 ( Baptolinus ; characters; Italy).

— Lucht, 1987: 103 ( Baptolinus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Denmark; Sweden).

— Bordoni, 1986b: 387 ( Baptolinus ; Slovakia; Slovenia; Carniola).

— Tóth, 1989: 40 ( Baptolinus ; characters; Hungary).

— Janák, 1993a: 47 ( Atrecus ; Czech Republic; Slovakia).

— Ciceroni, 1993: 77 ( Atrecus ; Italy).

— Ciceroni and Zanetti, 1995: 26 ( Atrecus ; Italy).

— Assing, 1998c: 350 ( Atrecus ; characters).

- DISTRIBUTION: Europe.

macrocephalus Nordmann, 1837: 124 ( Gyrohypnus ; Type locality: insula Sitka).

— Erichson, 1839b: 297 ( Othius ; characters; America borealis occidentalis, ad littora ins. Sitchae). — Mannerheim, 1843b: 227 ( Othius ; characters; Sitka Island).

— Mannerheim, 1852: 312 ( Othius ; characters; notes; Sitka).

— Provancher, 1877: 252 ( Baptolinus ; cited as melanocephalus ; characters; Quebec).

— Casey, 1906: 426 ( Baptolinus ; characters; New Hampshire; British Columbia; Queen Charlotte Island; Prince of Wales Island).

— Bernhauer and Schubert, 1914: 316 ( Baptolinus ; catalog).

— Van Dyke, 1924: 17 ( Baptolinus ; Alaska).

— Fall, 1926: 149 ( Baptolinus ; Alaska).

— Hatch, 1957: 232 ( Atrecus ; characters; British Columbia; Washington; Oregon; Idaho).

— Smetana, 1982: 38 ( Atrecus ; characters; Alberta; British Columbia; New Brunswick; Newfoundland; Northwest Territories; Nova Scotia; Ontario; Quebec; Alaska; California; Idaho; Maine; Montana; New Hampshire; New York; Oregon; Vermont; Washington).

— Dearborn and Donahue, 1994: 14 ( Atrecus ; Maine).

— Downie and Arnett, 1996: 396 ( Atrecus ; characters; USA).

- DISTRIBUTION: USA, Canada.

newtoni Smetana, 1982: 53 ( Atrecus ; Type locality: Washington: Pierce Co., Mt. Rainier N. P., 4.7 mi. W Longmire, 2200 feet. Also cited from Oregon).

— Smetana, 1990c: 83 ( Atrecus ; Oregon).

- DISTRIBUTION: USA.

nigriceps Mannerheim, 1830 , see: affinis Paykull, 1789 .

parvioculatus Assing, 1998c: 346 ( Atrecus ; Type locality: Turquie: S. Vit, Ayder, 1400 m, Vallée de la Firtina, Rize).

- DISTRIBUTION: Turkey.

pilicornis Paykull, 1790: 135 ( Staphylinus ; Type locality: Not cited but presumably Suecia). — Schrank, 1798: 642 ( Staphylinus ; characters; notes).

— Paykull, 1800: 379 ( Staphylinus ; characters; Sweden).

— Gyllenhal, 1810: 359 ( Staphylinus ; characters; Sweden).

— Zetterstedt, 1828: 81 ( Xantholinus ; characters; Lapland).

— Curtis, 1829: 26 ( Xantholinus ; catalog; Britain).

— C. Sahlberg, 1830: 335 ( Staphylinus ; characters; Finland).

— Mannerheim, 1830: 34 ( Gyrohypnus ; Finland; Sweden).

— Mannerheim, 1831: 448 ( Gyrohypnus ; Finland; Sweden).

— Stephens, 1833: 254 ( Othius ; characters; England).

— Nordmann, 1837: 119 ( Gyrohypnus ; Europe).

— Zetterstedt, 1838: 66 ( Xantholinus ; characters; habitat; Lapland).

— Erichson, 1839a: 422 ( Othius ; characters; Germany).

— Heer, 1839: 248 ( Othius ; characters; Switzerland).

— Erichson, 1839b: 296 ( Othius ; characters; Sweden; Germany).

— Stephens, 1839: 401 ( Othius ; characters; England).

— Märkel, 1845: 235 ( Othius ; with Formica fuliginosa ).

— Jacobsen, 1849: 675 ( Othius ; characters; notes; Denmark).

— Hochhuth, 1849: 100 ( Othius ; characters; Lenkoran).

— Redtenbacher, 1849: 690 ( Othius ; characters; Austria).

— Hardy, 1851: 37 ( Othius ; notes; England).

— Fairmaire and Laboulbène, 1856: 499 ( Othius ; characters; France).

— Kraatz, 1857d: 661 ( Baptolinus ; characters; Germany).

— Redtenbacher, 1857: 179 ( Othius ; characters; Austria).

— Redtenbacher, 1857: 982 ( Baptolinus ; characters; Austria).

— Thomson, 1859: 27 (type species of Gyrohypnus ).

— Thomson, 1860: 187 ( Gyrohypnus ; characters; Scandinavia).

— Fauvel, 1865a: 298 [= 1865b: 48] ( Baptolinus ; habitat; France).

— Fauvel, 1873b: 86 [= 1873a: 372] ( Baptolinus ; characters; France; Lapland; Scandinavia; Russia; Germany; Switzerland; Austria; Italy). — Redtenbacher, 1874: 222 ( Baptolinus ; characters; Austria).

— Seidlitz, 1875: 279 ( Baptolinus ; characters; Baltic region).

— J. Sahlberg, 1876: 33 ( Baptolinus ; Finland).

— Mulsant and Rey, 1877a: 241 ( Baptolinus ; characters; France).

— Sharp, 1889: 249 ( Baptolinus ; Japan).

— Ganglbauer, 1895: 473 ( Baptolinus ; characters; north and middle Europe).

— Reitter, 1908: 107 ( Baptolinus ; characters; northern and central Europe).

— Reitter, 1909: 133 ( Baptolinus ; characters; Germany).

— Bernhauer and Schubert, 1914: 316 ( Baptolinus ; catalog).

— Johansen, 1914: 429 ( Baptolinus ; characters; Denmark).

— Šulc, 1922 ( Baptolinus ; osmeterium; anatomy; histology; physiology).

— Bernhauer, 1926g: 2 ( Baptolinus ; Kamchatka region).

— Portevin, 1929: 368 ( Baptolinus ; characters; France).

— Scheerpeltz, 1933: 1321 ( Baptolinus ; catalog).

— V. Hansen, 1952: 23 ( Baptolinus ; characters).

— Coiffait, 1956a: 42 ( Baptolinus ; characters in key).

— Smetana, 1958: 119 ( Atrecus ; characters; ecology; Czechoslovakia).

— Palm, 1963b: 54 ( Baptolinus ; characters; Sweden; Norway; Finland).

— Nakane, 1963a: 90 ( Atrecus ; characters; habitus photograph; Japan).

— Lohse, 1964: 167 ( Baptolinus ; characters; central Europe).

— Horion, 1965: 106 ( Baptolinus ; Russia; Finland; Sweden; Norway; Ukraine; Poland; Czechoslovakia; Hungary; Romania; Italy; Switzerland; France; Dalmatia; Bosnia Hercegovina; Croatia; Slovenia; Spain?; Austria; Germany).

— Pototskaia, 1967: 49 ( Atrecus ; larval characters).

— Coiffait, 1972: 362 ( Baptolinus ; characters; references; Europe; Siberia; Caucasus).

— Smetana 1975c: 168 ( Baptolinus ; Mongolia).

— Pototskaia, 1976: 172 ( Atrecus ; larval characters).

— Legner and Moore, 1977: 176 ( Atrecus ; found under bark).

— Bordoni, 1982: 402 ( Baptolinus ; characters; Italy).

— Bordoni, 1986b: 386 ( Baptolinus ; Slovakia).

— Lucht, 1987: 103 ( Baptolinus ; checklist; Germany; Poland; Czechoslovakia; Austria; France; Sweden).

— Tóth, 1989: 41 ( Baptolinus ; characters).

— Lohse and Lucht, 1989: 159 ( Atrecus ; characters).

— J. Li, 1993: 33 ( Atrecus ; China).

— Janák, 1993a: 47 ( Atrecus ; Czech Republic; Slovakia).

— Muona and Viramo, 1995: 9 ( Atrecus ; collected from dung of Ursus arctos ; Finland).

— Ciceroni and Zanetti, 1995: 26 ( Atrecus ; Italy).

— Raitschev, 1997a: 100 ( Baptolinus ; notes; Bulgaria).

— Assing, 1998c: 350 ( Atrecus ; characters).

— Ryabukhin, 1999: 56 ( Atrecus ; catalog; habitat; distribution; northeastern Russia).

— Naomi, Kuranishi, Saito, and Maruyama, 2000: 109 ( Atrecus ; Kamchatka).

— Olberg and Andersen, 2000: 223 ( Atrecus ; attracted to trunk-window traps baited with Fomes fomentarius and Phellinus spp. ; Norway).

- DISTRIBUTION: Europe, Russia, Azerbaijan, Mongolia, China, Japan.

suturalis Motschulsky, 1845b: 359 ( Othius ; Type locality: Kamtschatka; [Note: See lectotype designation by Assing, 1998c]).

— Bernhauer and Schubert, 1914: 316 ( Baptolinus ; catalog).

— Assing, 1998c: 344 ( Atrecus ; lectotype designation; characters; Russia; synonym of pilicornis ).

— Ryabukhin, 1999: 56 ( Atrecus ; cited as valid species; catalog; habitat; distribution; northeastern Russia).

punctiventris Fall, 1901: 221 ( Baptolinus ; Type locality: Lake Tahoe, California; [Note: See lectotype designation by Smetana, 1982]. Also cited from Utah and Vancouver Island [Note: According to Smetana, 1982: 48, the Canadian specimen is probably a misidentification of quadripennis ).

— Casey, 1906: 427 ( Baptolinus ; characters; California). — Bernhauer and Schubert, 1914: 316 ( Baptolinus ; catalog).

— Hatch, 1957: 233 ( Atrecus ; characters; British Columbia; Washington; Oregon; [Note: According to Smetana, 1982: 51, Hatch’s Canadian record may be a misidentification of quadripennis ]).

Smetana, 1982: 47 ( Atrecus ; lectotype designation: Lake Tahoe, Cal ; characters; California; Nevada; Oregon; Washington) .

- DISTRIBUTION: USA.

fraternus Casey, 1906: 428 ( Baptolinus ; Type locality: California, Siskyou Co.; [Note: See lectotype designation by Smetana, 1982]).

— Bernhauer and Schubert, 1914: 316 ( Baptolinus ; catalog).

— Smetana, 1982: 47 ( Atrecus ; lectotype designation; synonym of punctiventris ).

quadripennis Casey, 1906: 427 ( Baptolinus ; Type locality: Idaho, Coeur d’Alene; [Note: See lectotype designation by Smetana, 1982]).

— Bernhauer and Schubert, 1914: 316 ( Baptolinus ; catalog).

— Hatch, 1957: 233 ( Atrecus ; characters; British Columbia; Washington; Oregon; Idaho).

Smetana, 1982: 51 ( Atrecus ; lectotype designation; characters; Alberta; British Columbia; Idaho; Oregon; Utah; Washington; Wyoming) .

— Smetana, 1988a: 526 ( Atrecus ; collecting notes; Oregon).

- DISTRIBUTION: Canada, USA.

schuelkei Assing, 1998c: 348 ( Atrecus ; Type locality: China: W-Sichuan, Daxue Shan , Hailuogou Glacier Park Camp 1, 2100 m, 29.36.00N, 102.03.35E).

- DISTRIBUTION: China.

suturalis Motschulsky, 1845 , see: pilicornis Paykull, 1790 .

yunnanus Assing, 2000c: 75 ( Atrecus ; Type locality: China: Yunnan Prov., Heishui, 35 km N. Lijiang).

- DISTRIBUTION: China.

Othius

[105 species (4 doubtful); Palaearctic, Oriental, and Australian Regions]

Othius Stephens, 1829a: 23 (species included: fulgidus , pilicornis , ustulatus ). TYPE SPECIES: Staphylinus punctulatus Goeze , fixed by action of the Commission in Opinion 1250 (ICZN, 1983).

— Stephens, 1829: 284 (catalog; Britain).

— Stephens, 1832: 200 (characters).

— Stephens, 1833: 253 (characters; habitat).

— Westwood, 1838a: 16 (characters; type species).

— Erichson, 1839a: 420 (characters).

— Heer, 1839: 247 (characters).

— Erichson, 1839b: 294 (characters).

— Stephens, 1839: 385 (characters).

— Redtenbacher, 1849: 690 (characters).

— Schaum, 1852: 25 (list of species; Europe).

— Lacordaire, 1854: 63 (characters; notes; list of species).

— Wollaston, 1854: 574 (characters; notes).

— Fairmaire and Laboulbène, 1856: 498 (characters).

— Jacquelin du Val, 1856: 30 (characters).

— Gistel, 1856: 387 (list of species).

— Kraatz, 1857d: 653 (characters).

— Redtenbacher, 1857: 179 (characters; key to Austrian species).

— Thomson, 1858: 30 (characters).

— G. Waterhouse, 1858: 26 (catalog of British species).

— Thomson, 1859: 26 (characters; type species: fulvipennis ).

— Schaum, 1859: 27 (catalog; European species).

— Thomson, 1860: 185 (characters).

— LeConte, 1861a: 65 (characters).

— Gredler, 1863: 108 (list of species from Tirol).

— LeConte, 1863: 24 (list of North American species).

— Fauvel, 1866a: 347 [= 1866b: 98] (characters).

— Wencker and Silbermann, 1866: 31 (list of species; collecting notes; France).

— Gemminger and Harold, 1868: 607 (catalog).

— Jacquelin du Val, 1868: 65 (catalog; European species).

— Sharp, 1871d: 12 (list of British species).

— Fauvel, 1873b: 81 [= 1873a: 367] ( Cafius ; characters; France).

— Redtenbacher, 1874: 222 (characters; key to Austrian species).

— Fauvel, 1875a: XXIV [= 1875b: 226] (catalog).

— Mulsant and Rey, 1877a: 219 (characters; key to French species).

— Broun, 1880: 101 (characters).

— Heyden, 1880: 76 (list of species of Siberian region).

— Duvivier, 1883: 159 (catalog).

— Fowler, 1888: 294 (notes; key to British species).

— Seidlitz, 1891: 379 (key to some European species).

— Seidlitz, 1891a: 404 (key to some European species).

— Heyden, Reitter, and Weise, 1891: 105 (list of species of Europe and Caucasus).

— Heyden, 1893: 43 (list of species of Siberian region).

— Ganglbauer, 1895: 468 (larval and adult characters; key to middle European species).

— Fauvel, 1897d: 289 (catalog of species of Barbary; Madeira; Canaries).

— Casey, 1906: 422 (characters).

— Heyden, Reitter, and Weise, 1906: 162 (list of species of Europe and Caucasus).

— Reitter, 1908: 103, 104 (characters; key to species).

— Reitter, 1909: 132 (characters; key to species).

— Petri, 1912: 70 (list of species of Siebenbürgen).

— Bernhauer and Schubert, 1914: 317 (world catalog; 28 species).

— Johansen, 1914: 426 (characters; key to Danish species).

— Winkler, 1925: 373 (catalog for Palaearctic region).

— Porta, 1926: 91 (characters; key to species of Italy).

— Portevin, 1929: 367 (key to French species).

— Roubal, 1930: 365 (catalog; Slovakia).

— Cameron, 1932: 41 (characters; key to species of British India).

— Joy, 1932: 127 (key to British species).

— Scheerpeltz, 1933: 1321 (world catalog supplement).

— Normand, 1935: 369 (list of species of Tunisia).

— Blackwelder, 1944: 145 (checklist of species from Mexico, Central and South America, and

the West Indies).

— Tottenham, 1949: 371 (synonym of Gyrohypnus ; type species: punctulatus Goeze ).

— Blackwelder, 1952: 279 (type species: fulgidus ; discussion).

— V. Hansen, 1952: 23 (characters; key to Danish species).

— Coiffait, 1956a: 37, 38 (characters; key to species; cited as synonym of Gyrohypnus but believes rules of priority should be suspended for this case).

— Kocher, 1958: 141 (checklist of species; Morocco).

— Smetana, 1958: 122 (characters; habitus of one species; type species: laeviusculus ; key to Czechoslovakian species).

— Smetana, 1959a: 209 (list of species collected in Albania).

— Ferreira, 1962a: 16 (list of Portuguese species).

— Székessy, 1963: 51 (characters).

— Palm, 1963b: 54 (characters; key to species of Sweden).

— Lohse, 1964: 167 (characters; key to central European species).

— Pototskaia, 1967: 46 (larval characters).

— Coiffait and Saiz, 1968: 345 (list of Chilean species).

— Scheerpeltz, 1968a: 63 (catalog of Austrian species).

— Kasule, 1970: 74, 76 (larval characters).

— Coiffait, 1972: 327 (characters; larval characters; type species: punctulatus ; key to species of western Palaearctic region; references).

— Bordoni, 1973j: 691 (list of species from islands near Sicily).

— Tikhomirova, 1973a: 180 (checklist of species of USSR).

— Palm, 1976: 211 (key to species of Canary Islands).

— Pope, 1977: 28 (list of British species).

— Topp, 1978: 311 (larval characters).

— Palm, 1979: 269 (key to species of Madeira; notes).

— Muona, 1979: 16 (list of Scandinavian species).

— Uhlig, 1979: 252 (some species collected in Germany).

— M. Dvoř, 1979: 115 (some species collected in Slovakia).

— Smetana, 1979: 50 (requested that International Commission on Zoological Nomenclature set aside all type species designations and designate Staphylinus punctulatus Goeze as type species and place Othius on the Official list of Generic Names in Zoology).

— Uhlig, Vogel, and Sieber, 1980: 246 (some species collected in Germany).

— Burakowski, Mroczkowski, and Stefańska, 1980: 25 (catalog; Poland).

— Uhlig and Vogler, 1981: 109 (list of some species collected in Germany; habitat and collecting notes).

— Bordoni, 1982: 378 (characters; habitus; key to species of Italy).

— ICZN, 1983: 85 ( Othius placed on Official List of Generic Names in Zoology and Staphylinus punctulatus is fixed as the type species).

— Shibata, 1983: 81 (checklist of species of Japan).

— Shibata, 1986a: 123 (supplemental list of species from Taiwan).

— Muona and Viramo, 1986: 14 (list of species of northeastern Finland).

— Segers, 1986: 41 (checklist of species; Belgium).

— Tóth, 1989: 34 (characters; key to species of Hungary).

— Lohse and Lucht, 1989: 159 (supplement to key to species of Lohse, 1964).

— Angelini, 1991: 192 (list of species from high elevations of Calabria, Italy).

— Silfverberg, 1992: 18 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Siitonen, 1993: 230 (list of Finnish species).

— Welch, 1993: 228 (ovariole structure and development).

— Janák, 1993a: 47 (list of species of Czech Republic and Slovakia).

— Hernández, Outerelo, and Gamarra, 1994: 190 (checklist of species; Canary Islands).

— Assing and Wunderle, 1995a (revision of and key to species of Madeira; notes).

— Ciceroni and Zanetti, 1995: 26 (list of species of Italy).

— M. Hansen, 1996: 108 (list of Danish species).

— Assing, 1997a (review of and key to species of Canary Islands).

— Assing, 1997 (revision of and key to species of western Palaearctic; characters for species groups).

— Assing, 1998 (revision of and key to species of Himalayan region).

— Assing, 1998a (new records for, colonization by, and phylogeny of species of Madeira Canary Islands).

— Assing, 1999 (revision of and key to eastern Palaearctic species east of Himalaya).

— Assing and Schülke, 1999: 8 (key to species of central Europe).

Othiellus Casey, 1906: 422 , 423 (species included: laeviusculus , melanocephalus ). TYPE SPECIES: Othius laeviusculus Stephens , fixed by subsequent designation by Tottenham, 1939d: 339.

— Bernhauer and Schubert, 1914: 317 (synonym of Othius ).

— Cameron, 1932: 41 (synonym of Othius ).

— Tottenham, 1939d: 229 (type species designation: laeviusculus ).

— Tottenham, 1949: 371 (synonym of Gyrohypnus ; type species: laeviusculus Stephens ).

— Blackwelder, 1952: 279 (type species).

— Smetana, 1958: 122 (synonym of Othius ).

— Coiffait, 1972: 327 (synonym of Othius ).

— Tikhomirova, 1973a: 180 (subgenus of Othius ; checklist of species of USSR).

— Erichson, 1839a: 421 ( Othius ; characters; Germany).

— Heer, 1839: 248 ( Othius ; characters; Switzerland).

— Erichson, 1839b: 295 ( Othius ; characters; Europe).

— Stephens, 1839: 402 ( Othius ; characters; England).

— Grimm, 1845: 134 ( Othius ; with Formica fuliginosa ; Germany).

— Jacobsen, 1849: 674 ( Othius ; characters; notes; Denmark).

— Redtenbacher, 1849: 690 ( Othius ; characters; Austria).

— Hardy, 1851: 36 ( Othius ; notes; England).

— Fairmaire and Laboulbène, 1856: 498 ( Othius ; characters; France).

— Kraatz, 1857d: 656 ( Othius ; characters; Germany).

— Redtenbacher, 1857: 179 ( Othius ; characters; Austria).

— Thomson, 1860: 185 ( Othius ; characters; Scandinavia).

— Hochhuth, 1862: 69 ( Othius ; Russia).

— Fauvel, 1865a: 298 [= 1865b: 48] ( Othius ; habitat; France).

— Fauvel, 1873b: 85 [= 1873a: 371] ( Othius ; characters; France; Europe).

— Redtenbacher, 1874: 223 ( Othius ; characters; Austria).

— Seidlitz, 1875: 279 ( Othius ; characters; Baltic region).

— J. Sahlberg, 1876: 31 ( Othius ; Finland).

— Mulsant and Rey, 1877a: 231 ( Othius ; characters; France).

— Fowler, 1888: 295 ( Othius ; characters; Britain).

— Ganglbauer, 1895: 470 ( Othius ; characters; north and middle Europe).

— Poppius, 1905: 10 ( Othius ; Orkney Islands; Shetland Islands).

— Casey, 1906: 423 ( Othiellus ).

— Reitter, 1908: 106 ( Othius ; characters; northern and central Europe).

— Reitter, 1909: 133 ( Othius ; characters; Germany).

— Sharp and Muir, 1912: 499 ( Othius ; characters of male genitalia).

— Bernhauer and Schubert, 1914: 318 ( Othius ; catalog).

— Johansen, 1914: 427 ( Othius ; characters; Denmark).

— Šulc, 1922 ( Othius ; osmeterium; anatomy; histology; physiology).

— Portevin, 1929: 368 ( Othius ; characters; France).

— Scheerpeltz, 1933: 1322 ( Othius ; catalog).

— West, 1937: 41 ( Othius ; Faroe Islands).

— V. Hansen, 1952: 25 ( Othius ; characters; Denmark).

— Coiffait, 1956a: 40 ( Othius ; characters in key).

— Smetana, 1958: 126 ( Othius ; synonym of angustus ).

— Larsson and Gígja, 1959: 81 ( Othius ; biological notes; Iceland).

— Palm, 1963b: 55 ( Othius ; characters; Sweden; Denmark; Norway; Finland).

— Lohse, 1964: 168 ( Othius ; characters; central Europe).

— Horion, 1965: 110 ( Othius ; widespread in northern Europe, in central Europe south to Spain and Italy).

— Pototskaia, 1967: 48 ( Othius ; larval characters).

— Kasule, 1970: 76 ( Othius ; larval characters).

— Coiffait, 1972: 348 ( Othius ; synonym of angustus ).

— Pope, 1977: 28 ( Othius ; synonym of angustus ).

— Lucht, 1987: 103 ( Othius ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Lohse and Lucht, 1989: 160 ( Othius ; cited as valid name; characters).

— Assing, 1993: 33 ( Othius ; bionomics; Germany).

— Ciceroni and Zanetti, 1995: 26 ( Othius ; synonym of angustus ).

— Gerend and Braunert, 1997: 197 ( Othius ; cited as valid species; notes; Luxembourg).

— Assing, 1997: 24 ( Othius ; lectotype designation; synonym of angustus ).

— Uhlig, 1998: 136 ( Othius ; synonym of angustus ).

sexpunctatus Haliday, 1841: 187 ( Othius ; Type locality: Holywood).

— Kraatz, 1857d: 656 ( Othius ; synonym of melanocephalus ).

— Ganglbauer, 1895: 470 ( Othius ; synonym of melanocephalus ).

— Reitter, 1908: 106 ( Othius ; synonym of melanocephalus ).

— Bernhauer and Schubert, 1914: 318 ( Othius ; synonym of melanocephalus ).

— Smetana, 1958: 127 ( Othius ; synonym of angustus ).

— Coiffait, 1972: 348 ( Othius ; synonym of angustus ).

stenocephalus Eppelsheim, 1881: 506 ( Othius ; Type locality: Helenendorf; [Note: See lectotype designation by Assing, 1997]).

— Reitter, 1908: 106 ( Othius ; characters; Caucasus).

— Bernhauer and Schubert, 1914: 317 ( Othius ; variety of laeviusculus ).

— Scheerpeltz, 1933: 1322 ( Othius ; variety of laeviusculus ).

— Coiffait, 1972: 348 ( Othius ; characters; references; Balkans; Caucasus).

— Coiffait, 1978d: 163 ( Othius ; Turkey).

— Assing, 1997: 30 ( Othius ; subspecies of angustus ; lectotype designation; characters; notes; Turkey; Caucasus, Russia, Georgia).

— Assing, 1999c: 667 ( Othius ; subspecies of angustus ; Turkey; Georgia; Caucasus).

— Solodovnikov, 2000a: 325 ( Othius ; subspecies of angustus ; notes; Turkey).

sipeki Roubal, 1919: 15 ( Othius ; aberration of melanocephalus ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from [ Bohemia]: Pribram).

— Scheerpeltz, 1933: 1322 ( Othius ; aberration of melanocephalus ).

— Smetana, 1958: 127 ( Othius ; aberration of angustus ).

angustus Broun, 1893 , see: Maorothius .

antarcticus Fairmaire, 1885 , see: Antarctothius .

apicalis Cameron, 1926b: 347 ( Othius ; Type locality: Chakrata District, Chulli Khud, 6500 ft.; [Note: See lectotype designation by Assing, 1998]. Also cited from Mussorie District; Arni Gad).

— Cameron, 1932: 43 ( Othius ; characters; India).

— Scheerpeltz, 1933: 1321 ( Othius ; catalog).

— Paulian, 1941: 219 ( Othius ; larval characters; India).

Assing, 1998: 295 ( Othius ; characters; lectotype designation: Chakrata District, Chulli Khud , 6500 ft.; Nepal; India; Sikkim) .

— Assing, 1999c: 684 ( Othius ; correct measurements for TiL and TaL cited in Assing, 1998,

by factor of 2.02).

- DISTRIBUTION: India, Nepal.

arieiroensis Palm, 1979: 272 ( Othius ; Type locality: Madeira: between Pico Arieiro and Poiso, alt. about 1600 m).

— Assing and Wunderle, 1995a: 60 ( Othius ; characters; bionomics; Madeira).

— Assing, 1998a: 215 ( Othius ; Madeira).

- DISTRIBUTION: Madeira.

arisanus Shibata, 1973 , see: punctatus Bernhauer, 1922 .

atavisticus Rüschkamp, 1932, see: subuliformis Stephens, 1833 .

atavus Assing, 1999: 32 ( Othius ; Type locality: China: Yunnan prov., Gaoligongshan Mts. ,

90 km W of Baoshan).

- DISTRIBUTION: China.

ater Stephens, 1833 , see: Gyrohypnus .

baculifer Assing and Wunderle, 1995a: 61 ( Othius ; Type locality: Madeira: Bica da Cana , 1550 m).

— Assing, 1998a: 215 ( Othius ; Madeira).

- DISTRIBUTION: Madeira.

badius Zheng, 1995: 341 , 346 ( Othius ; Type locality: Sichuan, Mt. Emei, alt. 1752–2070 m). — Assing, 1999: 43 ( Othius ; characters; China).

- DISTRIBUTION: China.

bhutanensis Assing, 1999c: 681 ( Othius ; Type locality: Bhutan: Paro District, Chiley-La, 3000–3500 m).

- DISTRIBUTION: Bhutan.

bordonii Assing, 1999: 13 ( Othius ; Type locality: Kouy-Tchéou, Rég. de Pin-Fa).

- DISTRIBUTION: China?.

bovinus Coquerel, 1860 , see: laeviusculus Stephens, 1833 .

brachypterus Wollaston, 1864: 580 ( Othius ; Type locality: Gomera).

— Wollaston, 1865: 499 ( Othius ; habitat; Canary Islands).

— Fauvel, 1902b: 99 ( Othius ; Canary Islands).

— Bernhauer and Schubert, 1914: 317 ( Othius ; catalog).

— Coiffait, 1954: 165 ( Othius ; Canary Islands).

— Coiffait, 1956a: 41 ( Othius ; characters in key).

— Coiffait, 1972: 354 ( Othius ; characters; references; Canary Islands).

— Palm, 1976: 212 ( Othius ; characters; Canary Islands).

— Assing, 1997a: 104, 112 ( Othius ; characters; notes; Canary Islands).

— Assing, 1998a: 215 ( Othius ; Canary Islands).

— Assing, 1999a: 346 ( Othius ; Canary Islands).

— Assing, 1999c: 668 ( Othius ; Canary Islands).

- DISTRIBUTION: Canary Islands.

brevicornis Wollaston, 1857 , see: jansoni Wollaston, 1854 .

brevipennis Kraatz, 1857d: 657 ( Othius ; Type locality: Oesterreich; [ Note : See lectotype designation by Assing, 1997]).

— Redtenbacher, 1857: 982 ( Othius ; characters; Austria).

— Fauvel, 1873b: 85 [= 1873a: 371] ( Othius ; characters; Austria).

— Redtenbacher, 1874: 223 ( Othius ; characters; Austria).

— Ganglbauer, 1895: 472 ( Othius ; characters; eastern Alps).

— Reitter, 1908: 107 ( Othius ; characters; Austrian Alps).

— Bernhauer and Schubert, 1914: 317 ( Othius ; catalog).

— Scheerpeltz, 1933: 1321 ( Othius ; catalog).

— Walles, 1938: 67, 70 ( Othius ; Poland).

— Scheerpeltz and Höfler, 1948: 156 ( Othius ; fungus inhabitant).

— Smetana, 1958: 130 ( Othius ; characters; ecology; Czechoslovakia).

— Lohse, 1964: 169 ( Othius ; characters; central Europe).

— Horion, 1965: 114 ( Othius ; Germany; Austria; Italy; Slovenia; Poland; Slovakia).

— Coiffait, 1972: 352 ( Othius ; characters; references; Europe).

— Bordoni, 1982: 388 ( Othius ; characters; Italy).

— Frank, 1982: 31 ( Othius ; list of parasites).

— Lucht, 1987: 103 ( Othius ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland).

— Tóth, 1989: 39 ( Othius ; characters; Hungary).

— Janák, 1993a: 47 ( Othius ; Slovakia).

— Ciceroni and Zanetti, 1995: 27 ( Othius ; Italy).

Assing, 1997: 93 ( Othius ; lectotype designation; characters; notes; Germany; Austria; Czech Republic; Slovakia; Hungary; Slovenia; Croatia; Bosnia-Herzegovina) .

— Assing and Solodovnikov, 1998: 305 ( Othius ; characters; Poland).

— Assing and Schülke, 1999: 12 ( Othius ; characters).

— Assing, 1999c: 677 ( Othius ; notes; Austria; Slovakia; Slovenia; Italy; Bosnia Herzegovina).

- DISTRIBUTION: Poland, Germany, Austria, Czech Republic, Slovakia, Hungary, Slovenia, Croatia, Bosnia Hercegovina.

dilutus Motschulsky, 1858a: 210 ( Othius ; Type locality: Styrie; [Note: See lectotype designation by Assing, 1997]).

— Fauvel, 1873b: 83 [= 1873a: 369] ( Othius ; synonym of myrmecophilus ).

— Mulsant and Rey, 1877a: 228 ( Othius ; synonym of mymecophilus).

— Ganglbauer, 1895: 472 ( Othius ; synonym of brevipennis ).

— Reitter, 1908: 107 ( Othius ; synonym of brevipennis ).

— Bernhauer and Schubert, 1914: 317 ( Othius ; synonym of brevipennis ).

— Smetana, 1958: 130 ( Othius ; synonym of brevipennis ).

— Coiffait, 1972: 352 ( Othius ; synonym of brevipennis ).

— Bordoni, 1982: 388 ( Othius ; synonym of brevipennis ).

— Assing, 1997: 93 ( Othius ; lectotype designation; synonym of brevipennis ).

brookesi Cameron, 1952 , see: Maorothius .

cachemiricus Coiffait, 1982a: 252 ( Othius ; Type locality: Cachemire: Daksum, 2400–

2700 m).

— Assing, 1998: 331 ( Othius ; characters; India).

- DISTRIBUTION: India.

californicus Mannerheim, 1843 , see: Parothius .

casarum Gistel, 1857: 79 ( Othius ; [Note: Nomen dubium]; Type locality: Alpes Tyrolis, Salisburgi, et Helvetiae).

- DISTRIBUTION: Austria, Switzerland.

castaneipennis Gistel, 1857: 69 ( Othius ; [Note: Nomen dubium]; Type locality: Tyrolis alpes).

- DISTRIBUTION: Austria.

chongqingensis Zheng, 1995 , see: latus Sharp, 1874 .

chrysurus Reitter, 1891b: 195 ( Othius ; Type locality: Taschkent; [Note: See lectotype designation by Assing, 1997]).

— Reitter, 1908: 105 ( Othius ; characters; Tashkent).

— Bernhauer and Schubert, 1914: 317 ( Othius ; catalog).

— Scheerpeltz, 1933: 1321 ( Othius ; catalog).

— Coiffait, 1972: 341 ( Othius ; characters; Turkestan).

Assing, 1997: 104 ( Othius ; lectotype designation; characters; notes; Tajikistan; Uzbekistan; Kazakhstan) .

— Assing, 1999c: 679 ( Othius ; correct measurements for TiL and TaL cited in Assing, 1997, by factor of 2.02; collecting notes; Tajikistan; Uzbekistan; Kirghizia).

- DISTRIBUTION: Kazakhstan, Uzbekistan, Tajikistan, Kyrgyzstan.

clavifer Assing, 1998: 319 ( Othius ; Type locality: Nepal: Sankhua Sabha Distr. , Thudam).

— Assing, 1999c: 684 ( Othius ; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).

- DISTRIBUTION: Nepal.

coiffaiti Lohse, 1963 , see: philonthoides Wollaston, 1864 .

collapsus Assing, 1999: 78 ( Othius ; Type locality: Taiwan: Nantou Hsien, near Tsifeng ,

2300 m).

- DISTRIBUTION: Taiwan.

coniceps Assing, 1999: 83 ( Othius ; Type locality: Taiwan: Nantou Hsien, near Hohuanshan , 3100 m).

- DISTRIBUTION: Taiwan.

conifer Assing, 1998: 315 ( Othius ; Type locality: Nepal: Taplejung Distr. , pass Deorali

W Yamputhin, 3400 m).

— Assing, 1999c: 684 ( Othius ; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).

- DISTRIBUTION: Nepal.

contumax Assing, 1999: 81 ( Othius ; Type locality: Taiwan: Nantou Hsien, Nenkaoshan, Tenchi Hut , 2890 m).

- DISTRIBUTION: Taiwan.

corniger Assing, 1998: 321 ( Othius ; Type locality: Nepal: Taplejung Distr. , upper Simbua Khola Valley, near Tseram, 3250–3350 m).

— Assing, 1999c: 684 ( Othius ; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).

- DISTRIBUTION: Nepal.

corpulentus Coiffait, 1972: 340 , 343 ( Othius ; subspecies of transsilvanicus ; Type locality:

Mt. Sarco, Hu. merid.; [Note: See lectotype designation by Assing, 1997]. Originally cited from sud des Karpathes).

Assing, 1997: 87, 129 ( Othius ; lectotype designation: Mt. Sarco , Hu. merid.; valid species; characters; notes; Romania) .

— Assing, 1999c: 676 ( Othius ; Romania; Serbia?).

- DISTRIBUTION: Romania.

crassus Motschulsky, 1858a: 210 ( Othius ; Type locality: Alpes de la Carniole, Doga-Niva; [Note: See lectotype designation by Assing, 1997]).

— Fauvel, 1873b: 83 [= 1873a: 369] ( Othius ; synonym of lapidicola ).

— Ganglbauer, 1895: 472 ( Othius ; characters; eastern Alps; north Carpathians).

— Reitter, 1908: 107 ( Othius ; characters; Carpathians; Caucasus).

— Bernhauer and Schubert, 1914: 317 ( Othius ; catalog).

— Scheerpeltz, 1933: 1322 ( Othius ; catalog).

— Smetana, 1958: 130 ( Othius ; characters; ecology; Czechoslovakia).

— Lohse, 1964: 169 ( Othius ; characters; central Europe).

— Horion, 1965: 113 ( Othius ; Austria; Italy; Poland; Slovakia; Romania).

— Coiffait, 1972: 351 ( Othius ; characters; references; Alps; Carpathians; France; Caucasus).

— Bordoni, 1982: 386 ( Othius ; characters; Italy).

— Lucht, 1987: 103 ( Othius ; checklist; Germany; Poland; Czechoslovakia; Austria).

— Tóth, 1989: 38 ( Othius ; characters; Hungary).

— Janák, 1993a: 47 ( Othius ; Slovakia).

— Ciceroni and Zanetti, 1995: 27 ( Othius ; Italy).

Assing, 1997: 73 ( Othius ; lectotype designation; characters; notes; Austria; Slovenia; probably restricted range and citations from France, the Caucasus and other regions and countries are probably misidentifications) .

— Assing and Schülke, 1999: 14 ( Othius ; characters).

— Assing, 1999c: 674 ( Othius ; notes; Austria; Slovenia).

- DISTRIBUTION: Austria, Slovenia.

pallidus Brancsik, 1874: 135 ( Othius ; Type locality: Styria; [Note: See lectotype designation by Assing, 1997]. Originally cited from Bachergebirge bei Marburg in Steiermark).

— Ganglbauer, 1895: 472 ( Othius ; synonym of crassus ).

— Reitter, 1908: 107 ( Othius ; synonym of pallidus ).

— Bernhauer and Schubert, 1914: 317 ( Othius ; synonym of crassus ).

— Smetana, 1958: 130 ( Othius ; synonym of crassus ).

— Coiffait, 1972: 352 ( Othius ; synonym of crassus ).

— Bordoni, 1982: 386 ( Othius ; synonym of crassus ).

— Assing, 1997: 73 ( Othius ; lectotype designation: Styria; synonym of crassus ).

deceptivus Cuvier, 1833 , see: punctulatus Goeze, 1777 .

deharvengi Coiffait, 1982: 53 ( Othius ; Type locality: Nepal: versant sud-est du Col de Goropani , ouest de Pokhara, 2150 m).

— Assing, 1998: 329 ( Othius ; characters; Nepal).

— Assing, 1999c: 669 ( Othius ; Nepal).

- DISTRIBUTION: Nepal.

discrepans Assing, 1999: 45 ( Othius ; Type locality: China: Sichuan, Ziling Mt. , 1600–2400 m).

- DISTRIBUTION: China.

dilutus Motschulsky, 1858 , see: brevipennis Kraatz, 1857 .

dimidiatus Motschulsky, 1860 , see: Atrecus .

donisthorpei Chitty, 1903 [Note: Nomen nudum], see: punctulatus Goeze, 1777 .

elongatus Coiffait, 1956 , see: lapidicola Märkel and Kiesenwetter, 1848 .

excisus Assing, 1999: 46 ( Othius ; Type locality: China: S. Sichuan, S. Xichang, Mt. Luoji , 2300–2500 m).

- DISTRIBUTION: China.

extraordinarius Assing, 1998: 303 ( Othius ; Type locality: N-Indien: Himach. Pr., Umg. Kufri,

ca. 2600 m).

— Assing, 1999c: 684 ( Othius ; correct measurements for TiL and TaL cited in Assing, 1998, by

factor of 2.02).

- DISTRIBUTION: India. fastigatus Assing and Solodovnikov, 1998: 301 ( Othius ; Type locality: Georgia: Caucasus

Swanetien).

- DISTRIBUTION: Georgia. fibulifer Assing, 1999: 53 ( Othius ; Type locality: China: W Sichuan, Daxue Shan, E Theto-La

[recte: Tsheto-La] Pass, W. Kangding, 3500 m, 30.00.42 N, 101.51.47 E).

— Assing, 1999c: 683 ( Othius ; Sichuan).

- DISTRIBUTION: China. flavocaudatus Cameron, 1926b: 346 ( Othius ; Type locality: Chakrata District, Sijla Gad,

5000 feet; [Note: See lectotype designation by Assing, 1998]).

— Cameron, 1932: 42 ( Othius ; characters; India).

— Scheerpeltz, 1933: 1322 ( Othius ; catalog).

— Assing, 1998: 301 ( Othius ; characters; lectotype designation: Chakrata District, Sijla Gad,

5000 feet; Nepal; India).

— Assing, 1999c: 684 ( Othius ; correct measurements for TiL and TaL cited in Assing, 1998, by

factor of 2.02).

- DISTRIBUTION: Nepal, India. fulminans Gravenhorst, 1802 , see: punctulatus Goeze, 1777 . fulvipennis Fabricius, 1793 , see: punctulatus Goeze, 1777 . fulvipes Motschulsky, 1858 , see: Gabrius . fulvopterus Geoffroy, 1785 , see: punctulatus Goeze, 1777 . fuscicornis Heer, 1841 , see: laeviusculus Stephens, 1833 . gansuensis Assing, 1999 , see: latus Sharp, 1874 . glabricornis Stephens, 1833 , see: Atrecus . goui Zheng, 1995: 342 , 346 ( Othius ; Type locality: Sichuan: Wolong, 1830 m).

- DISTRIBUTION: China. grandis Hochhuth, 1849: 99 ( Othius ; Type locality: Cauc. Gotsch.; [Note: See lectotype desig-

nation by Gusarov, 1992a]. Originally cited from Lenkoran; Transkaukasien; Taurien).

— Hochhuth, 1862: 69 ( Othius ; Russia).

— Reitter, 1891b: 195 ( Othius ; characters; Caucasus).

— Reitter, 1908: 105 ( Othius ; characters; Caucasus; Lenkoran; Crimea).

— Bernhauer and Schubert, 1914: 317 ( Othius ; catalog).

— Scheerpeltz, 1933: 1322 ( Othius ; catalog).

— Coiffait, 1965e: 120 ( Othius ; Turkey).

— Pototskaia, 1967: 48 ( Othius ; larval characters).

— Bolov, 1969a: 328 ( Othius ; collecting notes; Russia).

— Coiffait, 1972: 342 ( Othius ; characters; references; Russia; Turkey).

— Petrenko, 1980: 81 ( Othius ; Caucasus).

— Gusarov, 1992a: 784 [= 1993a: 73] ( Othius ; lectotype designation: Cauc. Gotsch.).

— Assing, 1997: 55 ( Othius ; characters; notes; Turkey; Georgia; Russia).

— Solodovnikov, 1998a: 16 ( Othius ; notes; Russia, Krasnodar territory).

— Assing, 1999c: 672 ( Othius ; correct measurements for TiL and TaL cited in Assing, 1997, by

factor of 2.02; Caucasus).

— Solodovnikov, 2000a: 324 ( Othius ; notes; Turkey).

- DISTRIBUTION: Turkey, Azerbaijan, Georgia, Russia, Ukraine. hamatus Assing, 1999: 70 ( Othius ; Type locality: Taiwan: Kaohsiung Hsien, Peinantashan

trail, 2250 m).

- DISTRIBUTION: Taiwan.

hebes Assing and Solodovnikov, 1998: 299 ( Othius ; Type locality: Armenia: Caucasus , Armen. Geb. Also cited from Georgia and Azerbaijan).

— Assing, 1999c: 678 ( Othius ; Caucasus).

- DISTRIBUTION: Armenia, Georgia, Azerbaijan, Caucasus.

ibericus Coiffait, 1956 , see: punctulatus Goeze, 1777 .

infestus Assing, 1998: 311 ( Othius ; Type locality: Nepal: Taplejung Distr. , upper Simbua Khola Valley, near Tseram, 3250–3350 m).

— Assing, 1999c: 684 ( Othius ; correct measurements for TiL and TaL cited in Assing, 1998, by factor of 2.02).

- DISTRIBUTION: Nepal.

intermedius Korge, 1962: 151 ( Othius ; Type locality: Kanarische Inseln: Teneriffa: Montana Cagancho , ca. 1600 m).

— Coiffait, 1972: 357 ( Othius ; characters; Canary Islands).

— Palm, 1976: 214 ( Othius ; characters; Canary Islands).

— Assing, 1997a: 107, 113 ( Othius ; characters; notes; Canary Islands).

— Assing, 1998a: 216 ( Othius ; Canary Islands).

- DISTRIBUTION: Canary Islands.

jadwigae Roubal, 1911: 5 ( Othius ; Type locality: Kr. Poljana; [Note: See lectotype designation by Assing, 1997]).

— Bernhauer and Schubert, 1914: 317 ( Othius ; cited as jadvigae; catalog).

— Coiffait, 1972: 356 ( Othius ; characters; Caucasus).

— Assing, 1997: 61 ( Othius ; lectotype designation; characters; notes; Caucasus).

— Assing, 1999c: 684 ( Othius ; correct measurements for TiL and TaL cited in Assing, 1997, by factor of 1.54).

— Solodovnikov, 2000a: 328 ( Othius ; notes; Georgia).

- DISTRIBUTION: Russia, Caucasus.

jaegeri Assing, 1998: 334 ( Othius ; Type locality: Nepal: Telbrung Danda , 3700 m,

SE Annapurna mts.).

- DISTRIBUTION: Nepal.

jansoni Wollaston, 1854: 576 ( Othius ; Type locality: Madeira: Ribeiro de Santa Luzia;

Pico d’Arribentao above Funchal; [Note: See lectotype designation by Assing and Wunderle, 1995a]).

— Wollaston, 1857: 187 ( Othius ; characters; habitat; Madeira Islands).

— Wollaston, 1865: 499 ( Othius ; habitat; notes; Madeira Islands).

— Fauvel, 1902b: 99 ( Othius ; Madeira).

— Bernhauer and Schubert, 1914: 317 ( Othius ; catalog).

— Smetana, 1963b: 36 ( Othius ; Madeira).

— Coiffait, 1972: 354 ( Othius ; characters; references; Madeira).

— Palm, 1979: 271 ( Othius ; characters; collecting notes; Madeira).

Assing and Wunderle, 1995a: 58 ( Othius ; lectotype designation: [Note: Locality not cited]; bionomics; characters; Madeira) .

— Assing, 1998a: 215 ( Othius ; Madeira).

— Assing, 1999c: 672 ( Othius ; Madeira).

- DISTRIBUTION: Madeira.

vestitus Wollaston, 1857: 186 ( Othius ; Type locality: Madeira: Lombo dos Pecegueiros; Cruzinhas at more than 4000 ft.).

— Fauvel, 1902b: 99 ( Othius ; synonym of jansoni ).

— Bernhauer and Schubert, 1914: 317 ( Othius ; synonym of jansoni ).

— Coiffait, 1972: 354 ( Othius ; synonym of jansoni ).

— Palm, 1979: 272 ( Othius ; valid species; collecting notes; characters; Madeira).

— Assing and Wunderle, 1995a: 58 ( Othius ; synonym of jansoni ).

brevicornis Wollaston, 1857: 187 ( Othius ; Type locality: Madeira: Dezerta Grande ).

— Wollaston, 1865: 499 ( Othius ; notes; Madeira Islands).

— Fauvel, 1902b: 99 ( Othius ; synonym of jansoni ).

— Lucht, 1987: 103 ( Othius ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).

— Tóth, 1989: 35 ( Othius ; characters; Hungary).

— Ciceroni, 1993: 76 ( Othius ; Italy).

— Scheerpeltz, 1964a: 299 ( Othius ; Crete).

— Welch, 1993: 228 ( Othius ; ovariole number and ovary structure).

— Janák, 1993a: 47 ( Othius ; Czech Republic; Slovakia).

— Sparacio, 1995: 159 ( Othius ; characters; notes; Sicily).

— Ciceroni and Zanetti, 1995: 27 ( Othius ; Italy).

— Vorst, 1996 ( Othius ; notes on biology and distribution; Netherlands).

Assing, 1997: 34 ( Othius ; lectotype designation: [Note: Locality not cited]; characters; notes; Great Britain; Morocco; Algeria; Tunisia; Portugal; Spain; Balearic Islands; France; Germany; Switzerland; Austria; Italy; Poland; Czech Republic; Slovakia; Hungary; Slovenia; Croatia; Bosnia-Herzegovina; Bulgaria; Macedonia; Greece; Cyprus; Turkey; Syria; Caucasus ) .

— Assing and Schülke, 1999: 9 ( Othius ; characters).

— Assing, 1999c: 667 ( Othius ; Morocco; Algeria; Spain; France; Germany; Italy; Austria; Czech Republic; Bosnia Herzegovina; Yugoslavia; Macedonia; Greece; Cyprus; Turkey; Iran).

— Solodovnikov, 2000a: 325 ( Othius ; notes; Russia).

— Assing and Wunderle, 2001: 36 ( Othius ; Cyprus).

- DISTRIBUTION: Tunisia, Algeria, Morocco, Europe, Russia, Azerbaijan, Turkey,

Syria, Iran.

punctipennis Lacordaire, 1835: 409 ( Staphylinus ; Type locality: Paris).

— Erichson, 1839b: 296 ( Othius ; characters; Austria; Switzerland; Italy; France; England).

— Heer, 1841: 581 ( Othius ; characters; Switzerland).

— Hochhuth, 1849: 100 ( Othius ; characters; Lenkoran).

— Redtenbacher, 1849: 690 ( Othius ; characters; Austria).

— Rosenhauer, 1856: 70 ( Othius ; Malaga).

— Fairmaire and Laboulbène, 1856: 498 ( Othius ; characters; France).

— Kraatz, 1857d: 655 ( Othius ; characters; Germany).

— Redtenbacher, 1857: 179 ( Othius ; characters; Austria).

— Kraatz, 1858a: 65 ( Othius ; Greece).

— G. Waterhouse, 1858: 26 ( Othius ; synonym of laeviusculus ).

— Hochhuth, 1862: 69 ( Othius ; Russia).

— Fauvel, 1869: 490 ( Othius ; synonym of laeviusculus ).

— Fauvel, 1873b: 84 [= 1873a: 370] ( Othius ; synonym of laeviusculus ).

— Redtenbacher, 1874: 222 ( Othius ; characters; Austria).

— Mulsant and Rey, 1877a: 234 ( Othius ; characters; France).

— Reitter, 1908: 106 ( Othius ; synonym of laeviusculus ).

— Bernhauer and Schubert, 1914: 317 ( Othius ; synonym of laeviusculus ).

— Scheerpeltz, 1933: 1322 ( Othius ; synonym of laeviusculus ).

— Smetana, 1958: 126 ( Othius ; synonym of laeviusculus ).

— Coiffait, 1972: 344 ( Othius ; synonym of laeviusculus ).

— Bordoni, 1982: 391 ( Othius ; synonym of laeviusculus ).

— Assing, 1997: 34 ( Othius ; synonym of laeviusculus ).

fuscicornis Heer, 1841: 581 ( Othius ; Type locality: Aigle).

— Kraatz, 1857d: 655 ( Othius ; synonym of punctipennis ).

— Reitter, 1908: 106 ( Othius ; synonym of laeviusculus ).

— Bernhauer and Schubert, 1914: 317 ( Othius ; synonym of laeviusculus ).

— Smetana, 1958: 126 ( Othius ; synonym of laeviusculus ).

— Coiffait, 1972: 344 ( Othius ; synonym of laeviusculus ).

— Bordoni, 1982: 391 ( Othius ; synonym of laeviusculus ).

— Assing, 1997: 34 ( Othius ; synonym of laeviusculus ).

bovinus Coquerel, 1860: 154 ( Quedius ; Type locality: Mers-el-Kébir).

— Fauvel, 1869: 486 ( Othius ; synonym of laeviusculus ).

— Fauvel, 1873b: 84 [= 1873a: 370] ( Othius ; synonym of laeviusculus ).

— Fauvel, 1902b: 99 ( Othius ; synonym of laeviusculus ).

— Bernhauer and Schubert, 1914: 317 ( Othius ; synonym of laeviusculus ).

— Scheerpeltz, 1933: 1322 ( Othius ; synonym of laeviusculus ).

— Coiffait, 1972: 344 ( Othius ; synonym of laeviusculus ).

— Bordoni, 1982: 391 ( Othius ; synonym of laeviusculus ).

— Assing, 1997: 34 ( Othius ; synonym of laeviusculus ).

lapidicola Märkel and Kiesenwetter, 1848a: 321 ( Othius ; Type locality: Kärnthner Alpen; [Note: See lectotype designation by Assing, 1997]).

— Fairmaire and Laboulbène, 1856: 498 ( Othius ; characters; France).

— Kraatz, 1857d: 657 ( Othius ; characters; Germany).

— Fauvel, 1865a: 299 [= 1865b: 49] ( Othius ; habitat; France).

— Fauvel, 1873b: 83 [= 1873a: 369] ( Othius ; characters; France; Germany; Switzerland; Austria; Italy; Scandinavia).

— Fauvel, 1874: 338 [= 1874b: 44] ( Othius ; Lapland; Spain).

— Redtenbacher, 1874: 222 ( Othius ; characters; Austria).

— J. Sahlberg, 1876: 31 ( Othius ; characters; Finland).

— Mulsant and Rey, 1877a: 225 ( Othius ; characters; France).

— Ganglbauer, 1895: 471 ( Othius ; characters; north and middle Europe).

— Reitter, 1909: 133 ( Othius ; characters; Germany).

— Bernhauer and Schubert, 1914: 318 ( Othius ; catalog).

— Portevin, 1929: 368 ( Othius ; characters; France).

— Scheerpeltz, 1933: 1322 ( Othius ; catalog).

— V. Hansen, 1952: 26 ( Othius ; characters).

— Smetana, 1958: 127 ( Othius ; characters; ecology; Czechoslovakia).

— Palm, 1963b: 56 ( Othius ; characters; Sweden; Norway; Finland).

— Lohse, 1964: 168 ( Othius ; characters; central Europe).

— Horion, 1965: 111 ( Othius ; Canaries; Turkey; Russia; Finland; Sweden; Norway; France; Switzerland; Czech Republic; Slovakia; Poland; Bosnia Hercegovina; Montenegro; Serbia; Macedonia; Bulgaria; Greece; Italy; Spain; Austria; Germany).

— Coiffait, 1966: 22 ( Othius ; Turkey).

— Coiffait, 1972: 347 ( Othius ; characters; references; Europe; Caucasus; Spain; Italy; Balkans; Turkey).

— Coiffait, 1978d: 163 ( Othius ; Turkey).

— Bordoni, 1982: 392 ( Othius ; characters; Italy).

— Frank, 1982: 31 ( Othius ; list of parasites).

— Lucht, 1987: 103 ( Othius ; checklist; Germany; Poland; Czechoslovakia; Austria; Sweden).

— Tóth, 1989: 36 ( Othius ; characters).

— Nowosad, 1990: 101 ( Othius ; collected from mole nest; notes; Poland).

— Ciceroni, 1993: 76 ( Othius ; Italy).

— Wittwer, 1993: 249 ( Othius ; Switzerland).

— Janák, 1993a: 47 ( Othius ; Czech Republic; Slovakia).

— Hodge and Jones, 1995: 37 ( Othius ; characters; Britain).

— Ciceroni and Zanetti, 1995: 27 ( Othius ; Italy).

Assing, 1997: 8 ( Othius ; lectotype designation; characters; notes; Norway; Sweden; Finland; Russia; France; Germany; Switzerland; Austria; Italy; Czech Republic; Slovenia; Hungary; Romania; Croatia; Bosnia-Herzegovina; Yugoslavia; Macedonia; Albania; Bulgaria; Greece; Turkey; Armenia) .

— Uhlig, 1998: 136 ( Othius ; note).

— Assing and Schülke, 1999: 10 ( Othius ; characters).

— Assing, 1999c: 663 ( Othius ; notes on elevation of collections; Finland; Russian Federation; France; Germany; Switzerland; Austria; Italy; Czech Republic; Slovakia; Romania; Croatia; Bosnia Herzegovina; Bulgaria; Greece; Turkey; Georgia; Caucasus).

— Solodovnikov, 2000a: 325 ( Othius ; notes; Turkey).

- DISTRIBUTION: Europe, Georgia, Armenia, Turkey.

suturalis Motschulsky, 1858a: 211 ( Othius ; [preoccupied]; Type locality: Alpes de la Carniole, Koschutna).

— Fauvel, 1873b: 83 [= 1873a: 369] ( Othius ; synonym of lapidicola ).

— Ganglbauer, 1895: 471 ( Othius ; synonym of lapidicola ).

— Bernhauer and Schubert, 1914: 318 ( Othius ; synonym of lapidicola ).

— Smetana, 1958: 127 ( Othius ; synonym of lapidicola ).

— Coiffait, 1972: 347 ( Othius ; synonym of lapidicola ).

— Bordoni, 1982: 392 ( Othius ; synonym of lapidocola).

— Assing, 1997: 8 ( Othius ; synonym of lapidicola ).

longicornis Thomson, 1871: 369 ( Othius ; Type locality: Norrland).

— Fauvel, 1873b: 83 [= 1873a: 369] ( Othius ; synonym of lapidicola ).

— J. Sahlberg, 1876: 31 ( Othius ; synonym of lapidicola ).

— Ganglbauer, 1895: 471 ( Othius ; synonym of lapidicola ).

— Bernhauer and Schubert, 1914: 318 ( Othius ; synonym of lapidicola ).

— Smetana, 1958: 127 ( Othius ; synonym of lapidicola ).

— Coiffait, 1972: 347 ( Othius ; synonym of lapidicola ).

— Bordoni, 1982: 392 ( Othius ; synonym of lapidicola ).

— Assing, 1997: 8 ( Othius ; synonym of lapidicola ).

albanicus Reitter, 1918: 154 ( Othius ; Type locality: Albanien: Miloti ; [Note: See lectotype designation by Assing, 1997]).

— Scheerpeltz, 1933: 1321 ( Othius ; catalog).

— Coiffait, 1968b: 93 ( Othius ; notes; Greece).

— Coiffait, 1972: 346 ( Othius ; characters; Albania; Greece; Turkey; Bulgaria).

— Assing, 1997: 8, 128 ( Othius ; lectotype designation; synonym of lapidicola ).

elongatus Coiffait, 1956b: 139 ( Othius ; Type locality: Turquie d’Asie , Villayet d’Afyon: Dinar. Also cited from Slovakia).

— Lohse, 1963: 177 ( Othius ; synonym of lapidicola ).

— Lohse, 1964: 168 ( Othius ; synonym of lapidicola ).

— Coiffait, 1965e: 120 ( Othius ; cited as valid species; Turkey).

— Coiffait, 1972: 343 ( Othius ; characters; Turkey).

— Assing, 1997: 8 ( Othius ; synonym of lapidicola ).

rhodicus Coiffait, 1976b: 85 ( Othius ; Type locality: Rhodes: Monte Profitis Ilias, 800 m). — Assing, 1997: 8 ( Othius ; synonym of lapidicola ).

kastamonuensis Bordoni, 1976: 234 ( Othius ; Type locality: Turchia: vil. Kastamonu,

Ilgaz dag gecidi, 1800–2300 m).

— Assing, 1997: 8 ( Othius ; synonym of lapidicola ).

latus Sharp, 1874: 51 ( Othius ; Type locality: Japan; [Note: See lectotype designation by Assing, 1999]. Originally cited from Copper Temple , Nagasaki; northern China).

— Bernhauer and Schubert, 1914: 318 ( Othius ; catalog).

— Scheerpeltz, 1933: 1322 ( Othius ; catalog).

— Ito, 1993a: 141 ( Othius ; characters; Japan).

— J. Li, 1993: 32 ( Othius ; China).

Assing, 1999: 20 ( Othius ; lectotype designation: Japan; characters; Russia; Japan; China) .

- DISTRIBUTION: Japan, China, Russia.

stoetzneri Bernhauer, 1931: 1 ( Othius ; Type locality: Szetschwan: Kwanshien; [Note: See lectotype designation by Assing, 1999]).

— Scheerpeltz, 1933: 1323 ( Othius ; catalog).

— Assing, 1999: 20 ( Othius ; lectotype designation; synonym of latus latus ).

ozakii Ito, 1993a: 143 ( Othius ; subspecies of latus ; Type locality: Japan: Aomori Pref.: Jyogakura, Mt. Hakkoda).

— Assing, 1999: 20 ( Othius ; synonym of latus latus ).

chongqingensis Zheng, 1995: 343 , 346 ( Othius ; Type locality: Sichuan: Mt. Jin-yun, Chongqing City , 800–919 m).

— Assing, 1999: 20 ( Othius ; synonym of latus latus ).

gansuensis Assing, 1999: 27 ( Othius ; subspecies of latus ; Type locality: China: Gansu,

28 km S. Lanzhou, 2200 m).

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Philonthus

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

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