Cephennomicrus muluensis, Jałoszyński, Paweł, 2015

Jałoszyński, Paweł, 2015, Four new species of Cephennomicrus Reitter (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia, Zootaxa 3911 (2), pp. 273-279 : 278-279

Cephennomicrus muluensis sp. n.

( Figs. 4 , 8 , 15–16 )

Type material. Holotype: EAST MALAYSIA (Borneo: Sarawak): ♂, two labels: " MALAYSIA: Sarawak, / Mulu NP, 100 km SEE / Niri, 200 m, 19.-24.VIII / 2003, leg. A. SCHULZ" [white, printed]; " Cephennomicrus / muluensis m. / det. P. Jałoszyński, '14 / HOLOTYPUS " [red, printed] ( SMNS).

Diagnosis. BL nearly 1 mm; pronotum with antebasal groove and two pairs of pits; each elytron with six macrosetae; aedeagus in ventral view with sides parallel from base to apical third, where median lobe gradually narrows to form subtriangular apex; dorsal wall in subapical region with numerous dense and short setae.

Description. BL 0.95 mm. Body of male ( Figs. 4 , 8 ) elongate and strongly convex, brown with slightly lighter macrosetae.

Head broadest at posterior margin of short but distinct tempora, HL 0.15 mm, HW 0.25 mm; eyes large, strongly convex and coarsely faceted; vertex and frons confluent and convex; supraantennal tubercles feebly marked. Punctures on head dorsum deep and distinct, separated by spaces subequal to diameters of punctures; setae indiscernible under magnification 80x. Antennae slender and with club composed of enlarged antennomeres X–XI, AnL 0.43 mm, antennomeres I–II elongate; III–VIII about as broad as long; IX distinctly transverse; X elongate; XI longer than X and about twice as long as broad; antennomeres X–XI with several strongly erect, long setae among short and suberect basic setation.

Pronotum ( Fig. 8 ) semioval, broadest near middle, at insertion of median macroseta; PL 0.28 mm, PW 0.39 mm; anterior and lateral margins broadly rounded; anterior corners visible only in anterodorsal view, distinct and sharp-angled but with rounded apices; posterior margin slightly arcuate; posterior corners obtuse; pronotal base with two lateral pairs of small but distinct pits and transverse groove laterally connected to internal pits. Punctures on pronotal disc similar to those on frons and vertex; pronotal disc only with macrosetae: two lateral pairs and one posterior pair in front of mesoscutellum.

Elytra ( Fig. 8 ) oval, broadest near middle; EL 0.53 mm, EW 0.45 mm, EI 1.17; humeral calli well-marked, elongate. Punctures on elytra much less distinct than those on pronotum and head, similarly dense but smaller and shallower; each elytron only with macrosetae: four lateral, one apical and one subapical. Hind wings well developed.

Legs moderately long and slender, unmodified.

Aedeagus ( Figs. 15–16 ) in ventral view elongate and approximately drop shaped with flattened base; AeL 0.16 mm; median lobe with subtriangular apical part; dorsal wall in subapical area with short and dense setae; internal armature symmetrical, lightly sclerotized, without long projections; parameres slender and exceeding apex of median lobe, recurved, each with one apical and one subapical seta.

Female. Unknown.

Distribution. East Malaysia, Sarawak.

Etymology. Locotypical, after Mount Mulu.

Remarks. This remarkable species resembles members of the enigmatic Cephennomicrus pinguis group from Southeast Asia that may be placed in a separate genus in the future. The aedeagus of C. muluensis has a group of dense setae present in a subapical dorsal area, a character rarely occurring in the genus. A similar setal patch, located closer to the basal aedeagal orifice, can be found in C. gumnos Jałoszyński , a species known from Papua New Guinea ( Jałoszyński 2010c). Despite clear similarities in genital structures, the external morphology of these two species is strikingly different: C. gumnos is a large species (BL 1.18–1.19 mm) with a very stout body that is strongly constricted between the pronotum and elytra, a nearly impunctate posterior third of the pronotum and elytra, a pronotum without the transverse antebasal pronotal groove and with five thin and curved lateral macrosetae on each elytron.

Figures

FIGURES 1 – 4. Dorsal habitus of holotype males. 1 ― Cephennomicrus curvidens sp. n.; 2 ― Cephennomicrus gentingensis sp. n.; 3 ― Cephennomicrus santubongensis sp. n.; 4 ― Cephennomicrus muluensis sp. n.

FIGURES 5 – 8. Body outline and distribution of macrosetae. 5 ― Cephennomicrus curvidens sp. n.; 6 ― Cephennomicrus gentingensis sp. n.; 7 ― Cephennomicrus santubongensis sp. n.; 8 ― Cephennomicrus muluensis sp. n.

FIGURES 9 – 16. Aedeagi of holotype males in ventral (9, 11, 13, 15) and lateral (10, 12, 14, 16) views. 9 – 10 ― Cephennomicrus curvidens sp. n.; 11 – 12 ― Cephennomicrus gentingensis sp. n.; 13 – 14 ― Cephennomicrus santubongensis sp. n.; 15 – 16 ― Cephennomicrus muluensis sp. n.