Bensonella lakainguta Hwang, 2014

Páll-Gergely, Barna & White, Tom S., 2023, Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae), Journal of Natural History 56 (45 - 48), pp. 2011-2029 : 2025-2026

Bensonella lakainguta Hwang, 2014

( Figures 7–8 , 9 (e–h))

Boysidia (Bensonella) plicidens – Pilsbry, 1917 (in Pilsbry 1916 –1918): 198 (partim), pl. 34, figs 3, 9–10

Boysidia (Bensonella) plicidens – Habe 1956: 109, figs 9–11

Bensonella plicidens – Minato 1988: 41

Bensonella plicidens – Schileyko 1998: 139, fig. 159

Bensonella plicidens – Maassen 1999: 126

Bensonella plicidens lakainguta Hwang, 2014: 18 , figs 2–4

Boysidia (Bensonella) qingliangfengensis F. Fang, J. Wang and Y. Chen, 2015: 692 , fig. 1 new synonym

Material examined

NHMUK 201901108 , 12 shells, Omi, Japan, Hirase coll. 457 ; NHMUK 20191128 , three shells, Rijozen , Omi, Japan ; NHMW-MO-38313 (1 shell), Mt. Fujiwara, 300 m, Japan, coll. Blume ex coll. Azuma , 24 September 1951 ; NHMW-MO-55810 (5 shells), Japan, Mie Pref., Fujiwara-dake , 300 m, coll. Edlauer ex coll. Kuiper ; NHMW-MO-38645 (4 shells), Yoro, Mino ; NHMW-MO-69480 (2 shells), Riozen, Omi, Japan, coll. Oberwimmer ex coll. Jetschin ; NHMW-MO-113726 (3 shells), Omi, Japan, coll. Rušnov ; HNHM 105323 (figured shell), Japan, Kochi Prefecture, Kami-shi, Ryugado Prefectural National Park , above the cave, 500 m along a side-road, 290 m, 33.601917°N, 133.746217°E, Leg. Hunyadi, A., Murányi, D. and Páll-Gergely, B., 08 August 2016.

Remarks

In the original description, Hwang (2014) mentioned three differences between Indian B. plicidens and the subspecies Bensonella plicidens lakainguta . The most important of them was the submarginal elevations of columellar, basal and palatal barriers, i.e. anterior to the hooks those barriers become more elevated. However, this trait can also be seen in Indian B. hooki sp. nov. ( Figure 9 (c,d)); see differences between B. lakainguta and B. hooki sp. nov. outlined under the description of the latter.

Boysidia (Bensonella) qingliangfengensis ( Figure 7 (e,f)) was described from the border region of the Chinese Anhui and Zhejiang Provinces (very close to Pilsbry̾s (1917) record from Hangzhou), and is claimed to be larger than B . plicidens, yet looks identical for all other characters. Therefore , it is treated here as a junior synonym of B . lakainguta Hwang, 2014.

Maassen (1999) mentioned that he compared Bensonella karoensis with Japanese shells of Bensonella plicidens (= Bensonella lakainguta ). Furthermore, he erroneously stated that Bensonella plicidens inhabited Thailand.

Figures

Figure 7. Bensonella lakainguta Hwang, 2014. a–c: Holotype of Bensonella lakainguta (NMNS-7244-00); D: Bensonella lakainguta Hwang, 2014, Japan,Kochi Prefecture, Kami-shi,Ryugado Prefectural National Park, above the cave, 500 m along a side-road, 290 m, 33.601917°N, 133.746217°E, Leg. A. Hunyadi, D. Murányi and B. Páll-Gergely, 8 October 2016 (HNHM 105323); e–f: Holotype of Boysidia (Bensonella) qingliangfengensis F. Fang, J. Wang and Y. Chen, 2015. Photos: a–c and e–f: images from the original descriptions, d: B. Páll-Gergely. Figs e and f are not to scale.

Figure 8. Bensonella lakainguta Hwang, 2014, Japan, Omi, NHMUK 201901108, specimen 1. All photos: B. Páll-Gergely.

Figure 9. a, b: Bensonella plicidens (Benson, 1849), NHMUK 56.9.15.69A, specimen 1 (same as Figure 4); c,d: Bensonella hooki sp. nov., holotype, NHMUK 20191114 (same as Figure 6); e,f: Bensonella lakainguta Hwang, 2014, HNHM 105323 (same as Figure 7(d)); g, h: Bensonella lakainguta, NHMUK 201901108 (same as Figure 8). All images: B. Páll-Gergely.