Neobisnius Ganglbauer 1895

Chani-Posse, Mariana, Newton, Alfred F., Hansen, Aslak Kappel & Solodovnikov, Alexey, 2018, Checklist and taxonomic changes for Central and South American Philonthina (Coleoptera: Staphylinidae), Zootaxa 4449 (1), pp. 1-95 : 45-50

Neobisnius Ganglbauer 1895

Neobisnius Ganglbauer 1895: 464 . Type species: Gabrius villosulus Stephens, 1833 , designated by Lucas (1920: 434).

Notes: Neobisnius is a worldwide distributed genus including 72 species at present. Among them, 35 species have been reported from the Neotropical region. The New World species of the genus were revised by Frank (1981).

Neobisnius aeneipennis Bruch 1915 , nomen nudum

Neobisnius aeneipennis Bruch 1915: 500 , attributed to Bernhauer in litteris; nomen nudum. Locality: Catamarca. Distribution: Argentina. Note: not in Fernández et al. (2007) or Bachmann et al. (2017).

Neobisnius armuellensis Bierig 1933

Neobisnius armuellensis Bierig 1933: 56 . Type locality: Puerto Armuelles, Panamá. Type depository: FMNH 4 ST. Distribution: Costa Rica, Panama, Colombia. Note: redescription, distribution ( Frank 1981: 28).

Neobisnius brasilianus Wendeler 1956

Neobisnius brasilianus Wendeler 1956: 230 . Type locality: Brasilien, Nova Teutonia . Type depository: ZMHB (Wendeler) HT ♀ [ MCP seen]. Distribution : Brazil (Santa Catarina) . Note: redescription, distribution ( Frank 1981: 25); distribution within Brazil (Asenjo et al. 2013).

Neobisnius concolor ( Sharp 1885)

Actobius concolor Sharp 1885: 458 . Type locality: Panama, Volcan de Chiriqui 4000–6000 ft, Peña Blanca 3000– 4000 ft. Type depository: BMNH 13 ST: 10 Chiriqui, 1 Peña Blanca [MCP seen]; FMNH 2 ST: 2 Chiriqui. Distribution: Panama;?Mexico. Note: comb. nov. ex Actobius ( Bernhauer & Schubert 1914: 322) ; status dubious, may be synonym of N. sobrinus Erichson , redescription, distribution ( Frank 1981: 27).

Neobisnius demmeli Bierig 1933

Neobisnius demmeli Bierig 1933: 54 . Type locality: cercanía de Cayajabos, Sierra del Rosario, Prov. P. del Rio, Cuba. Type depository: FMNH 2 ST [orig. 5]; USNM 1 ST. Distribution: Cuba. Note: revised status, ex synonym of N. humilis Erichson , redescription, distribution ( Frank 1981: 32); distribution within Cuba ( Peck 2005).

Neobisnius edznai Frank 1981

Neobisnius edznai Frank 1981: 44 . Type locality: Edzná, Campeche, Mexico, 150 ft. Type depository: BMNH HT ♂, etc. Distribution: Mexico, Guatemala. Note: distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Neobisnius facilis ( Sharp 1885)

Actobius facilis Sharp 1885: 458 . Type locality: San Jerónimo, Baja Verapaz, Guatemala. Type depository: BMNH HT ♂: from San “Gerónimo” [MCP seen]. Distribution: Mexico, Guatemala, El Salvador, Nicaragua. Notes: comb. nov. ex Actobius ( Bernhauer & Schubert 1914: 323) ; redescription, distribution ( Frank 1981: 37); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Neobisnius flavipes Bierig 1933

Neobisnius flavipes Bierig 1933: 50 . Type locality: Puerto Armuelles , Panamá . Type depository: FMNH? HT, #?PT [not seen]. Distribution : Panama. Note: redescription, distribution ( Frank 1981: 23).

Neobisnius flavomaculatus Bernhauer 1908

Neobisnius flavomaculatus Bernhauer 1908: 336 . Type locality: Santa Catarina, Brazil & Grenada, W. I. Type depository: FMNH 3 ST. Distribution: Colombia, Bolivia, Brazil, Paraguay, Uruguay, Grenada;? Argentina. Notes: Argentina distribution ( Bruch 1928: 444); redescription, distribution Grenada ( Frank 1981: 15); distribution within Brazil (Asenjo et al. 2013).

= Neobisnius flavomaculatus Bruch 1915: 501 , attributed to Bernhauer in litteris (see flavomaculatus Bernhauer 1908: 336 ). Locality: San Luis. Distribution: Argentina. Notes: there are two specimens deposited at MACN under this name, one from San Luis and the other one from Buenos Aires ( Argentina). The specimen from San Luis is, without any doubt, the one referred by Bruch (1915). It does not agree with the original description of the species ( Bernhauer 1908: 336) and it is N. parcepunctatus Bernhauer. The specimen from Buenos Aires is N. flavomaculatus indeed.

Neobisnius fortis ( Sharp 1885)

Actobius fortis Sharp 1885: 457 . Type locality: Panama, Volcan de Chiriqui 4000–6000 ft, Bugaba 800–1500 ft. Type depository: BMNH 2 ST: 1 Chiriqui, 1 Bugaba [MCP seen]. Distribution: Panama, Colombia, Brazil, Argentina. Note: comb. nov. ex Actobius ( Bernhauer & Schubert 1914: 323) ; redescription, distribution ( Frank 1981: 36); distribution within (and including) Brazil (Asenjo et al. 2013)

Neobisnius fraternus Bernhauer 1921

Neobisnius fraternus Bernhauer 1921c: 105 . Type locality: Jujuy prov., Argentina . Type depository: FMNH 1 ST. Distribution : Argentina. Notes: redescription, distribution ( Frank 1981: 29).

Neobisnius funerulus Cameron 1922

Neobisnius funerulus Cameron 1922: 119 , attributed to Fauvel in litteris. Type locality: St. Vincent. Type depository: BMNH HT. Distribution: Lesser Antilles ( St. Vincent, St. Lucia). Notes: redescription, distribution ( Frank 1981: 34); distribution within Lesser Antilles ( St. Vincent) ( Peck 2010).

Neobisnius gratus ( LeConte 1863)

Philonthus gratus LeConte 1863: 38 . Type locality: Colorado River, near Fort Yuma, California [North America]. Type depository: MCZ LT + 4 PLT. Distribution: USA, Mexico;? Argentina. Notes: as Actobius ( Horn, 1884: 231) ; comb. nov. ex Actobius ( Bernhauer 1912b: 176) ; lectotype designation, redescription, distribution ( Frank 1981: 16); Argentina ( Bruch 1928: 444).

Neobisnius humilis ( Erichson 1840)

Philonthus humilis Erichson 1840: 512 . Type locality: ins. Puerto Rico. Type depository: ZMHB LT ♂, 3 PLT. Distribution: Cuba, Jamaica, Hispaniola, Puerto Rico, Cayman Is., Virgin Is.;? Grenada. Notes: comb. nov. ex Philonthus ( Bernhauer & Schubert 1914: 323) ; lectotype designation ( Smetana 1963: 4); redescription, distribution ( Frank 1981: 18, not including Grenada); distribution, also within Cuba ( Peck 2005); distribution within (and including) Cayman Is. (Frank et al. 2011); Grenada and other Lesser Antilles ( Blackwelder 1944, but not confirmed by Frank (1981), probably refers to former synonym.

= Neobisnius limbatus Cameron 1922: 118 . Type locality: Haiti. Type depository: BMNH (Cameron) 4 ST (as “paratypes”, see Frank 1981: 18). Distribution: Haiti. Note: as synonym of N. humilis ( Blackwelder 1943) .

Neobisnius jocosus ( Horn 1884)

Actobius jocosus Horn 1884: 232 . Type locality: Delaware; North Carolina [no specific locality]. Type depository: MCZ LT ♀. Distribution: USA (e), Mexico, Guatemala, Honduras, El Salvador. Notes: comb. nov. ex Actobius ( Bernhauer & Schubert 1914: 323) ; lectotype designation, redescription, distribution ( Frank 1981: 27); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

= Actobius alternans Sharp 1885: 461 . Type locality: Guatemala, San Gerónimo, Torola, Paso Antonio; Paso Antonio, 400 ft., Escuintla, Guatemala (LT). Type depository: BMNH LT, 3 PLT, additional 4 PLT?: 2 San Gerónimo, 2 Torola [MCP seen]. Notes: comb. nov. ex Actobius ( Bernhauer & Schubert 1914: 322) ; lectotype designation, as synonym of N. jocosus ( Frank 1981: 27) .

Neobisnius lathrobioides (Baudi di Selve, 1848) [adventive in CASA]

Philonthus lathrobioides Baudi di Selve 1848: 130 . Type locality: Pedemontio. Type depository: MRSN/MCSN? #? ST. Distribution: Europe, Egypt, Tunisia, Algeria, Morocco, Turkey, Azerbaijan, Iran; adventive in Azores, Canary Is., Madeira, Canada, USA, Chile, Australia, Madagascar. Notes: lectotype designation; valid species ( Bordoni 1975: 82); redescription, distribution ( Frank 1981: 9).

Neobisnius ludicrus ( Erichson 1840)

Philonthus ludicrus Erichson 1840: 514 . Type locality: Guyana Venezuelensi. Type depository: ZMHB LT ♂, 1 PLT. Distribution: USA, Mexico, Guatemala, Honduras, Costa Rica, Panama, Colombia, Venezuela, Bolivia, Brazil, Trinidad, Virgin Is., Barbados, Grenada, St. Vincent, St. Lucia. Notes: comb. nov. ex Philonthus ( Bernhauer & Schubert 1914: 323) ; lectotype designation, redescription and distribution ( Frank 1981: 20); distribution, also within Mexico ( Navarrete-Heredia et al. 2002), Barbados ( Peck 2009a), Lesser Antilles ( St. Vincent) ( Peck 2010); Brazil (Asenjo et al. 2013).

= Philonthus umbripennis LeConte 1863: 38 . Type locality: Middle states [ USA] . Type depository: MCZ HT ♀. Note : new synonym of N. ludicrus ( Frank 1981: 20) .

= Philonthus deletus Sharp 1876a: 167 . Type locality: Tapajos [Pará, Brazil]. Type depository: BMNH 4 ST: from Tapajos [MCP seen]. Distribution: Brazil. Note: comb. nov. ex Philonthus , as synonym of N. ludicrus ( Frank 1981: 20) .

= Neobisnius fauveli Smetana 1963: 2 (proposed for N. agnatus of Horn 1884: 229 (misidentification)). Type locality: Louisiana, USA. Type depository: IRSNB HT ♂, 1PT. Note: treated as a subspecies of N. ludicrus by Frank (1981), for dark specimens from Texas to northwest Florida, but treated as an implicit or explicit synonym of N. ludicrus by subsequent authors (e.g., Frank 1986, Navarrete-Heredia et al. 2002, respectively).

Neobisnius maculicollis Scheerpeltz 1972

Neobisnius maculicollis Scheerpeltz 1972: 107 . Type locality: Buenos Aires: Ezeiza Argentina . Type depository: HNHM HT. Distribution : Argentina. Note: not in Frank (1981).

Neobisnius maximus Bernhauer 1910

Neobisnius maximus Bernhauer 1910: 383 . Type locality: Cordoba, Veracruz, Mexico. Type depository: FMNH 1 ST. Distribution: Mexico. Note: redescription, distribution ( Frank 1981: 37, only type locality); distribution within Mexico ( Navarrete-Heredia et al. 2002).

Neobisnius mixtus ( Sharp 1885)

Actobius mixtus Sharp 1885: 460 . Type locality: San Jerónimo, Baja Verapaz, Guatemala. Type depository: BMNH LT ♂, 2 PLT: FMNH 3 PLT. Distribution: Mexico, Guatemala, Panama. Note: comb. nov. ex Actobius ( Bernhauer & Schubert 1914: 323) ; lectotype designation, redescription, distribution ( Frank 1981: 24); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Neobisnius moestus ( Sharp 1887)

Actobius moestus Sharp 1887: 790 . Type locality: Mexico (near the city), Mexico. Type depository: BMNH 2 ST. Distribution: Mexico. Note: comb. nov. ex Actobius ( Bernhauer & Schubert 1914: 323) ; redescription, distribution ( Frank 1981: 45); distribution within Mexico ( Navarrete-Heredia et al. 2002).

Neobisnius nigrocaeruleus Cameron 1922

Neobisnius nigrocaeruleus Cameron 1922: 118 . Type locality: Haiti. Type depository: BMNH (Cameron) #? ST. Distribution: Cuba, Jamaica, Haiti, Dominican Rep. Notes: redescription, distribution ( Frank 1981: 26); distribution, also within Cuba ( Peck 2005).

= Neobisnius carbonarius Bierig 1933: 49 . Type locality: Cayajabos, Sierra del Rosario, Prov. P. del Rio, Cuba. Type depository: FMNH HT ♂ . Distribution: Cuba. Note: as synonym of N. nigrocoeruleus ( Frank 1981) .

Neobisnius nitidulus ( Sharp 1885)

Actobius nitidulus Sharp 1885: 460 . Type locality: Guatemala City, Guatemala. Type depository: BMNH LT, 3+ PLT; FMNH 2 PLT. Distribution: Mexico, Guatemala, El Salvador. Note: comb. nov. ex Actobius ( Bernhauer & Schubert 1914: 323) ; lectotype designation, redescription, distribution ( Frank 1981: 34); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

= Actobius politus Sharp 1885: 461 . Type locality: Mexico. Type depository: BMNH HT ♀. Distribution: Mexico. Note: comb. nov. ex Actobius ( Bernhauer & Schubert 1914: 323) ; as synonym of N. nitidulus ( Frank 1981) .

Neobisnius occidentoides Frank 1981

Neobisnius occidentoides Frank 1981: 47 . Type depository: BMNH HT ♂, etc. Type locality: Lake Corpus Cristi St. Pk., San Patricio Co., Texas, USA. Distribution: Canada, USA, Mexico. Notes: distribution ( Brunke & Marshall 2011); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Neobisnius omnirufus Frank 1981

Neobisnius omnirufus Frank 1981: 31 . Type depository: MCZ HT ♂, etc. Type locality: Argentina: Mendoza . Distribution: Argentina.

Neobisnius parcepunctatus Bernhauer 1912

Neobisnius parcepunctatus Bernhauer 1912b: 176 . Type locality: Argentinien, Chaunar-Region . Type depository: FMNH 4 ST. Distribution : Argentina. Note: redescription, distribution ( Frank 1981: 12).

Neobisnius richteri Bernhauer 1927

Neobisnius richteri Bernhauer 1927: 245 . Type locality: Buenos Aires prov. Type depository: FMNH 1 ST. Distribution: Argentina, Brazil (Santa Catarina). Note: redescription, distribution ( Frank 1981: 35); distribution within Brazil (Asenjo et al. 2013).

Neobisnius sanguinipennis Scheerpeltz 1972

Neobisnius sanguinipennis Scheerpeltz 1972: 110 . Type locality: Rio Negro: El Bolsón, 350m. Type depository: HNHM HT. Distribution: Argentina. Note: not in Frank (1981).

Neobisnius scutellaris Bernhauer 1908

Neobisnius scutellaris Bernhauer 1908: 336 . Type locality: Paraguay. Type depository: FMNH 1 ST. Distribution: Argentina, Uruguay, Bolivia, Paraguay, Brazil. Note: distribution ( Frank 1981: 30); distribution within Brazil (Asenjo et al. 2013).

Neobisnius semipunctatus ( Fairmaire & Germain 1862)

Othius semi-punctatus Fairmaire & Germain 1862: 434. Type locality: Santiago [Chile]. Type depository: MNHN? #?ST. Distribution: Chile. Notes: status, types, Chile only, other localities misidentified ( Fagel 1954); redescription ( Frank 1981: 10); distribution ( Coiffait & Sáiz 1968, Frank 1981).

Neobisnius semirufus Bernhauer, 1921

Neobisnius semirufus Bernhauer 1921c: 105 . Type locality: Salta prov., Argentina. Type depository: FMNH HT. Distribution: Argentina, Bolivia. Note: redescription, distribution ( Frank 1981: 33).

Neobisnius senilis ( Horn 1884)

Actobius senilis Horn 1884: 226 . Type locality: Garland, Costilla Co., Colorado, USA. Type depository: MCZ LT ♂, 2 PLT. Distribution: Canada, Mexico. Notes: comb. nov. ex Actobius ( Bernhauer & Schubert 1914: 323) ; lectotype designation, redescription ( Frank 1981: 45); distribution ( Frank 1981, Navarrete-Heredia et al. 2002); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

= Philonthus piger Horn 1884: 219 (in key). Note: This name appeared in a key to some species of Philonthus , with characters but without other data, but in "Additions and notes" at the end of the same publication Horn (1884: 240) indicated this name was a synonym of Actobius senilis described in the same work (see above).

Neobisnius simplex ( Sharp 1885)

Actobius simplex Sharp 1885: 459 . Type locality: Mirador, Veracruz, Mexico. Type depository: BMNH HT ♂. Distribution: Mexico, Guatemala, El Salvador, Costa Rica, Panama, Brazil. Note: comb. nov. ex Actobius ( Bernhauer & Schubert 1914: 323) ; redescription, distribution ( Frank 1981: 42); distribution, also within Mexico ( Navarrete-Heredia et al. 2002); distribution within Brazil (Asenjo et al. 2013).

= Neobisnius cavifrons Bierig 1933: 51 . Type locality: Puerto Armuelles , Chiriquí, Panama . Type depository: FMNH HT, #?PT; SDEI 1 +PT. Distribution : Panama. Note: as synonym of N. simplex ( Frank 1981) .

Neobisnius sobrinus ( Erichson 1840)

Philonthus sobrinus Erichson 1840: 512 . Type locality: Pensylvania. Type depository: ZMHB LT, 3 PLT. Distribution: Canada, USA, Mexico, Guatemala, Costa Rica, Colombia, Venezuela, Brazil. Notes: comb. nov. ex Philonthus ( Bernhauer & Schubert 1914: 323) ; lectotype designation ( Smetana 1963: 4); redescription and distribution ( Frank 1981: 39); distribution, also within Mexico ( Navarrete-Heredia et al. 2002); distribution within Brazil (Asenjo et al. 2013).

= Philonthus agnatus Erichson 1840: 513 . Type locality: valle Araguensi Columbiae [Rio Araguas in Santander]. Type depository: ZMHB LT ♂, 3 PLT. Distribution: Colombia. Notes: comb. nov. ex Philonthus ( Bernhauer & Schubert 1914: 322) ; lectotype designation ( Smetana 1963: 4); as synonym of N. sobrinus ( Frank 1981) .

= Actobius ocreatus Horn 1884: 228 . Type locality: California. Type depository: MCZ LT ♂ . Distribution : USA: California. Notes: lectotype designation, as synonym of N. sobrinus ( Frank 1981) .

= Actobius arduus Sharp 1885: 458 . Type locality: Puebla, Mexico. Type depository: BMNH LT ♂, 14+ PLT. Distribution: Mexico. Notes: comb. nov. ex Actobius ( Bernhauer & Schubert 1914: 322) ; lectotype designation, as synonym of N. sobrinus ( Frank 1981) .

= Neobisnius simulator Smetana 1963: 3 . Type locality: Curling, W Newfoundland, Canada . Type depository: CNC (not MZLU as originally stated) HT ♀. Distribution : Canada (Newfoundland). Notes : type deposition correction, as synonym of N. sobrinus ( Frank 1981) .

Neobisnius terminalis ( LeConte 1863) Philonthus terminalis LeConte 1863: 38 . Type locality: Maryland [North America]. Type depository: MCZ LT + 1

PLT? (see Frank 1981: 13). Distribution: Canada, USA, Mexico, Guatemala, Costa Rica, Panama. Notes: as Neobisnius ( Bierig 1933: 56) ; lectotype designation, redescription ( Frank 1981: 13); distribution ( Frank 1981, Brunke & Marshall 2011, Navarrete-Heredia et al. 2002).

= Actobius elegantulus Horn 1884: 232 . Type locality: Arizona. Type depository: MCZ LT ♂. Distribution: USA: widespread (as subspecies of N. terminalis ). Notes: lectotype designation, as subspecies of N. terminalis ( Frank 1981) ; as synonym of N. terminalis (e.g., Navarrete-Heredia et al. 2002).

= Actobius delicatulus Sharp 1885: 461 . Type locality: Guatemala, Pantaleón, 1700 feet, Paso Antonio, 400 feet; Pantaleón, Escuintla, Guatemala, 1700 ft. Type depository: BMNH 4 ST: 2 Pantaleón, 2 San Antonio [MCP seen]. Distribution: Guatemala. Notes: comb. nov. ex Actobius ( Bernhauer & Schubert 1914: 322) ; as synonym of N. terminalis ( Frank 1981) .

= Actobius adustus Sharp 1887: 790 . Type locality: Tehuantepec, Oaxaca, Mexico. Type depository: BMNH 8 ST [MCP seen]. Distribution: Mexico. Notes: comb. nov. ex Actobius ( Bernhauer & Schubert 1914: 322) ; as synonym of N. terminalis ( Frank 1981) .

Neobisnius vigii Frank 1981

Neobisnius vigii Frank 1981: 43 . Type depository: CNC HT ♂, ZMHB 1 PT. Type locality: Colombia: Madgalena , Parque Tayrona, 21 mi E of Santa Marta . Distribution : Colombia. Note: PT from " La Vigi ", not located.