Calolydella omissa Fleming & Wood

Fleming, AJ, Wood, D. Monty, Smith, M. Alex, Hallwachs, Winnie & Janzen, Daniel H, 2018, Revision of the Mesoamerican species of Calolydella Townsend (Diptera: Tachinidae) and description of twenty-three new species reared from caterpillars in Area de Conservacion Guanacaste, northwestern Costa Rica, Biodiversity Data Journal 6, pp. 11223-11223 : 11223

Calolydella omissa Fleming & Wood sp. n.

Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016605 ; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0016605; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-43825, BOLD:AAD9874, ASTAP809-07; Taxon: scientificName: Calolydellaomissa; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: omissa; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Cabanya; verbatimElevation: 340; verbatimLatitude: 10.877; verbatimLongitude: -85.2308; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Letis buteoDHJ03; verbatimEventDate: 02-Nov-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: 06-SRNP-43825 ; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: 06-SRNP-43825; individualCount: 5; sex: 3M, 2F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 06-SRNP-43825; Taxon: scientificName: Calolydellaomissa; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: omissa; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Cabanya; verbatimElevation: 340; verbatimLatitude: 10.877; verbatimLongitude: -85.2308; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Letis buteoDHJ03; verbatimEventDate: 02-Nov-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Description

Male (Fig. 26a, b, c). Length: 7-8mm. Head (Fig. 26b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, with a single row of fine setulae outside of frontal setae; parafacial silver throughout. Thorax (Fig. 26a, c): gold on dorsal surface, silver laterally (>50% coverage); with four regular thoracic vittae; postpronotum with four setae; 3:3 acrostichal setae; 2:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite bare or rarely with up to two hair-like setae; scutellar discal setae situated as wide apart as subapical scutellar setae. Wing vein R4+5 with 6-7 setulae dorsally, reaching from base almost to crossvein R-M. Abdomen (Fig. 26a): with uninterrupted transverse marginal pollinose bands; abdominal pollinosity gold dorsally, silver ventrally, and with an orange spot lateroventrally at base of ST1+2; T3 with one pair of median marginal setae, lacking discal setae; T4 with one pair of discal setae. Terminalia sternite 5 (Fig. 27c): with two small lobes and a wide U-shaped median cleft, 0.45X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; entire lobe of sternite with short setae, of varying lengths. Cerci (Fig. 27b), in dorsal view, separated by a narrow gap but rejoining at apex; each cercus long and very slightly tapered, almost of equal width along entire length; cercus slender with an upward bend when viewed laterally; setose along its entire length. Surstylus (Fig. 27a) slender and digitiform with a slight upward curve, appearing 0.2X shorter than cercus when viewed laterally; with short setae along entire length; tip of surstylus very slightly curved inwards when viewed dorsally.

Female (Fig. 26d, e, f). Length: 5-6mm. Fronto-orbital plate 1.4X wider than in male.

Diagnosis

Calolydella omissa can be distinguished from all other species of Calolydella by the following combination of traits: fronto-orbital plate gold, parafacial silver, abdomen with uninterrupted transverse silver pollinose bands across all tergites, anatergite bare (rarely setose), and T3 without discal setae.

Etymology

The specific epithet is derived from the Latin adjective " omissa ", meaning dropped or absent, in reference to the absence of discal setae on T3, a character state unique to this species.

Distribution

Costa Rica, ACG, Alajuela Province, Cabanya, 340m.

Ecology

Calolydella omissa has been reared once from Letis Janzen03 ( Lepidoptera : Erebidae ), in rain forest.