Cymbionotum (Subgenus),

Ball, George E. & Shpeley, Danny, 2005, Taxonomic review of the Tribe Melaenini (Coleoptera: Carabidae), with observations on morphological, ecological and chorological evolution, Zootaxa 1099 (1), pp. 1-120: 1-120

publication ID 10.11646/zootaxa.1099.1.1

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Cymbionotum (Subgenus)


Procoscinia, new subgenus 

Type species. Cymbionotum fernandezi  , n. sp., (here designated).

Subgeneric name. Based on two Latin terms: the prefix pro, meaning before, implying primitiveness; and Coscinia  , the original (but homonymous) name of the genus now recognized as Cymbionotum  . Thus, the subgeneric name implies a generalized member of the genus within which it is included.

Recognition. Precinctive in the Neotropical Region of the Western Hemisphere, adults of Procoscinia  have the posteriolateral dentiform projections distinctly anteriad the posterior margin of the pronotum. The posterior margin of the pronotum is not beaded, and the proepipleuron (pep) is not visible from a dorsal perspective ( Fig. 14AView FIGURE 14; cf. Fig. 14BView FIGURE 14). Males have sclerite x of the endophallus about one third length of phallus, and in infolded position near base of phallus ( Fig. 15FView FIGURE 15, scx).

Descriptive notes. With character states of genus Cymbionotum  , restricted as follows. Size and ratios of body parts ( Tables 3–9). Average size (SBL and EW, Tables 3 and 4) for genus intermediate, but broad overlap with schueppelii  group, and distinctly smaller than members of basale  and semelederi  species groups. Pronotum relatively broad, values for PL/PWM ( Table 7) more than 0.90; base broad, values for ratio PWB/PWA ( Table 9) more than 0.65.

Color. Dorsal surface uniformly rufous, or head and pronotum slightly darker than elytra. Appendages: antennae, labrum and mandibles rufous; palpi testaceous; femora, tibiae and tarsi rufotestaceous.

Microsculpture. Dorsal surface with mesh pattern isodiametric, though microlines very fine and in places effaced, or microlines absent; scutellum with mesh pattern evident throughout, more or less isodiametric, sculpticells convex, surface beaded.

Vestiture. Dorsal surface with setation very dense ( Figs. 2AView FIGURE 2, 3FView FIGURE 3), setae of moderate length ( Table 1 and Fig. 2EView FIGURE 2)

Head. Clypeus: anterior margin medially without a dentiform projection, or distinct swelling.

Antennae. Antennomere 2 pyriform (cf. Fig. 11DView FIGURE 11), and relatively longer (about two thirds length of antennomere 3, Table 11); antennomeres 5–10 moderately asymmetrical

(cf. Fig. 11FView FIGURE 11)

Labrum. Anterior margin distinctly, but broadly concave.

Included taxa. This subgenus includes two species: C. fernandezi  , new species, and C. negrei Perrault. 

Habitat. The members of Procoscinia  occupy the tropical rain forest vegetation zone (Table 17), but may be confined to riparian situations.

Geographical distribution. The range of Procoscinia  ( Fig. 16View FIGURE 16) is confined to northern South America.

Relationships and chorological affinities. This subgenus is the putative, primitive adelphotaxon of subgenus Cymbionotum  , from which it is widely isolated (by the South Atlantic Ocean) geographically ( Figs. 16View FIGURE 16, 18, 20, 21, 23).