Cymbionotum (Procoscinia) fernandezi, Ball and Shpeley

Ball, George E. & Shpeley, Danny, 2005, Taxonomic review of the Tribe Melaenini (Coleoptera: Carabidae), with observations on morphological, ecological and chorological evolution, Zootaxa 1099 (1), pp. 1-120: 1-120

publication ID 10.11646/zootaxa.1099.1.1

publication LSID


persistent identifier

treatment provided by


scientific name

Cymbionotum (Procoscinia) fernandezi, Ball and Shpeley

n. sp.

Cymbionotum (Procoscinia) fernandezi, Ball and Shpeley   , n. sp.

( Figs. 1B View FIGURE 1 , 2A, 2E View FIGURE 2 , 3F View FIGURE 3 , 12A–I View FIGURE 12 , 13 A–J View FIGURE 13 , 14A View FIGURE 14 , 15A–E View FIGURE 15 , 16 View FIGURE 16 )

Type material. Thirty eight specimens (20 males and 17 females). HOLOTYPE male, card­mounted, labeled: " COLOMBIA Bolivar/ Zambrano bosque seco/ Lote Papayo cultivo/ Gmelina arborea   . Luz/ 10–15. V. 1999 /Col. H.M.­M.A.­M.G./ No. 967597". A microvial, pinned beneath the whole specimen, contains its genitalia (UNCB). PARATYPES, 4 labeled same as holotype: male, point­mounted, head and prothorax missing (broken and parts lost, following for habitus picture, Fig. 1B View FIGURE 1 ); female, pointmounted, reproductive tract removed, in microvial pinned beneath whole specimen; female, card­mounted, with left elytron removed, and attached to card; and female, cardmounted, disassembled for SEM photographs of mouthparts and ovipositor sclerites). Eighteen males, 15 females, labeled: " COLOMBIA Bolivar Zam­/brano. Hda. Monterrey For­/estal BsT 9¼42'N, 74¼53'W/ 20m. May 1999 "; "Lote Papayo Interior de/ cultivo Gmelina arborea   / trampa de luz,/ H. Moreno y M. çvila/ No. 967543" With the agreement of H. Moreno, the paratypes were distributed as follows: BMNH, 2; CASC, 2; CNIF, 7; IAVH, 7; MNHP, 2; MRAC, 2; UASM, 2; UNCB, 11; and UASM, 2   .

Specific epithet. This Latinized eponym, genitive case, is based on the surname of Prof. Fernando Fernandez, who loaned the type specimens to us, and with whom we have cooperated in extending knowledge of Colombian Carabidae   .

Type locality. Colombia: Bolivar : Zambrano   .

Recognition. Compared to C. negrei   , adults of C. fernandezi   are on average, larger, both in length and width, with wider vertex, wider head in relation to pronotum, and wider pronotal base ( Table 12). The phalli ( Figs. 15D, F View FIGURE 15 ) of these two species are similar in form, but that of C. fernandezi   ( Fig. 15B View FIGURE 15 ) is broader, with the preapical portion of the shaft bent more gradually ventrad, and ventral surface generally dark (cf. Fig. 15F View FIGURE 15 ).

Description. With recognition features as noted above, and other details specified in the subgeneric description, and illustrations, as follows: habitus, Fig. 1B View FIGURE 1 ; integumentary setal length, Fig. 2E View FIGURE 2 ; density of elytral punctation, Fig. 3F View FIGURE 3 ; labrum and epipharynx, Figs. 12I and J View FIGURE 12 , respectively; mandibles, Figs. 12A–H View FIGURE 12 ; maxillae, Figs. 12K–L View FIGURE 12 ; labium, Figs. 13A–C View FIGURE 13 ; pronotum, dorsal aspect, Fig. 14A View FIGURE 14 ; hind wings, Fig. 15A View FIGURE 15 ; middle tibia and tarsomere 5, Figs. 13D and E View FIGURE 13 , respectively; phallus and endophallic armature, Fig. 15B View FIGURE 15 ; gonocoxa of ovipositor, Figs. 13F–J View FIGURE 13 ; internal genitalia of female, Fig. 15E View FIGURE 15 .

Habitat, habits, and seasonal occurrence. All specimens were collected in May, at an ultraviolet light, within a plantation of an exotic tree ( Gmelina arborea Roxb.   , white teak, family Verbenaceae   ), a species indigenous in the Oriental Region and southern China. We infer that C. fernandezi   is mesophilic, occupying disturbed areas in the tropical rain forest zone (Table 17, Fig 24), at less than 200 m altitude with adults active in midyear.

Geographical distribution ( Fig. 16 View FIGURE 16 ). This species is known only from the Rio Magdalena valley, Colombia.

Relationships and chorological affinities. Based on shared morphological features, geographical distribution and allopatric ranges, C. fernandezi   and C. negrei   are postulated to be adelphotaxa.

Material examined. We have seen the type series, only. For details, see " Type material", above.