Melaenini

Ball, George E. & Shpeley, Danny, 2005, Taxonomic review of the Tribe Melaenini (Coleoptera: Carabidae), with observations on morphological, ecological and chorological evolution, Zootaxa 1099 (1), pp. 1-120: 1-120

publication ID

http://doi.org/ 10.11646/zootaxa.1099.1.1

publication LSID

lsid:zoobank.org:pub:CA0A0ED3-D49D-4198-A409-407EF86A164E

persistent identifier

http://treatment.plazi.org/id/227887BE-FFFF-FFAF-250D-F7CCFA88FBC4

treatment provided by

Felipe

scientific name

Melaenini
status

 

Key to the genera, subgenera and species of Melaenini  

1. Head dorsally with single supraorbital seta over each eye. Dorsal surface of body with fixed setae only, without general setose vestiture. Antennomeres 1–4 with setae near distal margin of each article. Elytron with striae distinctly impressed, intervals convex ................................. Melaenus Dejean   ................ 2

– Head dorsally with two supraorbital setae over each eye. Dorsal surface of body with general setose vestiture. Antennomeres 1–4 with setae over surface of each article. Elytron with striae shallow (stria 1), or indistinct; intervals flat............... ......................................................................... Cymbionotum Baudi di Selve   ... 3

2 (1). Pronotum each side with two posteriolateral projections. Specimen from locality in Africa ( Fig. 10 View FIGURE 10 )......................................................... M. elegans Dejean   , p. 39

– Pronotum each side with one posteriolateral projection. Specimen from locality in India ( Fig. 10 View FIGURE 10 ) ........................................................... M. piger Fabricius   , p. 38

3 (1) Antennomere 2 ( Fig. 11E, A View FIGURE 11 2 View FIGURE 2 ) globose, spherical in outline, not pyriform; antennomeres 5–10 ( Fig. 11G, A View FIGURE 11 6 View FIGURE 6 , A 7 View FIGURE 7 ) markedly asymmetric ............................ ................................................................................ semelederi   species group ... 4

– Antennomere 2 ( Fig. 11D, A View FIGURE 11 2 View FIGURE 2 ) pyriform, margins markedly constricted proximad base; antennomeres 5–10 ( Fig. 11F, A View FIGURE 11 6 View FIGURE 6 , A 7 View FIGURE 7 ) moderately asymmetric........ 6

4 (3). Elytral vestiture denser ( Figs. 2C View FIGURE 2 , 3B View FIGURE 3 ) ................. C. semelederi Chaudoir   , p. 68

– Elytral vestiture sparser ( Fig. 2D View FIGURE 2 , 3A View FIGURE 3 ) .............................................................. 5

5 (4). Phallus in left lateral aspect markedly curved preapically and endophallus with two microtrichial fields (Fig. 17J). Specimen from locality in or near Afghanistan (Fig. 18) ................................................................. C. striatum Reitter   , p. 73

– Phallus in left lateral aspect gradually curved preapically and endophallus with single microtrichial field (Fig. 17K). Specimen from locality in or near Iran (Fig. 18) ........................................................................ C. mandli Jedlička   , p. 74

6 (3). Dorsal surface uniformly dark rufous. Pronotum ( Fig. 14A View FIGURE 14 ) with posteriolateral angles distinctly anteriad posterior margin, proepipleuron not visible. Specimen from locality in Western Hemisphere (northern South America— Fig. 16 View FIGURE 16 )........... ............................................................... Procoscinia, new subgenus   ............... 7

Dorsal surface various in color: uniformly dark concolorous to uniformly pale concolorous, to bicolored (Figs. 17A–C, 17F­H, 19A­B, 19E­F, 19I­J, 19L­N, 22A­C). Pronotum ( Fig. 14B View FIGURE 14 ) with posteriolateral angles near posterior margin, proepipleuron visible. Specimen from locality in Eastern Hemisphere (Afrotropical, southern Palaearctic , Oriental Regions­ Figs. 16 View FIGURE 16 , 20, 21, 23) .................... ...................................................... Subgenus Cymbionotum (sensu stricto)   ...... 8  

7 (6). Dorsal surface with shallow microlines, mesh pattern isodiametric. Specimen from locality in northern Colombia........................... C. fernandezi   , n. sp., p. 56

– Dorsal surface without microlines, smooth. Specimen from locality in northern Venezuela....................................................................... C. negrei Perrault   , p. 58

8 (6). Body size large (SBL more than 5.0 mm, EW more than 1.5 mm). Elytron bicolored (Figs. 17A–C), pale anteriorly, dark posteriorly. Femora and tibiae piceous to rufous ...................................................................... basale   species group......9

– Body size smaller (SBL less than 4.5 mm, EW less than 1.5 mm). Elytron concolorous or bicolored. Femora and tibiae rufous to testaceous ......................... 10

9 (8). Dorsal vestiture ( Fig. 2D View FIGURE 2 ) of elytron relatively long setae. Specimen from locality in Palaearctic southwestern Asia or in the Afrotropical Region west to Niger ( Fig. 16 View FIGURE 16 ) .............................................................. C. semirubricum Reitter   , p. 61

– Dorsal vestiture of elytron relatively shorter setae. Specimen from locality in northwestern Afrotropical Region ( Mali and Sénégal, Fig. 16 View FIGURE 16 )............................ ......................................................................................... C. basale Dejean   , p. 65

10 (8). Elytron with punctation in straight lines, stria­like. Dorsal integument uniformly rufotestaceous. Clypeus (cf. Fig. 11B View FIGURE 11 ) without dentiform projection .................. ...................................................................................... C. namwala   , n. sp., p. 99

– Elytron with punctation scattered over surface (if lines of punctation apparent, punctures also in between). Dorsal integument various in color. Clypeus with ( Fig. 11C View FIGURE 11 ) or without ( Fig. 11B View FIGURE 11 ) dentiform projection ..................................... 11

11 (10). Eyes somewhat flattened, only slightly projected laterad dorsolateral carina of head ( Fig. 11B View FIGURE 11 ). Values for VW/HW 0.87 or more. Dorsal integument concolorous, uniformly pale .......................................................................................... 12

– Eyes rounded, moderately projected laterad dorsolateral carina of head ( Fig. 11C View FIGURE 11 ). Values for VW/HW 0.87 or less. Dorsal integument various: distinctly or indistinctly bicolored, or uniformly pale........................................................... 15

12 (11). Clypeus anteriomedially with dentiform projection ( Fig. 11C View FIGURE 11 , ct) ................. 13

– Clypeus anteriomedially without dentiform projection ( Fig. 11B View FIGURE 11 ) ................. 14

13 (12). Scutellum with microsculpture mesh pattern isodiametric, sculpticells convex .. ............................................................................. C. candidum Andrewes   , p. 100

– Scutellum smooth, without microlines .................. C. subcaecum   , n. sp., p. 101

14 (12). Scutellum with slightly transverse mesh pattern over much of surface, smooth apically. Specimen from locality in southwestern Asia, west to eastern Palaearctic Africa (Fig. 23) ........................................ C. transcaspicum Semenov   p. 102

– Scutellum with slightly transverse mesh pattern near anterior margin, smooth apically. Specimen from locality in Afrotropical Region, east to Anatolian peninsula in Palaearctic Region (Fig. 23) ...... C. microphthalmum Chaudoir   , p. 104

15 (11). Specimen from locality in South Africa or Botswana (Fig. 20). Male genitalia as in Fig. 19D.............................................................. C. capicola Péringuey   , p. 83

– Specimen from locality farther north in Africa, and eastward in Palaearctic southwestern Asia, to Vietnam, in the Oriental Region. Male genitalia otherwise (Figs. 19C, 19G–H, 19K, 19O and 22D)........................................................... 16

16 (15). Head with postocular transverse impression with microsculpture mesh pattern ( Figs. 11B–C View FIGURE 11 ) isodiametric to slightly transverse ............................................ 17

– Head with postocular transverse impression smooth, without microlines, or microlines not joined in form of a network...................................................... 20

17 (16). Specimen from locality in Palaearctic southwestern Asia, or in eastern Afrotropical Region, north of South Africa (Fig. 21); elytron bicolored (Figs. 19L–N; cf. Fig. 19F) ................................................................... C. pictulum Bates   , p. 93

– Specimen from locality in Oriental Region (Fig. 21) ...................................... 18

18 (17). Elytron concolorous, rufous, or bicolored, with piceous or black mark small, confined to disc (Figs. 19E–F). Specimen from locality in Burma, or farther east (Fig. 21) .................... .................................................. C. helferi Chaudoir   , p. 88

– Elytron bicolored, dark marks more extensive, extended to lateral margin. Specimen from locality in India, Sri Lanka, or Nepal............................................ 19

19 (18). Phallus in left lateral aspect gradually curved preapically (Fig. 19K). Specimen from locality in southern India (Calcutta area southward) or Sri Lanka.............. ................................................................................... C. fluviale Andrewes   , p. 92

– Phallus in left lateral aspect markedly curved preapically (Fig. 19H). Specimen from locality in northern India or Nepal................... C. fasciger Chaudoir   , p. 91

20 (16). Elytron concolorous, pale ................................................................................ 21

– Elytron bicolored (Figs. 19A–B, 22A–C) ........................................................ 24

21 (20). Scutellum entirely smooth or surface wrinkled basally, without microlines ........ ........................................................................ C. rufotestaceum Fairmaire   , p. 81

– Scutellum basally with microlines in form of mesh pattern ............................ 22

22 (21). Dissected specimen female (one of two species) ................................................. ..................... C. schueppelii Dejean   (in part), p. 78, C. fasciatum   (in part), p. 95

– Dissected specimen male ................................................................................. 23

23 (22). Male genitalia (Fig. 19C) with microtrichial field (mf) of endophallus extensive; phallus with preapical portion shorter, curved ventrad rather sharply ................. .................................................................... C. schueppelii Dejean   (in part), p. 78

– Male genitalia (Fig. 22D) with microtrichial field (mf) of endophallus very small; phallus with preapical portion longer, curved ventrad more gradually ..... ...................................................................... C. fasciatum Dejean   (in part), p. 98

24 (20). Elytron bicolored, with dark marking (Figs. 19A–B) extended to entire posterior margin ...................................................................... C. schueppelii Dejean   , p. 78

– Elytron bicolored, fasciate (Figs. 22A–C), fascia narrow or broad, but complete posterior margin not dark, at most bicolored............... C. fasciatum Dejean   , p. 95