Morophagoides Petersen, 1957

Osada, Yohei, Sakai, Makoto & Hirowatari, Toshiya, 2015, A revision of the genus Morophagoides Petersen (Lepidoptera, Tineidae) from Japan, Zootaxa 3973 (2), pp. 351-368 : 352

Genus Morophagoides Petersen, 1957

Morophagoides Petersen, 1957: 593 . Type species: Scardia ussuriensis Caradja, 1920 .

Description. Head: Vertex and face roughly clothed with yellowish white hairs. Antenna filiform and strongly ciliated; pecten consisting of 5-14 bristles; flagellomere clothed with a row of gray scales. Maxillary palpus clothed with gray scales. Labial palpus forwardly directed or upturned, covered with yellowish white scales mesally and gray scales laterally, 2nd segment with 6-7 lateral bristles. Thorax: Mesonotum grayish cream, but anterior margin and posterior end clothed with black scales; tegula clothed with black scales anteriorly and grayish cream scales posteriorly; metanotum clothed with grayish cream scales on scutellum and posterior part of scutum. Legs extensively covered with black scales, apical portion of tibia and each tarsomere grayish cream; hind tibia bearing dorsally and ventrally dense long hairs. Abdomen: Extensively clothed with pale gray to grayish cream scales. Eighth abdominal segment without coremata.

Wing markings: Forewing upperside grayish cream in ground color; distinct dark maculae on subbasal portion and middle of costa and middle of posterior margin; some small dark speckles arranged along costa and termen; irregularly darkened areas around discoidal cell. Fringe with yellowish white and brown scales alternately. Hindwing upperside glassy grayish white, end of free veins indistinctly darkened. Fringe with yellowish white scales posteriorly and yellowish white and brown scales alternately.

Wing venation: Forewing all free veins present; R1 arising from 1/3 of discoidal cell; R5 ending posterior to apex of wing; 1A+2A twice as long as 1A; apex of discoidal cell rounded. Hindwing all free veins present; basal 2/ 5 of costa weakly arched; M branched in discoidal cell; distance between base of CuA1 and CuA2 1.1 times as long as medial cell.

Male genitalia: Tegumen and vinculum fused into a ring; tegumen and uncus mesally fused with each other; gnathos with a pair of thin elongate plates; vinculum and saccus broad, the latter gradually tapered to pointed apex. Valva with a setose conical dorsoventrally-directed medial process; posterior part of valva with setae; valval apodeme well developed. Phallus almost straight; suprazonal sheath about 1/4 as long as phallus; subzonal sheath about 3/4 as long as phallus; vesica with patch of large spicular microtrichia, with digitiform subapical carina. Juxta absent.

Female genitalia: Eighth abdominal tergum developed with setae; eighth sternum broad basally, evenly tapered posteriorly, swollen mesally, and with a pair of setose lobes posteriorly. Antrum well developed and strongly sclerotized. Ductus bursae ornamented with microtrichia. Corpus bursae ovate, with a pair of small pouch-shaped signa invaginated into wall.

Remarks. The genus Morophagoides is distributed in the Palaearctic, Oriental ( Taiwan), Nearctic and Neotropical regions, and as described above some species are well known as pests of the shiitake mushroom, Lentinus edodes . The four new species are placed in this genus because the genital structure of these species is similar to the other Japanese species of the genus.