Orictites plurisetosus, Balkenohl, 2017

Balkenohl, Michael, 2017, Revision of the genus Orictites ANDREWES, 1931 (Coleoptera, Carabidae, Clivinini), Contributions to Natural History 35, pp. 1-68 : 32-34

Orictites plurisetosus sp. nov.

Type material: Holotype: ♂, labels: white, black printed: “ PHILIPPINEN – Mindoro 28km S Calapan Balete 27.–29.11.1992 leg. Schillhammer (13)” ( NHMW).

Description ( Figs 5 , 23 ; pp. 9, 24)

Measurements: Body length 3.84 mm, width 1.02 mm; ratio length/width of pronotum 1.06; ratio length/width of elytra 2.06.

Colour: Shiny. Brown. Pronotum, all reflexed margins, lines, striae and punctures darker. Labrum and palpi yellowish. Antennae and legs brown.

Head: Slightly narrower than pronotum. Clypeus with small rounded tooth laterally, anterior margin slightly excised, with blunt raised elevation between the two clypeal setae, with punctures reaching up to frons, margin of wings convex, flattened, divided from middle part of clypeus by slight notches and carinae, divided from supraantennal plates by distinct obtusely angled notches and ridge, clypeus and wings reflexed margined, supraantennal plates vaulted up to posterior eye level, minutely punctured (100x), with reflexed rounded margin, elongated as supraorbital carina, frons separated from clypeus by indistinct transverse depression, with costate nearly parallel running frontal carinae at each side posteriorly, with broad furrow between frontal carina and supraorbital carina, sharp furrow between frons and frontal carina, frons moderately convex, punctured. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle. Eyes and genae prominent, nearly as wide as anterior angles of pronotum, moderately convex, genae enlarged, rounded, not as high and shorter than eyes, enclosing eyes posteriorly, forming obtuse angle at neck. Grooves for reception of antennae short, length 0.4 of eye diameter. Neck constriction marked as line of punctures, neck laterally covered with reticulation and punctures. Labrum straight, 7-setose, ciliate laterally, with isodiametric reticulation. Mandibles robust, shorter than head, flattened, stout, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles obtusely angled towards base (opened position), both mandibles somewhat hollowed out dorsally. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2 nd segment, 2 nd segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck with medium sized punctures and some reticulation, Submentum and mentum distinctly separated; submentum with four bigger, about equally spaced setigerous punctures; mentum at base bilaterally with rounded elevation, with some longitudinal carinae, with fine reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming obtuse triangle, bisetose, not projecting as far as lateral lobes. Antennae of moderate length, reaching up to posterior third of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with fine reticulation, segments 5–10 moniliform (L/ W 1.0), antennomeres cylindrical, with shiny areas, segments 4–11 densely pubescent, with a few additional long setae.

Pronotum: Square, longer than wide, sides straight, rounded at anterior angles. Anterior margin straight. Reflexed lateral margin smooth, slightly convex for a short distance before posterior angles. Margin from posterior angle to base running as slightly sigmoid line. Lateral channel distinctly broadened between the two lateral setigerous punctures, smooth. Anterior setigerous puncture located at the end of anterior quarter adjoining convexity of disc of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angles indistinctly projecting, rounded, formed by reflexed lateral margin, posterior angle developed as sharp distinct tooth, projecting antero-laterally. Anterior transverse line sharp, deep. Median line deep and moderately broad, surpassing anterior transverse line without joining, fine at base, not joining base. Surface irregularly covered with big punctures, with distinct punctured basal impression. Basal channel broad, deep. Flange raised keel-like (lateral view).

Elytra: Subcylindrical, sides indistinctly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles, with additional row of fine punctures laterally. Reflexed margin with one or two scars posterior humerus, margin slightly thickened at humerus, bending with channel over rounded humerus up to 5 th stria. No tooth at humerus. Basal tubercle distinct, with setigerous puncture, situated at declivity of first stria. Striae 1–4 free at base, all striae deep, distinctly punctate, striae 1 and 2 ending free at apex, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura raised. All intervals distinctly convex, 3 rd and 4 th broader and distinctly raised at base, both with tubercle at base, 7 th and 8 th carinate. All intervals shiny on disc. Interval 3 with 16/18, 5 with 19, and 7 with 15 setigerous punctures, somewhat irregularly placed on the respective interval.

Hind wings: Fully developed.

Ventral surface: Epipleuron shiny, with punctures and indistinct reticulation, submarginal furrow distinct. Proepisternum and episternum nearly smooth, prosternite smooth, keeled at middle, mesosternum smooth. Sternites with transverse to isodiametric reticulation, each with big punctures at base, slightly denser laterally, 3 rd to 5 th with paralateral ambulatory setae at each side, ventral strigae distinct, sternite 6 smooth at middle, with two widely separated apical setae at each side.

Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, no dorsal furrow, smooth, basal tarsomere elongated, as long as the following four together. Mesotibia with preapical spine, furnished with strong seta. All tarsomeres relatively broad.

Male genitalia ( Fig. 23 ): Moderately long, moderately sclerotized. Median lobe slender, moderately arcuate, slightly twisted basally, apex stick-like. Endophallus basally strongly folded, with some bristles towards apex. Parameres of about same length, dorsal one broader, slightly distorted, asetose.

Female genitalia: Unknown.

Variation: The number of setigerous punctures on the 3 rd interval of the elytra seems to vary intraindividually.

Diagnosis: A small medium-brown species with the knob on the clypeus developed as elevation and indistinct transverse depression behind, no humeral tooth, and a tubercle at the base of interval 3 and 4 of the elytra. Distinguished from all other species by the series of setigerous punctures on intervals 3, 5, and 7, and the eyes and genae which are projecting laterally.

Distribution: Known from Mindoro ( Philippines) only.

Etymology: The name refers to the numerous setae on the intervals 3, 5 and 7 of the elytra.

Figures

Figs 5–8: Orictites spp., habitus. 5: O. plurisetosus sp. nov., holotype, male; 6: O. anteriortenuis sp. nov., holotype, male; 7: O. omnipunctatus sp. nov., holotype, male; 8: O. ampliosetosus sp. nov., holotype, male.

Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm.