Orictites desuntsetosus, Balkenohl, 2017

Balkenohl, Michael, 2017, Revision of the genus Orictites ANDREWES, 1931 (Coleoptera, Carabidae, Clivinini), Contributions to Natural History 35, pp. 1-68 : 29-32

Orictites desuntsetosus sp. nov.

Type material: Holotype: ♂, Labels: white, black printed: “leg. Jäch (11) PHILIPPINEN – Luzon 30km E Lucena City Quezon NP 23.11.1992 ” ( NHMW) . Paratypes: 3 ♀, same data as holotype ( NHMW / CBB) .

Description ( Figs 4 , 22 , 34 ; pp. 8, 24, 35)

Measurements (n = 4): Body length 4.38–4.65 mm ( = 4.49 mm), width 1.25– 1.31 mm ( = 1.29 mm), ratio length/width of pronotum 0.98–0.99 ( = 0.98), ratio length/width of elytra 2.25–2.31 ( = 2.29).

Colour: Shiny. Brown. Reflexed margins, lines, striae, and punctures darker. Labrum and palpi yellowish. Antennae and legs brown.

Head: A quarter narrower than pronotum. Clypeus deeply excised, with small rounded tooth laterally nearly fused with clypeal wings, with blunt raised elevation between the two clypeal setae, anteriorly smooth, posterior part of elevation covered with big punctures, margin of wings convex, flattened, divided from middle part of clypeus by slight carinae, divided from supraantennal plates by distinct obtusely angled notches and ridge, clypeus and wings reflexed margined, supraantennal plates vaulted up to posterior eye level, smooth, with reflexed rounded margin, as supraorbital carina, frons separated from clypeus by flat transverse depression with punctures, with costate posteriorly diverging frontal carinae at each side, with broad furrow between frontal carina and supraorbital carina, sharp furrow between frons and frontal carina, frons moderately convex, smooth. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle. Eyes relatively big, convex, genae distinct, enclosing eyes posteriorly, forming obtuse angle at neck. Grooves for reception of antennae short, length 0.3 of eye diameter. Neck constriction marked by punctures with wider distance, neck laterally covered with reticulation and densely scattered set medium sized punctures. Labrum straight, 7-setose, ciliate laterally, with isodiametric reticulation. Mandibles robust, shorter than head, flattened, stout, broadened at base, acutely curved at apex, carinae of right scrobe interrupted at middle, both mandibles obtusely angled towards base (opened position), both mandibles somewhat hollowed out dorsally. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2 nd segment, 2 nd segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck with punctures. Submentum and mentum distinctly separated; submentum with four bigger, about equally spaced setigerous punctures; mentum at base bilaterally with rounded elevation, with some longitudinal carinae, with fine reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming obtuse triangle, bisetose, not projecting as far as lateral lobes. Antennae relatively long, reaching up to posterior setigerous puncture of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with fine reticulation, segments 5–10 submoniliform (L/ W 0.89), antennomeres somewhat flattened, with shiny areas, segments 4–11 densely pubescent, with a few additional long setae.

Pronotum: Square, as long as wide, sides straight, slightly diverging posteriorly. Anterior margin straight. Reflexed lateral margin smooth. Margin from posterior angle to base running as sigmoid line. Lateral channel conspicuously broad between the two lateral setigerous punctures, nearly smooth. Anterior setigerous puncture located at the end of anterior quarter approximated to the convexity of the disc of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angle indistinctly projecting, angled, formed by reflexed lateral margin, posterior angle developed as distinct tooth, projecting antero-laterally. Anterior transverse line formed by some distant punctures. Median line deep, broad, surpassing anterior transverse line without joining, not joining base. Surface with some irregularly situated medium sized punctures, with basal impression. Basal channel broad, deep. Flange raised keel-like (lateral view).

Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles, with additional row of small punctures laterally. Reflexed margin with two indistinct scars (120x) in anterior third, bending with channel over rounded humerus up to 5 th stria. No tooth at humerus. Basal tubercle distinct, with setigerous puncture, situated at declivity of first stria. Striae 1–4 free at base, all striae deep, indistinctly punctate, striae 1 and 2 ending free at apex, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura raised. All intervals distinctly convex, 3 rd broader and distinctly raised at base, with tubercle at base, 8 th carinate. All intervals shiny on disc. Interval 3 or any other interval asetose.

Hind wings: Fully developed.

Ventral surface: Epipleuron shiny, with scattered punctures and indistinct reticulation, submarginal furrow distinct. Proepisternum and episternum nearly smooth, prosternite smooth, keeled at middle. Mesosternum smooth. Sternites with transverse to isodiametric reticulation, each with big punctures at base, denser laterally, 3 rd to 5 th with paralateral ambulatory setae at each side, ventral strigae distinct, sternite 6 smooth at middle, with two widely separated apical setae at each side.

Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, no dorsal furrow, smooth, basal tarsomere elongated, as long as the following four together. Mesotibia with preapical spine, furnished with strong seta. All tarsomeres relatively broad.

Male genitalia ( Fig. 22 ): Moderately sclerotized. Median lobe slender, moderately arcuate, not twisted, apex stick-like. Endophallus without visible teeth. Parameres of about same length, both slender, each distorted anterior apex, with minute seta preapically.

Female genitalia. ( Fig. 34 ): Coxostylus slender, nearly straight in basal part, gently curved to apex, with short obtuse knob-like tooth dorsally at base, with one seta preapically, with one blunt and one acute short ensiform seta, with five nematiform setae. Ramus of moderate size.

Variation: In one of the paratypes there are single punctures visible.

Diagnosis: A small medium-brown species with the knob on the clypeus developed as elevation and indistinct transverse depression behind, no humeral tooth, and a tubercle only at the base of interval 3 of the elytra. Distinguished from all other species by the completely missing setigerous punctures on intervals of the elytra and the deeply excised clypeus.

Distribution: Known from the Luzon Island ( Philippines).

Etymology: The name refers to the missing setae on the elytra.

Figures

Figs 1–4: Orictites spp., habitus. 1: O. minotaur ANDREWES, 1931, lectotype, male; 2: O. costulipennis BATES, 1892, holotype, male; 3: O. mjoebergi LouWERENS, 1964, lectotype, male; 4: O. desuntsetosus sp. nov., holotype, male.

Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm.

Figs 31–34: Orictites spp., coxostylus. 31: O. minotaur ANDREWES, 1931, scale bar 0.5 mm; 32: O. costulipennis BATES, 1892, scale bar 0.1 mm. 33: O. mjoebergi LouWERENS, 1964, scale bar 0.1 mm; 34: O. desuntsetosus sp. nov., scale bar 0.1 mm.