Defensina bucciniformis ( Hoernes & Auinger, 1880 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard M., 2021, The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea — striking diversity of a negelected group, Zootaxa 5025 (1), pp. 1-75 : 42-43

Defensina bucciniformis ( Hoernes & Auinger, 1880) nov. comb.

Figs 4I, P, 12D 1 –D 2 , E 1 –E 2 , F, G

[ Mitrella ] bucciniformis nov. form.—Hoernes 1880: 127 [nomen nudum].

* Columbella (Mitrella) bucciniformis nov. form.— Hoernes & Auinger 1880: 98, pl. 12, figs 3a–c.

Columbella (Mitrella) bucciniformis Hö. Au. — Boettger 1902: 17.

Columbella (Mitrella) inquidens n. sp. — Boettger 1906: 17.

Pyrene (Mitrella) inquidens (Boettger) — Zilch 1934: 203, pl. 16, fig. 89.

Mitrella ( Atilia ?) bucciniformis Hoernes und Auinger 1880 —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 184, pl. 45, fig. 9.

Mitrella (Mitrella) inquidens ( Boettger, 1906) — Atanacković 1985; 142, pl. 32, figs 1–2.

Type material. Lectotype (designated herein): NHMW1876 /0011/0018, SL: 8.7 mm, MD: 3.6 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 12, figs 3a–c), Figs 12D 1 –D 2 . Paralectotypes: NHMW1859 /0045/0601,SL: 8.8mm,MD: 3.5mm, LăpugiudeSus ( Romania) , Figs12E 1 –E 2 . NHMW1854 /0035/0133, SL: 8.7 mm, MD: 3.3 mm, Lăpugiu de Sus ( Romania) , Figs 4P, 12F . NHMW1855 /0043/0009, 17 spec., Lăpugiu de Sus ( Romania) . NHMW 2020 /0172/0001 (ex coll. B. Landau), 9 spec., Lăpugiu de Sus ( Romania) .

Additional material. SMF 360679 (= XII.2223 a), Senckenberg Museum, Frankfurt/Main, Germany, SL: 5.8 mm, MD: 2.8 mm, Coşteiu de Sus ( Romania), lectotype of Mitrella inquidens Boettger, 1906 , illustrated in Zilch (1934, pl. 16, fig. 89), Figs 4I, 12G .

Shell measurements and ratios. SL: 8.7–8.8 mm, MD: 3.3–3.6 mm, AA: 34–38°, SL/MD: 2.4–2.6, AL/AW: 2.3–2.5, LWH/AH: 1.4.

Description and Discussion. See under generic note. Columbella (Mitrella) inquidens Boettger, 1906 is a juvenile specimen, which lacks the characteristic apertural features of the adult specimens.

Palaeoenvironment. Unknown, probably middle to outer neritic environments.

Distribution in Central Paratethys. Badenian (middle Miocene): southern Pannonian Basin: Hrvaćani-Drenik ( Bosnia and Hercegovina) ( Atanacković 1985); Dacian Basin: Târnene, Staropatica ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960); Făget Basin: Lăpugiu de Sus, Coşteiu de Sus ( Romania) ( Boettger 1906). Only known from southern Paratethyan basins.

Genus Thiarella Sacco, 1890

Type species. Murex thiara Brocchi 1814 ; subsequent designation by Cossmann (1899: 46). Pliocene, Italy.

Discussion. Thiarella is characterised by a medium-sized, moderately broad to slender fusiform shell with high conical spire and long siphonal canal, flat-sided whorls with narrowly-incised suture and sculpture of widely-spaced axial ribs, forming a beaded subsutural swelling below adapical suture, giving the whorls a coronate aspect.

Sacco (1890b) included Murex thiara Brocchi, 1814 , from the Tortonian of Stazzano and San Agata ( Italy) and the Pliocene of Asti and Vaile ( Italy), Columbella (Thiarella) rosavandae Sacco, 1890 , from the Messinian of Tetti Borelli ( Italy), and Columbella (Thiarella) carinata Sacco, 1890 non Hinds, 1844, from the Tortonian of Stazzano and San Agata ( Italy) and the Messinian of Tetti Borelli ( Italy), in his new genus. These taxa co-occur at the same localities and the differences seen in the syntypes seem to be mainly a matter of ontogeny (see Ferrero Mortara et al. 1981, pl. 58, figs 1–2). We consider C. rosavandae and C. carinata to be subjective junior synonyms of Thiarella thiara . Therefore, we do not propose a replacement name for the preoccupied Columbella carinata Sacco, 1890 . Thiarella is represented only by Thiarella thiara and Thiara bronni , forming a continuous linage from the Langhian to the Pliocene.

Cossmann (1899: 46) introduced the replacement name Crenisutura, arguing that Thiarella was preoccupied by a mitrid by Swainson (1840). Swainson (1840: 131), however, spelled his genus Tiarella , whereas Sacco (1890b: 52) introduced his new genus as Thiarella . According to ICZN article 56.2 two names with a one-letter difference are not homonyms. The European Neogene Thiarella species are superficially similar with the extant Strombina angularis (G.B. Sowerby I, 1832) from Panama and Mexico in general shape and sculpture. However, that species does not have coronate whorls and the apertural dentition is quite different, with a strongly developed internal labial callus pad extending along the entire mid-portion of the outer lip, narrowing the aperture, and bearing coarse denticles; typical of the ‘ Strombina group’.

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width.

FIGURE 12 A1–A2. “Mitrella” bittneri (Hoernes &Auinger, 1880), NHMW1860/0001/0129a, lectotype, Steinebrunn (Austria). B1–B2. “Mitrella” bittneri (Hoernes & Auinger, 1880), NHMW1860/0001/0129b, paralectotype, Steinebrunn (Austria). C1– C2. “Mitrella” bittneri (Hoernes & Auinger, 1880), NHMW1860/0001/0129c, paralectotype, Steinebrunn (Austria). D1–D2. Defensina bucciniformis (Hoernes & Auinger, 1880), NHMW1876/0011/0018, lectotype, Lăpugiu de Sus (Romania). E1–E2. Defensina bucciniformis (Hoernes & Auinger, 1880), NHMW1859/0045/0601, paralectotype, Lăpugiu de Sus (Romania). F. Defensina bucciniformis (Hoernes & Auinger, 1880), NHMW1854/0035/0133, paralectotype, Lăpugiu de Sus (Romania). G. Defensina bucciniformis (Hoernes & Auinger, 1880), SMF 360679, lectotype of Mitrella inquidens Boettger, 1906, Coşteiu de Sus (Romania).