Ptomaphaminus marshalli Schilthuizen & Perreau

Schilthuizen, Menno, Perreau, Michel & Njunjic, Iva, 2018, A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae), ZooKeys 777, pp. 57-108 : 87

Ptomaphaminus marshalli Schilthuizen & Perreau sp. n. Figures 12o, p, 13x, y, 14a, b

Material.

Holotype: Malaysia, Sarawak, 4th Division, Gunung Mulu National Park, Slope, 29.4.78, Tps 4-6, v–viii.1978 (leg. P.M. Hammond & J.E. Marshall, in NHMUK, B.M.1978-49), 1 male. Paratypes: Sarawak. Gunung Mulu National Park, Slope, 29.4.78, Tps 4-6, v–viii.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M.1978-49), 2 males, 1 female; Gunung Mulu National Park, Slope, 29.4.78, Tps 1-3, v–viii.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M.1978-49), 6 males, 7 females; Gunung Mulu National Park, Slope, 7.4.78, baited traps, v–viii.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M.1978-49), 1 female. Sabah. Crocker Range Park, Inobong Station, 5°51.265'N, 116°06.363'E, 500 m elev., 21-23.ix.2012 (leg. M. Schilthuizen, Crocker Range / Kinabalu Expedition, RMNH.INS.555639), 1 male.

Description.

Length: 1.50-1.90 mm. General colour brown; the tarsi, the two first antennomeres, the base of the third antennomere, and of the tibiae yellowish. Winged. Eyes well developed. Pronotum 1.65 times as wide as long. Elytra 1.25 times as long as wide. Female sutural apex of elytra rounded (Figure 14b), similar to the male (Figure 14a). Spiculum gastrale of the male genital segment apically dilated into a short discoid expansion (Figure 13x). Aedeagus approximately 5.7 times shorter than the body length, the sides regularly arcuate in dorsal view (Figure 12o), moderately arcuate ventrally and obtusely angular dorsally in lateral view (Figure 12p). Apex of the median lobe with a large ventrally deflexed but not retroverted hook. Four lateroventral setae spaced from the apical third of the aedeagus and the apex. Internal stylus of the endophallus moderately sinuate. Female genital segment with reduced gonocoxites. Spermiduct not helical. Apex of the spermatheca conically narrowed, the apex shortly rounded, without apical sclerotised plate (Figure 13y).

Differential diagnosis.

Females of this species can be recognized by the apex of the spermatheca, which is conically narrowed, a very distinct morphology compared to all other species, which have a spermatheca with a widely rounded apex.

Habitat and distribution.

Known from lowland forests of Gunung Mulu, Sarawak, and of Gunung Kinabalu, Sabah, Malaysia.

Etymology.

Dedicated to J. E. Marshall, one of the collectors of the species during the expedition of the Natural History Museum of London in Sarawak.