Phlegmariurus wilsonii (Underwood & F.E Lloyd) B. Øllgaard (2012a: 481)
Øllgaard, Benjamin & Testo, Weston, 2021, The Lycopodiaceae of Panamá, Phytotaxa 526 (1), pp. 1-66 : 45-46
publication ID |
https://doi.org/ 10.11646/phytotaxa.526.1.1 |
DOI |
https://doi.org/10.5281/zenodo.5815569 |
persistent identifier |
https://treatment.plazi.org/id/286E8977-7B62-FD76-1085-FD84CE9C4668 |
treatment provided by |
Plazi |
scientific name |
Phlegmariurus wilsonii (Underwood & F.E Lloyd) B. Øllgaard (2012a: 481) |
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. Phlegmariurus wilsonii (Underwood & F.E Lloyd) B. Øllgaard (2012a: 481) View in CoL View at ENA . Fig. 20
Lycopodium wilsonii Underwood & F.E Lloyd (1906: 111) View in CoL .— Urostachys wilsonii (Underwood & F.Lloyd) Herter (1923: 163) View in CoL .— Huperzia wilsonii (Underwood & F.E Lloyd) B.Øllgaard (1987: 170) View in CoL . Type:— PUERTO RICO: Luquillo Mountains, Wilson 153 (NY lectotype, here designated). Note: According to the protolog the type is Wilson 271, but Wilson 153 (NY) is annotated by the author as sp. nov. and fits the description, while Wilson 271 (NY) is Phlegmariurus dichotomus View in CoL and is not annotated as sp. nov.
Lycopodium andinum Herter (1909a: 49) View in CoL non Rosenstock (1908: 239).— Lycopodium lindavianum Herter (1909b: 90) View in CoL .— Urostachys lindavianus (Herter) Nessel (1939: 146) View in CoL .— Huperzia lindaviana (Herter) Holub (1985: 74) View in CoL . Type:— ECUADOR: “In territorio rei publ. Ecuador”, Fraser s.n. (P [Herb. Drake] lectotype, designated by Øllgaard 1988: 97; isotype, G).
Lycopodium trichodendron Herter (1909a: 49) View in CoL .— Urostachys trichodendron (Herter) Herter (1923: 163) View in CoL . Type:— GUADELOUPE: L’Herminier s. n. (P00559253), Bory 103 (P00559254, syntypes).
Lycopodium stamineum Maxon (1912: 2) View in CoL .— Urostachys stamineus (Maxon) Nessel (1939: 148) View in CoL . Type:— PANAMÁ. Chiriquí: Upper Caldera watershed, betrween Camp I and the Divide, Holcombs Trail, above El Boquete, ca. 1750 m, Maxon 5636 ( US 676085 holotype).
Lycopodium arcanum Maxon (1938: 310) View in CoL . Type:— HONDURAS: Comayagua, above El Achiote, above plains of Siguatepeque, 1800 m, Yuncker, Dawson & Youse 6149 ( US holotype).
Urostachys mandiocanus (Raddi) Herter var. filifolius Herter (1954: 124) View in CoL . Type:— ECUADOR: Eastern slope of eastern cordillera, bank of Palora river, 1500 m, Rimbach 125 ( US holotype).
Plants erect, arcuate-spreading to pendulous, to 20 (–30) cm long. Shoots homophyllous, 1.5–2.5 (–3) cm in diameter including leaves, equally thick throughout, or in some slender individuals gradually tapering to 1–1.5 cm in terminal divisions. Stems excluding leaves (1–) 1.5–2 (–3) mm thick at the base, often tapering to (0.7–) 1 mm, prominently ridged by decurrent leaf bases, pale green to stramineous, often with bright red spots on leaf bases, sporangiate, often in seasonally produced zones, from 5–15 cm above the base and upward, 3–6 (–9) times dichotomous. Leaves usually uniform throughout, borne in alternating, often irregularly oblique whorls of 6–7, these 0.5–1.5 mm apart in basal divisions, upward often in whorls of 4–5, forming 8–14 indistinct longitudinal ranks, perpendicularly spreading to ascending, straight to upward curved, usually not twisted at the base, linear to filiform, (6–) 10–17 × 0.3–0.5 mm, quickly narrowed to ca. 0.2 mm wide due to involution, gradually tapering toward the tip, adaxially canaliculate to involute, often with a prominent vein abaxially near the base. Leaves of terminal divisions in old plants sometimes gradually reduced to 6(–4) mm long. Decurrent leaf bases usually not wider than the lamina base, often bright red. Sporangia 1–1.5 mm wide.
Distribution:— Mexico (Oaxaca) to Panamá, West Indies, northern South America south to Bolivia, Brazil (Mato Grosso).
Habitats: —Epiphytic (rarely terrestrial) in montane forest, elev. 900–1900 m elev. Usually an erect epiphyte, but sometimes becoming pendulous when very large; often with bright red, but usually not prominent leaf bases.
Notes:— Phlegmariurus wilsonii is related to P. dichotomus , P.mandiocanus ( Raddi 1819: 280) B. Øllgaard (2012a: 480) from Brazil, and P. pithyoides (Schld. & Cham.) B.Øllgaard from Central America, Hispaniola, Colombia, and Venezuela. With these it shares the bottle-brush-like aspect, and with the latter two the bright red coloration of the leaf bases. Pendulous individuals of P. wilsonii are rather similar to P. polycarpos , but the latter has shorter, more flattened, leaves, and uniformly falcate leaves due to a basal twist, and this species apparently is always pendulous. Pendulous individuals of P. wilsonii may be separated from P. watsonianus by the lack of auricles at the lamina base.
The absence or presence of red colour on the leaf bases seems uncorrelated with other characters. The intensity and size of the coloration is also variable. The direction of leaves is correlated to growth habit. In erect-growing plants the leaves are wide-spreading to slightly reflexed, while in pendent plants the leaves are somewhat ascending.
Specimens studied:—Chiriquí: Bajo Chorro, Boquete, 1830 m, Davidson 230 (F, MO, US). Palo Alto, 4.5 mi NE of Boquete, forest along W branch of headwaters of Río Palo Alto, 1920 m, Hammel 7304 (MO). Above El Boquete, 1750 m, Maxon 5636 ( US). Along trail between north fork of Río Palo Alto and Cerro Pate Macho, ca. 6 km NE of Boquete, 1600–2000 m, Smith 2414 (UC). Camino campamento de Fortuna sitio de presa—Trocha 3 de noviembre, entre Santa María y Pinola, 1000–1200 m, Correa et al. 2926 (PMA). Parque Nacional Volcán Barú, Alto Chiquero, 893 m, Mello-Silva et al. 3350 (PMA).
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Naturhistorisches Museum Wien |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Genus |
Phlegmariurus wilsonii (Underwood & F.E Lloyd) B. Øllgaard (2012a: 481)
Øllgaard, Benjamin & Testo, Weston 2021 |
Phlegmariurus dichotomus
W. H. Wagner 1993 |
Huperzia wilsonii (Underwood & F.E Lloyd) B.Øllgaard (1987: 170)
B. Ollgaard 1987: 170 |
Huperzia lindaviana (Herter)
Holub 1985: 74 |
Urostachys mandiocanus (Raddi)
Herter 1954: 124 |
Urostachys lindavianus (Herter)
Nessel 1939: 146 |
Urostachys stamineus (Maxon)
Nessel 1939: 148 |
Lycopodium arcanum
Maxon 1938: 310 |
Urostachys wilsonii (Underwood & F.Lloyd)
Herter 1923: 163 |
Urostachys trichodendron
Herter 1923: 163 |
Lycopodium stamineum
Maxon 1912: 2 |
Lycopodium andinum
Herter 1909: 49 |
Lycopodium lindavianum
Herter 1909: 90 |
Lycopodium trichodendron
Herter 1909: 49 |
Lycopodium wilsonii Underwood & F.E Lloyd (1906: 111)
Underwood & F. E Lloyd 1906: 111 |