Phlegmariurus tubulosus ­­­ (Maxon) B. Øllgaard (2012b: 19)

Øllgaard, Benjamin & Testo, Weston, 2021, The Lycopodiaceae of Panamá, Phytotaxa 526 (1), pp. 1-66 : 42

2.­­­ Phlegmariurus tubulosus ­­­ (Maxon) B. Øllgaard (2012b: 19) . Fig. 18B

Lycopodium tubulosum Maxon (1913: 178) .— Urostachys tubulosus (Maxon) Nessel (1939: 177) .— Huperzia tubulosa (Maxon) B. Øllgaard (1992: 697) . Type:— COSTA RICA: Pacayas, at the foot of the volcano Turrialba, 1400 m, Dec. 1908, Biolley 17398, US holotype).

Urostachys costaricensis Herter, (1958: 83) .— Lycopodium costaricense (Herter) Morton (1964: 72) . Type:— COSTA RICA: without special locality, Mar. 1908, Robert Ridgeway s.n. ( US 861.309).

Pendent, slender, 15–40 cm long. Shoots gradually heterophyllous, tapering from ca ­­­ 10–25 mm in diam. incl. leaves at the base, to 3–5 (–7) mm distally, sometimes only slightly tapering. Stems excl. leaves 1–1.5 (–2) mm thick at the base. ­­­ Leaves usually gradually modified, borne in irregular, alternating whorls of 3 (or 4), these 1.5–2 (–3) mm apart; ­­­ leaves of basal divisions spreading to ascending or loosely appressed, often twisted to a vertical position, ­­­ linear-lanceolate to linear-oblanceolate, 9–15 × 1–2 mm, almost flat or somewhat concave adaxially, softly to firmly herbaceous, with flat or slightly revolute, entire margins. Leaves of middle and distal divisions usually gradually shorter, narrower and more appressed, conform, or abaxially convex, with involute margins. ­­­ Leaves of fully sporangiate divisions 6-ranked, with a widened, clasping base, often contracted into a short to long, narrow, involute apex, 2–5 × 1–1.5 mm. Sporangia 1–1.2 mm wide.

­­­ Distribution:— Costa Rica, Panamá.

­­­ Habitats:—Epiphytic in montane forest, 1000–2100 m elev.

­­­ Notes:—Closely related to Phlegmariurus taxifolius , and differing mainly in its smaller size. Intermediate specimens occur, making the distinction of the species difficult.

­­­ Specimens­­­studied:— Chiriquí: Road Gualaca–Fortuna Dam site, 5.9 mi NW of Los Planes de Hornito, 1370 m, Croat 49886 (MO). Upper Caldera River, near Camp I, Holcomb’s Trail, above El Boquete, 1450–1650 m, Maxon 5717 ( US). Los Siguas Camp, S slope of Cerro de la Horqueta, 1700 m, Maxon 5430 ( US). Cerro Horqueta, Cord. de Talamanca, above Boquete, 1750–2100 m, Cochrane 6284 (MO). Near El Boquete, 1000–1500 m, Cornman 941, 1163 ( US), 1221 ( US, UC). Bajo Chorro, Boquete Distr., 1830 m, Davidson 341 (F, MO, US). Cerro Horqueta, 8 km NW of Boquete, montane forest, N slopes, 1800 m, Maas & Dressler 4966 (U). Palo Alto, 4.5 mi NE of Boquete, along trail to Cerro La Trompeta, 1770 m, Hammel 2139 (MO). Near Boquete, trail from Bajo Chorro along the Río Caldera, 2.5 km from trailhead, 1700 m, W. Testo 989 (PMA, VT). NE del campamento de Fortuna (hornito), sitio de presa 1000–1200 m, Correa et al. 2887 (PMA).