Phlegmariurus reflexus ­­­ (Lam.) B.Øllg. var. minor (Spring) B. Øllgaard (2012a: 481) (Spring)

Øllgaard, Benjamin & Testo, Weston, 2021, The Lycopodiaceae of Panamá, Phytotaxa 526 (1), pp. 1-66 : 37

Phlegmariurus reflexus ­­­ (Lam.) B.Øllg. var. minor (Spring) B. Øllgaard (2012a: 481) .

— Lycopodium reflexum Lam. var. minus Spring (1842: 26) .— Urostachys reflexus (Lam.) Herter var. minor (Spring) Nessel (1927: 393) .— Huperzia reflexa var. minor (Spring) B. Øllgaard (1988: 26) .— SYNTYPES: Gaudichaud s. n.; Brasil, Pr. Rio de Janeiro (P); Langsdorff s. n. (M-Mart.); Guillemin s. n.; Brazil, Serra dos Orgaos, fr. Majo (P); Martius s. n.; in sylvis prov. Paraënsis (M); Claussen s. n.; Brazil, in prov. Minarum (P).

Lycopodium brutum Herter (1909a: 47) .— Urostachys brutus (Herter) Herter (1923: 162) .— Urostachys reflexus (Lam.) Herter var. brutus (Herter) Nessel (1939: 116) . TYPE:— Trinidad: Hooker ded. 1845; (P holotype; L isotype?). The type matches the Ecuadorian plants of var. minor , with nearly smooth leaf margins.

Specimens­­­studied:— Bocas­­­del­­­Toro: N of La Fortuna Dam site, Hammel 2442 (MO, P). Chiriquí: Dist. Bugaba, Santa Clara, van der Werff & Herrera 7310 (AAU, MO, UC). Dist. Gualaca, Reserva Forestal de Fortuna, Sendero del km 63 y Sendero Hornitos, Araúz et al. 758 (PMA). Parque Internacional La Amistad (PILA), Mirador La Nevera, Araúz 763 (PMA). Along Río Caldera, 1 mi below El Boquete, 1250 m, Killip 5484 (US). Near El Boquete, 1000–1500 m, Cornman 887 (UC, US). Near El Boquete, 1000–1500 m, 20 Feb 1918, Cornman 1023 (MICH, MO, UC, US). Parque Nacional Volcán Barú, sendero El Salto, 2192 m, Ríos 192, 193 (PMA). Dist. Volcán, near Las Lagunas, Cerro Pando, 6000 ft., D’Arcy 10067 (MO). 6.5 mi S of Volcán, 33 ft., Webster 16633 (MO).Alto to Cerro Copete, 2300–3300 m, Woodson & Schery 364 (GH, MO). Province above Cerro Punta, ca. 2 mi E of Guadalupe, McAlpin 1454 (photo DUKE). Road bank 15 km from Río Sereno, Salazar 446 (PMA). Cerro Punta, Florpan s.n. (AAU). Cerro Punta, Las Nubes PILA, trail La Cascada, 2442 m, Florpan 6869 (AAU). Volcán de Chiriquí, 1.2 mi from Boquete, Armond 527 (photo DUKE). Woods 2.5 mi NW of Hato del Volcán, 1800 m, Armond 211 (photo DUKE). Trail to Bajo Mona, ca. 8 km from Bajo Boquete, 1500 m, Armond 35 (photo DUKE). Upland woods 5.2 mi NW of El Hato del Volcán, 1800 m, Armond 211 (photo DUKE). Fortuna, road around the lake, Valdespino et al. 957 (PMA). Fortuna 100–200 m after bridge before vivero of IRHE, Valdespino et al. 990 (PMA). Along Río Caldera (Boquete region) 3.5 km NW of Bajo Mono, 1600 m, Smith 2467 (UC). Along Río Caldera (Boquete region) 3.5 km NW of Bajo Mono, 1600 m, Smith 2468 (UC). Trail to sawmill, El Boquete, Cornman 887 (UC). Vicinity of Barriles, 6–16 km S of Volcán, 1000–1300 m, Dressler & Williams 3991 (PMA). Coclé:­­­ Mountains beyond La Pintada, 400–600 m, Hunter & Allen 602 (MO). 3 mi N of El Valle, 582 m, Armond 366 (MO, DUKE). El Valle, Ebinger 959 (US). Dist. Antón, Valle de Antón, camino a La Mesa, Araúz 799 (PMA). Panamá: Llano Cartí, Comarca Kuna-Yala, km 12.2, Valdespino et al. 1008 (PMA).